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The hard clam ('' Mercenaria mercenaria''), also known as the round clam, hard-shell (or hard-shelled) clam, or the quahog, is an edible marine
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, biv ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is es ...
that is native to the eastern shores of North America and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
from
Prince Edward Island Prince Edward Island (PEI; ) is one of the thirteen provinces and territories of Canada. It is the smallest province in terms of land area and population, but the most densely populated. The island has several nicknames: "Garden of the Gulf", ...
to the
Yucatán Peninsula The Yucatán Peninsula (, also , ; es, Península de Yucatán ) is a large peninsula in southeastern Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north ...
. It is one of many unrelated edible bivalves that in the United States are frequently referred to simply as
clam Clam is a common name for several kinds of bivalve molluscs. The word is often applied only to those that are edible and live as infauna, spending most of their lives halfway buried in the sand of the seafloor or riverbeds. Clams have two sh ...
s, as in the expression " clam digging". Older literature sources may use the systematic name ''Venus mercenaria''; this species is in the family
Veneridae The Veneridae or venerids, common name: Venus clams, are a very large family of minute to large, saltwater clams, marine bivalve molluscs. Over 500 living species of venerid bivalves are known, most of which are edible, and many of which are ...
, the venus clams. Confusingly, the "ocean quahog" is a different species, '' Arctica islandica'', which, although superficially similar in shape, is in a different family of bivalves: it is rounder than the hard clam, usually has black
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods a ...
, and there is no pallial sinus in the interior of the shell.


Alternative names

The hard clam has many alternative
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contra ...
s. It is also known as the Northern quahog, round clam, or chowder clam. In fish markets, there are specialist names for different sizes of this species of clam. The smallest legally harvestable clams are called countnecks or peanuts, next size up are littlenecks, then topnecks. Above that are the cherrystones, and the largest are called quahogs or chowder clams. Rice, M.A. (1992). The Northern Quahog: Biology of ''Mercenaria mercenaria''. Rhode Island Sea Grant Publication No. RIU-B-92-001, University of Rhode Island, Narragansett. 60 pp.
web link
The most distinctive of these names is ''quahog'' ( , , or , also spelt ''quahaug'', ''quohog'' or ''cohog''). The word comes from the Narragansett word "poquauhock", which is similar in
Wampanoag The Wampanoag , also rendered Wôpanâak, are an Indigenous people of the Northeastern Woodlands based in southeastern Massachusetts and historically parts of eastern Rhode Island,Salwen, "Indians of Southern New England and Long Island," p. 1 ...
and some other
Algonquian languages The Algonquian languages ( or ; also Algonkian) are a subfamily of indigenous American languages that include most languages in the Algic language family. The name of the Algonquian language family is distinguished from the orthographically simi ...
; it is first attested in North American English in 1794. Native polities on the eastern Atlantic seaboard made valuable beads called
wampum Wampum is a traditional shell bead of the Eastern Woodlands tribes of Native Americans. It includes white shell beads hand-fashioned from the North Atlantic channeled whelk shell and white and purple beads made from the quahog or Western N ...
from the shells, especially those colored purple; the species name ''mercenaria'' is related to the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
word for commerce. Today people living in coastal New England still use Algonquian words for the clam, as they have done for thousands of years. In many areas where
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lotus ...
is important, clam farmers have bred specialized versions of these clams with distinctions needed for them to be distinguished in the marketplace. These are quite similar to common "wild type" ''Mercenaria'' clams, except that their shells bear distinctive markings. These are known as the ''notata'' strain of quahogs, which occur naturally in low numbers wherever quahogs are found.


Distribution

Hard clams are quite common throughout
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian province ...
, north into
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
, and all down the Eastern seaboard of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
to
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, a ...
; but they are particularly abundant between
Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of mainland Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer mon ...
and
New Jersey New Jersey is a U.S. state, state in the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. It is bordered on the north and east by the state of New York (state), New York; on the ea ...
, where seeding and harvesting them is an important commercial form of
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lotus ...
. For example, the species is an important member of the suspension-feeding,
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
fauna of the lower
Chesapeake Bay The Chesapeake Bay ( ) is the largest estuary in the United States. The Bay is located in the Mid-Atlantic (United States), Mid-Atlantic region and is primarily separated from the Atlantic Ocean by the Delmarva Peninsula (including the parts: the ...
.
Rhode Island Rhode Island (, like ''road'') is a state in the New England region of the Northeastern United States. It is the smallest U.S. state by area and the seventh-least populous, with slightly fewer than 1.1 million residents as of 2020, but i ...
is situated right in the middle of "quahog country" and has supplied a quarter of the U.S.'s total annual commercial quahog catch. The quahog is the official shellfish of the U.S. state of
Rhode Island Rhode Island (, like ''road'') is a state in the New England region of the Northeastern United States. It is the smallest U.S. state by area and the seventh-least populous, with slightly fewer than 1.1 million residents as of 2020, but i ...
. The species has also been introduced and is farmed on the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
coast of North America and in
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
and continental
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
. It reproduces sexually by females and males shedding
gametes A gamete (; , ultimately ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. In species that produce ...
into the water.


Parasite

Quahog parasite unknown (QPX) is a
parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
that affects the hard shell clam ''Mercenaria mercenaria''. While little is known about the disease, research is currently under way in several laboratories. This research is fueled by the need to inform aquaculturists, who suffer financially because of the mortality rates in clams that QPX inflicts and the ensuing years in which runs must be left
fallow Fallow is a farming technique in which arable land is left without sowing for one or more vegetative cycles. The goal of fallowing is to allow the land to recover and store Organic compound, organic matter while retaining moisture and disrupting ...
to clear the disease. It was discovered along the coast of
Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of mainland Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer mon ...
, Massachusetts in 1995. Quahog parasite X (or quahog parasite unknown PX disease of the hard clam ''Mercenaria mercenaria'' is caused by a poorly known protistan parasite. Its DNA sequence analysis places the QPX parasite among the thraustochytrid stramenopiles. Thraustochytrids are common protists in marine sediments and the water column, but only a few thraustochytrids are known as parasites of marine animals. Although QPX disease was first recorded on the Atlantic coast of Canada in the early 1960s, it did not become a major economic problem until its appearance in cultured clams at Prince Edward Island, Massachusetts in 1992, and Virginia in 1997. Infected clams are characterized by the presence of blisters or pustules in the mantle and later by gaping and death.


Human use

In coastal areas of the New England states, Long Island, and New Jersey, restaurants known as
raw bars A raw bar is a small restaurant or a bar within a restaurant where live shellfish are shucked and served. Raw bars typically offer a variety of raw and cooked seafood and shellfish that is served cold. Seafood-based dishes may also be offered, an ...
or clam bars specialize in serving littlenecks and topnecks raw on an opened half-shell, usually with a cocktail sauce with
horseradish Horseradish (''Armoracia rusticana'', syn. ''Cochlearia armoracia'') is a perennial plant of the family Brassicaceae (which also includes mustard, wasabi, broccoli, cabbage, and radish). It is a root vegetable, cultivated and used worldwide ...
, and often with
lemon The lemon (''Citrus limon'') is a species of small evergreen trees in the flowering plant family Rutaceae, native to Asia, primarily Northeast India (Assam), Northern Myanmar or China. The tree's ellipsoidal yellow fruit is used for culin ...
. Sometimes littlenecks are steamed and dipped in butter, though not as commonly as their soft-shelled clam cousin the "steamer". Littlenecks are often found in-the-shell in sauces, soups, stews, and clams casino, or substituted for European varieties such as the cockle in southern European seafood dishes. The largest clams are quahogs or chowders and cherrystones; they have the toughest meat and are used in such dishes as clam chowder, clam cakes, and
stuffed clam Stuffed clams (or stuffies) are popular in New England, especially in Rhode Island, and consist of a breadcrumb and minced clam mixture that is baked on the half shell of a quahog hard shell clam. Other ingredients typically found in the basic ...
s, or are minced and mixed into dishes that use the smaller, more tender clams. Historically, Native Americans used the quahog as a component in
wampum Wampum is a traditional shell bead of the Eastern Woodlands tribes of Native Americans. It includes white shell beads hand-fashioned from the North Atlantic channeled whelk shell and white and purple beads made from the quahog or Western N ...
, the shell beads exchanged in the
North American fur trade The North American fur trade is the commercial trade in furs in North America. Various Indigenous peoples of the Americas traded furs with other tribes during the pre-Columbian era. Europeans started their participation in the North American fu ...
. The Narragansetts used the hard clam for food and ornaments. A population of hard clams exists in
Southampton Water Southampton Water is a tidal estuary north of the Solent and the Isle of Wight in England. The city of Southampton lies at its most northerly point, where the estuaries of the River Test and River Itchen meet. Along its salt marsh-fringed we ...
in
Hampshire Hampshire (, ; abbreviated to Hants) is a ceremonial county, ceremonial and non-metropolitan county, non-metropolitan counties of England, county in western South East England on the coast of the English Channel. Home to two major English citi ...
,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...
. Originally bred in the warm water outflows at
Southampton Power Station Southampton Power Station was a coal fired power station built by Southampton Corporation that operated between 1904 and 1977. History The Southampton Electric Lighting and Power Company supplied electricity to Southampton from 1891, from a ...
for use as eel bait, the population became self-sustaining and can now be found in Southampton Water and has also spread to
Portsmouth Harbour Portsmouth Harbour is a biological Site of Special Scientific Interest between Portsmouth and Gosport in Hampshire. It is a Ramsar site and a Special Protection Area. It is a large natural harbour in Hampshire, England. Geographically it ...
and Langstone Harbour.


Clams and red tide

The term "
red tide A harmful algal bloom (HAB) (or excessive algae growth) is an algal bloom that causes negative impacts to other organisms by production of natural phycotoxin, algae-produced toxins, mechanical damage to other organisms, or by other means. HABs are ...
" refers to an accumulation of a toxin, such as
saxitoxin Saxitoxin (STX) is a potent neurotoxin and the best-known paralytic shellfish toxin (PST). Ingestion of saxitoxin by humans, usually by consumption of shellfish contaminated by toxic algal blooms, is responsible for the illness known as paraly ...
, produced by marine algae. Filter-feeding shellfish are affected, such as clams, oysters, and mussels. The toxin affects the human
central nervous system The central nervous system (CNS) is the part of the nervous system consisting primarily of the brain and spinal cord. The CNS is so named because the brain integrates the received information and coordinates and influences the activity of all p ...
. Eating contaminated shellfish, raw or cooked, can be fatal. Some other kinds of algal blooms make the seawater appear red, but red tide blooms do not always discolor the water, nor are they related to tides.


Notes


References

{{DEFAULTSORT:Hard Clam Clams Veneridae Marine molluscs of North America Molluscs of the Atlantic Ocean Molluscs of the United States Molluscs of Mexico Fauna of the Northeastern United States Fauna of the Southeastern United States Molluscs described in 1758 Taxa named by Carl Linnaeus Symbols of Rhode Island Rhode Island cuisine Native American culture Seafood in Native American cuisine