Pyrorchis
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''Pyrorchis'', commonly known as beak orchids, is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s in the orchid family,
Orchidaceae Orchids are plants that belong to the family (biology), family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan distribution, cosmopolitan plants that ...
and is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
. It contains two species which were previously included in the genus ''
Lyperanthus ''Lyperanthus'', commonly known as beak orchids, is a genus of flowering plants from the orchid family, Orchidaceae, that is endemic to Australia. There are two species, one in Western Australia and the other in four eastern Australian states, di ...
'', also known as beak orchids. Both species have fleshy, oval leaves and form colonies which flower profusely after bushfires.


Description

Orchids in the genus ''Pyrorchis'' are
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
,
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
,
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
,
sympodial In botany, sympodial growth is a bifurcating branching pattern where one branch develops more strongly than the other, resulting in the stronger branches forming the primary shoot and the weaker branches appearing laterally. A sympodium, als ...
herbs with a few inconspicuous, fine roots and an oval-shaped
tuber Tubers are a type of enlarged structure that plants use as storage organs for nutrients, derived from stems or roots. Tubers help plants perennate (survive winter or dry months), provide energy and nutrients, and are a means of asexual reproduc ...
lacking a protective sheath. Replacement tubers are formed on the end of long, thin root-like
stolon In biology, a stolon ( from Latin ''wikt:stolo, stolō'', genitive ''stolōnis'' – "branch"), also known as a runner, is a horizontal connection between parts of an organism. It may be part of the organism, or of its skeleton. Typically, animal ...
s. There are between one and three broad, fleshy, egg-shaped to heart-shaped leaves at the base of the plant. The leaves are long, wide and bright green, sometimes with black markings. There are up to eight resupinate
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s on a stalk high. The stalk is often surrounded by two or three loose, leaf-like
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
s up to long. The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
is lance-shaped, about long and forms a hood over the
column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member ...
. The
lateral Lateral is a geometric term of location which may also refer to: Biology and healthcare * Lateral (anatomy), a term of location meaning "towards the side" * Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx * Lateral release ( ...
sepals are about the same length and the same size and shape as the petals. As is usual in orchids, one petal is highly modified as the central labellum. The labellum is gently curved near its base where it encloses the base of the
column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member ...
but more strongly curved near its tip, which is deeply fringed. The column is about , straight or gently curved with the
anther The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
at its tip. Flowering occurs between August and early December, depending on species and the fruit that follows flowering is a non-fleshy,
dehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part. Structures that op ...
capsule containing up to 500 seeds.


Taxonomy and naming

Robert Brown Robert Brown may refer to: Robert Brown (born 1965), British Director, Animator and author Entertainers and artists * Washboard Sam or Robert Brown (1910–1966), American musician and singer * Robert W. Brown (1917–2009), American printmaker ...
first formally described ''Lyperanthus nigricans'' in 1810, publishing the description in ''
Prodromus Florae Novae Hollandiae ''Prodromus Florae Novae Hollandiae et Insulae Van Diemen'' (Prodromus of the Flora of New Holland and Van Diemen's Land) is a book by the botanist Robert Brown Robert Brown may refer to: Robert Brown (born 1965), British Director, Animator and ...
''. In 1994, David Jones and Mark Clements described the genus ''Pyrorchis'' and included the species that Brown had described. The name "''Pyrorchis''" is derived from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
word ''pyr'' meaning "fire" and ''orchis'' meaning "orchid". There are two species: *'' Pyrorchis forrestii'' (
F.Muell. Baron Sir Ferdinand Jacob Heinrich von Mueller, (; 30 June 1825 – 10 October 1896) was a German-Australian physician, geographer, and most notably, a botanist. He was appointed government botanist for the then colony of Victoria (state), Vic ...
) D.L.Jones & M.A.Clem.
– pink beaks (Western Australia) *'' Pyrorchis nigricans'' ( R.Br.) D.L.Jones & M.A.Clem. – red beaks, elephants' ears ( N.S.W.,
Victoria Victoria most commonly refers to: * Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India * Victoria (state), a state of Australia * Victoria, British Columbia, Canada, a provincial capital * Victoria, Seychelles, the capi ...
,
Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...
,
South Australia South Australia (commonly abbreviated as SA) is a States and territories of Australia, state in the southern central part of Australia. With a total land area of , it is the fourth-largest of Australia's states and territories by area, which in ...
,
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
and A.C.T.)


Distribution and habitat

''Pyrorchis forrestii'' is only found in the south-west of
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
between Augusta and
Waychinicup National Park Waychinicup National Park is in Western Australia, southeast of Perth and east of Albany, Western Australia, Albany, along the coast of the Southern Ocean. Location and description The park is bordered by the Southern Ocean to the south, Moun ...
where it grows near the margins of swamps or near creeks. ''Pyrorchis nigricans'' occurs much more widely, growing in N.S.W.,
Victoria Victoria most commonly refers to: * Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India * Victoria (state), a state of Australia * Victoria, British Columbia, Canada, a provincial capital * Victoria, Seychelles, the capi ...
,
Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...
,
South Australia South Australia (commonly abbreviated as SA) is a States and territories of Australia, state in the southern central part of Australia. With a total land area of , it is the fourth-largest of Australia's states and territories by area, which in ...
, and the A.C.T. in a range of moist habitats. In drier areas of Western Australia, it is usually found in moist places near
granite Granite ( ) is a coarse-grained (phanerite, phaneritic) intrusive rock, intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly coo ...
outcrops.


Ecology

Both species of ''Pyrorchis'' produce small amounts of nectar and have a strong, sweet smell, especially in hot weather, suggesting pollination by bees although only ''P. forrestii'' has brightly coloured flowers. Both species flower profusely after fire, sometimes after other kinds of disturbance such as
mowing A mower is a person or machine that cuts (mows) grass or other plants that grow on the ground. Usually mowing is distinguished from reaping, which uses similar implements, but is the traditional term for harvesting grain crops, e.g. with reaper ...
, otherwise only one or two are seen after hot summer days.


Conservation

Both species of ''Pyrorchis'' are classified as "Not Threatened" by the Western Australian Government
Department of Parks and Wildlife The Department of Parks and Wildlife (DPaW) was the department of the Government of Western Australia responsible for managing lands described in the ''Conservation and Land Management Act 1984'' and implementing the state's conservation and en ...
.


See also

*
List of Orchidaceae genera This is a list of genera in the orchid family (Orchidaceae), originally according tThe Families of Flowering Plants- L. Watson and M. J. Dallwitz. This list is adapted regularly with the changes published in the ''Orchid Research Newsletter'' which ...


References


External links

*
Retired Aussies, ''Pyrorchis nigricans'', Undertaker Orchid
{{Taxonbar, from=Q3292390 Diurideae genera