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''Quercus pubescens'' (
synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
''virgiliana''), commonly known as the downy oak, pubescent oak or Italian oak, is a species of
white oak ''Quercus'' subgenus ''Quercus'' is one of the two subgenera into which the genus ''Quercus'' was divided in a 2017 classification (the other being subgenus ''Cerris''). It contains about 190 species divided among five sections. It may be calle ...
(genus ''
Quercus An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
'' sect. ''Quercus'') native to southern Europe and southwest Asia. It is found from northern
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
(
Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
) and France in the West to
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
and the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
in the East.


Description

''Quercus pubescens'' is a medium-sized
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
growing up to . Forest-grown trees grow tall, while open-growing trees develop a very broad and irregular crown. They are long-lived, to several hundred years, and eventually grow into very stout trees with trunks up to in diameter. Open-grown trees frequently develop several trunks. The
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
is very rough, light gray and divided into small flakes. Large trees develop very thick whitish bark cracked into deep furrows, similar to the
pedunculate oak ''Quercus robur'', the pedunculate oak, is a species of flowering plant in the beech and oak family, Fagaceae. It is a large tree, native to most of Europe and western Asia, and is widely cultivated in other temperate regions. It grows on soi ...
but lighter in colour. The twigs are light purple or whitish, with tomentum. The buds are small () and blunt, light brown. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are leathery usually long (rarely to 13 cm) and 3–6 cm wide, usually widest beyond the middle. The leaves group at the ends of twigs. The upper leaf surface is dark green and rough, the lower light green. Both leaf surfaces are covered with minute
pubescence Puberty is the process of physical changes through which a child's Human body, body matures into an adult body capable of sexual reproduction. It is initiated by hormone, hormonal signals from the Human brain, brain to the gonads: the ovary ...
which is sometimes lost in older leaves by late summer. The young expanding leaves are whitish or pinkish with very soft tomentum. The leaf shape is very variable, divided into 3–7 pairs of deep or shallow lobes, which are usually divided into a few sublobes. The lobes are usually blunt, rarely sharp. The apex is usually wide and round. The base of the leaf is heart shaped, widely rounded or sometimes pointed. The petioles are 4–15 mm (rarely to 22 mm) long, stout and pubescent. The leaves are persistent late into the autumn, remaining green up to early winter. They eventually turn russet or brown and fall off. The ''Quercus pubescens''
acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...
s are light brown to yellow, 8–20 mm long, usually thin and pointed. The acorn cups are light gray to almost white, with pointed, overlapping scales, covered with tomentum. The acorn stalks are thick and pubescent, up to 2 cm long. The acorns usually occur in groups of 2–5 on the same stalk.


Subspecies

Three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are accepted by ''Flora Europaea'': * ''Quercus pubescens'' subsp. ''pubescens'' – central and southern Europe. * ''Quercus pubescens'' subsp. ''anatolica'' O.Schwarz – southwest Asia, southeast Europe. * ''Quercus pubescens'' subsp. ''palensis'' (Palassou) O.Schwarz – northern Spain, Pyrenees.


Habitat

Downy oaks typically grow in dry, lime-rich
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
s. It is a sub-Mediterranean species, growing from the coastline to deep in the continent. Its optimum is in transitional
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
- oceanic climates, characterized by warm to hot, dry summers and cool (though not mild) winters with ample precipitation.


References


Flora Europaea: ''Quercus pubescens''
* Bean, W. J. (1976). ''Trees and shrubs hardy in the British Isles'' 8th ed., revised. John Murray. * Rushforth, K. (1999). ''Trees of Britain and Europe''. HarperCollins . *


''Quercus pubescens''
an
''Quercus virgiliana''
- information, genetic conservation units and related resources.
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that promotes the conservation and sustainable use of forest genetic resources in Europe as an integral part of sustainable forest management. It was established ...
(EUFORGEN) {{Authority control pubescens Flora of Western Asia Trees of Mediterranean climate Trees of Europe Plants described in 1796