The Combretaceae, often called the white mangrove family, are a
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s in the order
Myrtales
The Myrtales are an order of flowering plants placed as a sister to the eurosids II clade as of the publishing of the ''Eucalyptus grandis'' genome in June 2014.
The APG III system of classification for angiosperms still places it within the eu ...
. The family includes about 530 species of
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s,
shrubs, and
liana
A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ...
s in ca 10
genera
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...
.
The family includes the leadwood tree, ''
Combretum imberbe
''Combretum imberbe'' (leadwood, af, hardekool, st, mohwelere-tšhipi, ts, motswiri/mondzo, zu, impondondlovu) is a characteristic and often impressive bushwillow species of the southern Afrotropics. The medium to large tree has a sparse, s ...
''. Three genera, ''
Conocarpus
''Conocarpus'' is a genus of two species of flowering plants in the family Combretaceae, native to tropical regions of the world. One of the species is a widespread mangrove species, and the other is restricted to a small area around the southern ...
'', ''
Laguncularia
''Laguncularia'' is a genus of plants in the family Combretaceae
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lia ...
'', and ''
Lumnitzera
''Lumnitzera'' is an Indo-West Pacific mangrove genus in the family Combretaceae. An English common name is black mangrove. (However, "black mangrove" may also refer to the unrelated genus ''Avicennia''.) ''Lumnitzera'', named after the German ...
'', grow in
mangrove
A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...
habitats (mangals).
The Combretaceae are widespread in the
subtropics and
tropics
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred ...
. Some members of this family produce useful construction timber, such as idigbo from ''
Terminalia ivorensis
''Terminalia ivorensis'' is a species of tree in the family Combretaceae, and is known by the common names of Ivory Coast almond, idigbo, black afara, framire and emeri.
Description
''Terminalia ivorensis'' is found in Cameroon, Ivory Coast, Gh ...
''. The commonly cultivated ''
Quisqualis indica'' is now placed in the genus ''
Combretum
''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...
''. Many plants in the Quisqualis species contain the
Non-proteinogenic amino acid
In biochemistry, non-coded or non-proteinogenic amino acids are distinct from the 22 proteinogenic amino acids (21 in eukaryotesplus formylmethionine in eukaryotes with prokaryote organelles like mitochondria) which are naturally encoded in the ...
excitotoxin
Quisqualic acid
Quisqualic acid is an agonist of the AMPA, kainate, and group I metabotropic glutamate receptors. It is one of the most potent AMPA receptor agonists known. It causes excitotoxicity and is used in neuroscience to selectively destroy neurons ...
, a potent
AMPA agonist.
[Excitotoxic cell death and delayed rescue in human neurons derived from NT2 cells, M Munir, L Lu and P Mcgonigl, Journal of Neuroscience, Vol 15, 7847–7860]
White mangroves
The family name comes from the
type genus
In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name.
Zoological nomenclature
According to the International Code of Zoological Nomenclature, "The name-bearing type of a nomina ...
''
Combretum
''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...
''; it also includes the white mangrove, ''
Laguncularia racemosa
''Laguncularia racemosa'', the white mangrove, is a species of flowering plant in the leadwood tree family, Combretaceae. It is native to the coasts of western Africa from Senegal to Cameroon, the Atlantic Coast of the Americas from Bermuda an ...
'', found along seacoasts in tropical America and West Africa.
Genera
''
Plants of the World Online
Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants ...
'' currently includes:
[Plants of the World Online: Combretaceae R.Br. (retrieved 19 January 2022)]
/ref>
#''Combretum
''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...
'' Loefl.
#''Conocarpus
''Conocarpus'' is a genus of two species of flowering plants in the family Combretaceae, native to tropical regions of the world. One of the species is a widespread mangrove species, and the other is restricted to a small area around the southern ...
'' L.
#''Dansiea
''Dansiea'' is a genus of flowering plants belonging to the family Combretaceae
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, ...
'' Byrnes
#''Getonia
''Getonia'' is a monotypic genus of flowering plants belonging to the family Combretaceae. Its native range is India, Assam to Peninsula Malaysia.
Getonia floribunda
''Getonia floribunda'', commonly known as ukshi, is a large climbing shrub wh ...
'' Roxb. (synonym ''Calycopteris'')
#''Guiera
''Guiera'' is a flowering plant genus in the family Combretaceae. ''Guiera senegalensis'' is the only known species in the genus, found in Tropical Africa in dry areas from Senegal to Sudan (requiring much sunlight and light dry soil). The plant ...
'' Adans. ex Juss.
#''Laguncularia
''Laguncularia'' is a genus of plants in the family Combretaceae
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lia ...
'' C.F.Gaertn. (monotypic)
#''Lumnitzera
''Lumnitzera'' is an Indo-West Pacific mangrove genus in the family Combretaceae. An English common name is black mangrove. (However, "black mangrove" may also refer to the unrelated genus ''Avicennia''.) ''Lumnitzera'', named after the German ...
'' Willd.
#'' Macropteranthes'' F.Muell.
#''Strephonema
''Strephonema'' is a genus of flowering plants belonging to the family Combretaceae.
Its native range is Western and Western Central Tropical Africa
Although tropical Africa is mostly familiar to the West for its rainforests, this biogeog ...
'' Hook.f.
#'' Terminalia'' L. (synonyms: '''', ''Buchenavia
''Buchenavia'' is a genus of plant in family Combretaceae. It contains the following species (but this list may be incomplete):
* ''Buchenavia grandis'', Ducke
* ''Buchenavia hoehneana'', N. Mattos
* ''Buchenavia iguaratensis'', N. Mattos
* ''Buc ...
'', ''Bucida
''Bucida'' is a genus of flowering plants in the Indian almond family, Combretaceae. It contains the following species (but this list may be incomplete):
* '' Bucida angustifolia''
* ''Bucida buceras'' L. - Bullet tree (southern Mexico, Central ...
'', ''Pteleopsis
''Pteleopsis'' is a genus of plant in the family Combretaceae. ''Pteleopsis'' Engl. is often considered a synonym of ''Terminalia'' L., another genus of the same family which contains around 290 species.
Species include:
* ''Pteleopsis barbosae ...
'', ''Terminaliopsis'')
synonyms of ''
Combretum
''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...
''
*'' Meiostemon'' Exell & Stace
*'' Quisqualis'' L.
*''Thiloa
''Thiloa'' is a genus of the plant family Combretaceae
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lianas in ca ...
'' Eichler
References
External links
*
*
{{-
Myrtales families
Taxa named by Robert Brown (botanist, born 1773)