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''Quercus semecarpifolia'' is an Asian species of
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
. It is native to the Himalayas and nearby mountains in
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, Afghanistan, India, Nepal, and Pakistan, where it is referred to as kharsu. It is classified in subgenus ''Cerris'', section ''Ilex''. ''Quercus semecarpifolia'' is an
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
tree up to tall. The leaves are up to long, with a few teeth along the sides but rounded at the tip.Smith, James Edward in Rees. 1814 The cyclopædia; or, Universal dictionary of arts, sciences, and literature. volume 29, pages not numbered, ''Quercus'' number 20
/ref> It has been grown in middle Europe, Western Germany, winter-hardiness zone 7, withstanding -14 °C, without any damages. It gives a good, showy bush to small tree with lush green leaves. The epithet ''semecarpifolia'' refers to a resemblance between the leaves of this species and those of ''
Semecarpus anacardium ''Semecarpus anacardium'', commonly known as the marking nut tree, Malacca bean tree, marany nut, oriental cashew, dhobi nut tree and varnish tree, is a native of India, found in the outer Himalayas to the Coromandel Coast. It is closely related ...
.'' In its native range, it serves as a
keystone species A keystone species is a species that has a disproportionately large effect on its natural environment relative to its abundance. The concept was introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in main ...
, exhibiting up to 80% dominance at elevations between 2700 and 3300 meters. In areas where less degradation has occurred, ''Q. semecarpifolia'' reduces light at the forest floor, encouraging the growth of
shade tolerant In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in nursery (ho ...
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
species. Local people make use of kharsu more frequently than other native oaks for food, animal fodder, fuel, and timber. Its leaves are also known to support '' Antheraea proylei'' silk worms, goats, and prevent slug infestations.


Fossil record

Fossils of ''Quercus semecarpifolia'' have been described from the
fossil flora of Kızılcahamam district This fossil flora in Turkey stems from at least six Pliocene deposits in Güvem and Beşkonak villages, north of Ankara, north of Kızılcahamam and south of the Black Sea coast. They are to above sea level, extending north-south and eas ...
in
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
, which is of early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58line drawing, Flora of China Illustrations vol. 4, plate 368, figures 6-8 in lower half
semecarpifolia Trees of Afghanistan Trees of the Indian subcontinent Trees of Tibet Plants described in 1814 Taxa named by James Edward Smith {{Quercus-stub