The Cinquefoil Skipper (''Pyrgus cirsii'') is a species of
skipper butterfly (family Hesperiidae).
Description

This species has a
wingspan
The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
of 26–28 mm and, typically of its
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
, can be very difficult to identify in the field.
''Pyrgus cirsii'' is a medium-sized skipper. There is basically a possibility of confusion with other species of the genus ''
Pyrgus
''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' C ...
'', in particular with ''
Pyrgus armoricanus
''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and ''Ca ...
''. Characteristic of ''Pyrgus cirsii'' is a large square discoidal spot on the upper side of the forewing and almost always a second point-like mark over the basal spot on the upper side of the forewing. The hind wing underside is often ocher to reddish brown and stands out from other species of the genus.
It is sometimes considered a
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the
Carline Skipper
The carline skipper (''Pyrgus carlinae'') is a butterfly and a species of the skipper (family Hesperiidae). It is a montane butterfly only found in southwestern areas of the Alps. It can be an abundant species within this restricted range.
As ...
(''P. carlinae'') as the two are known to
interbreed
In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different variety (botany), varieties, subspecies, species or genus, genera through sexual reproduction. Generally, it means that each cell has gene ...
where their ranges overlap in the Alps but can usually be recognised by the prominent squarish, rather than c-shaped, white spot near the
costa
Costa may refer to:
Biology
* Rib (Latin: ''costa''), in vertebrate anatomy
* Costa (botany), the central strand of a plant leaf or thallus
* Costa (coral), a stony rib, part of the skeleton of a coral
* Costa (entomology), the leading edge of th ...
of the forewing and the usually deeper red-brown underwings. A reliable separation in this genus, which is difficult to determine according to external characteristics, is possible through genital examination.
Range
''Pyrgus cirsii'' is widespread from Spain to Central Europe.
According to current knowledge, there is also evidence from the Caucasus and Eastern Anatolia, as there are several isolated populations in north-eastern
Turkey
Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
and in
Armenia
Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
.
[Butterfly Conservation Armenia http://www.butterfly-conservation-armenia.org/pyrgus-cirsii.html]
Habitat
The current habitats of ''Pyrgus cirsii'' are sparsely overgrown, with open ground interspersed, full sun dry grasslands in stony, shallow and base-rich locations. Almost without exception, the current habitats are subject to regular and relatively intensive grazing, and they are also characterized by large pads of ''Potentilla verna'' agg. In the past, extremely sparse pine stands were also populated in Bavaria.
In addition to these qualitative characteristics, the overall size of the habitat obviously plays an important role. E. g. all Bavarian occurrences are or were in areas with large, contiguous grassland complexes. A minimum size of the living space with corresponding open ground structures or the very close connection of these dry turf areas is obviously essential.
Ecology
''Pyrgus cirsii'' forms one generation per year, at least in Bavaria. It appears relatively late in the year from late July to mid-September. As a rule, the flight season begins at the end of July / beginning of August with a peak in the second half of August. The last moths, mostly females, can be observed in mid-September.
The reports of finds in Bavaria range from 160 m in Main Franconia to around 500 m in the Northern
Franconian Jura
The Franconian Jura ( , , or ) is an upland in Franconia, Bavaria, Germany. Located between two rivers, the Danube in the south and the Main in the north, its peaks reach elevations of up to and it has an area of some 7053.8 km2. Emil Mey ...
, with a focus of 280 m to 400 m.
When visiting flowers, the
imago
In biology, the imago (Latin for "image") is the last stage an insect attains during its metamorphosis, its process of growth and development; it is also called the ''imaginal'' stage ("imaginal" being "imago" in adjective form), the stage in wh ...
s (males and females) behave opportunistically and use a wide range of different colored flowers. The males also like to suckle on moist soil.
''Pyrgus cirsii'' can occur in numbers in suitable habitats.
According to current knowledge, ''Pyrgus cirsii'' caterpillars probably lives exclusively on cinquefoil species (
Potentilla
''Potentilla'' is a genus containing over 500 species of Annual plant, annual, Biennial plant, biennial and Perennial plant, perennial herbaceous plant, herbaceous flowering plants in the rose family (biology), family, Rosaceae.
Potentillas m ...
,
Rosaceae
Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera.
The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...
). In Central Europe by far the most important is the spring cinquefoil (''Potentilla verna'' agg. = ''
Potentilla tabernaemontani''). Other proven caterpillar forage plants are ''
Potentilla cinerea,
Potentilla incana
''Potentilla incana'' is a plant species in the genus ''Potentilla
''Potentilla'' is a genus containing over 500 species of Annual plant, annual, Biennial plant, biennial and Perennial plant, perennial herbaceous plant, herbaceous flowering ...
''
''Potentilla arenaria'' ''
Potentilla hirta'' and ''
Potentilla pusilla''.
''Potentilla verna'' agg. has so far been proven as a larval food and egg-laying plant from
Bavaria
Bavaria, officially the Free State of Bavaria, is a States of Germany, state in the southeast of Germany. With an area of , it is the list of German states by area, largest German state by land area, comprising approximately 1/5 of the total l ...
. ''Potentilla verna'' agg. is also mentioned as a host plant from
Baden-Württemberg
Baden-Württemberg ( ; ), commonly shortened to BW or BaWü, is a states of Germany, German state () in Southwest Germany, east of the Rhine, which forms the southern part of Germany's western border with France. With more than 11.07 million i ...
and
Switzerland
Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
. ''
Potentilla incana
''Potentilla incana'' is a plant species in the genus ''Potentilla
''Potentilla'' is a genus containing over 500 species of Annual plant, annual, Biennial plant, biennial and Perennial plant, perennial herbaceous plant, herbaceous flowering ...
'' is given for
Alsace
Alsace (, ; ) is a cultural region and a territorial collectivity in the Grand Est administrative region of northeastern France, on the west bank of the upper Rhine, next to Germany and Switzerland. In January 2021, it had a population of 1,9 ...
.
After laying eggs in August and September, the caterpillar overwinters until the coming spring. The first newly hatched caterpillars were found on the
Swabian Alb
The Swabian Jura ( , more rarely ), sometimes also named Swabian Alps in English, is a mountain range in Baden-Württemberg, Germany, extending from southwest to northeast and in width. It is named after the region of Swabia. It is part of th ...
in March. The caterpillars live in the leaf bags characteristic of the ''
Pyrgus
''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' C ...
'' species and can therefore be searched for in a targeted manner. The development of the caterpillar to the imagos takes a relatively long time with over three months.
Both sexes of ''Pyrgus cirsii'' are based on the structure of the terrain and specifically seek out the areas with little vegetation. The males occupy territories here and defend them against their male counterparts. Almost all imagos that fly by are checked for a short period of time. Females that are ready to lay eggs fly very low and land on the sparse vegetation of the raw soil in search of places to lay their eggs. Recognizing the right food plant is obviously not possible "from the air", which is shown by the chance landing on very different plant species. After landing, it immediately checks the suitability of the plant by lowering the feelers on the leaves. If it is not a cinquefoil, the female continues to look for neighboring plants in the area, which are checked again in this way. If the search for suitable cinquefoil plants is successful, various leaves of the same are thoroughly drummed with the antennae. If a leaf is finally assessed as suitable, the eggs are usually laid on the underside of the leaf, sometimes also on the stem of the food plant.
Conservation
Like most ''Pyrgus'' species, ''Pyrgus cirsii'' needs a fairly large habitat and colonizes xerothermic limestone grasslands. The caterpillars are only found in the leanest, driest places. These places are in decline due to eutrophication, among other things from the air, decline in migratory sheep farming, in addition to the general loss of habitat due to loss of all kinds of grassland communities and climate changes. Increasing isolation might be a main reason for the strongly regressive population development of the species. As soon as the vegetation becomes denser and ''
Potentilla
''Potentilla'' is a genus containing over 500 species of Annual plant, annual, Biennial plant, biennial and Perennial plant, perennial herbaceous plant, herbaceous flowering plants in the rose family (biology), family, Rosaceae.
Potentillas m ...
'' stands between denser, higher plants, no successful larval development and usually no more oviposition can be expected.
For ''Pyrgus cirsii'', protective measures have been introduced and maintenance measures have been carried out in the
Steigerwald
The Steigerwald () is a hill region up to in the Bavarian-Franconian part of the South German Scarplands between Würzburg and Nuremberg. It is part of the Keuper Uplands, and within it, it is continued to the north-northeast and right of the ...
since 1998 as part of a species aid program. This resulted in clearing of bushes and clearing of grasslands to enlarge the current habitats. At the same time, de-bushed areas were reintegrated into regular grazing and are now becoming more or more intensive, i.e. grazed at a higher frequency. The current habitats are all grazed with sheep and goats at least twice a year, usually much more frequently.
Taxonomy
Some authors only accept the separation of ''P. carlinae'' and ''P. cirsii'' at the subspecies level.
External links
Fauna EuropaeaLepiforum.deButterfly Conservation Armenia
References
;
;Bibliography
* Database entry includes justification for why this species is vulnerable.
{{Taxonbar, from=Q1128556
Hesperiidae
Pyrgus
Butterflies described in 1839
Butterflies of Europe
Butterflies of Asia