Pyramidella Nisoides
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''Pyramidella'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of minute to medium-sized
sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
s, marine
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
in the subfamily Pyramidellinae of the family
Pyramidellidae Pyramidellidae, common name the pyram family, or pyramid shells, is a voluminous taxonomic family of mostly small and minute ectoparasitic sea snails, marine heterobranch gastropod molluscs. The great majority of species of pyrams are micromol ...
, the pyrams and their allies. ''Pyramidella'' is the
type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...
of the family Pyramidellidae.


Nomenclature

The generic name ''Pyramidella'' Lamarck, 1799, has been validated under the Plenary Powers of the
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
, Opinion 386 (1955). The name ''Obeliscus'' Anonymus, 1797 ''Obeliscus'' Gray, 1847">Obeliscus.html" ;"title=" ''Obeliscus"> ''Obeliscus'' Gray, 1847has also been used for this genus, whereas the name ''Pyramidella'' was used for the genus now known as ''Otopleura'' Fischer, 1885.


General description

The shell has an elongate-conic shape, slightly striated. The many Whorl (mollusc), whorls of the teleoconch are usually inflated and regularly increasing. They are longitudinally ribbed or smooth. The semioval Aperture (mollusc), aperture is entire, widened at its base and rounded in front. .The
columella Lucius Junius Moderatus Columella (, Arabic: ) was a prominent Roman writer on agriculture in the Roman Empire. His in twelve volumes has been completely preserved and forms an important source on Roman agriculture and ancient Roman cuisin ...
has one to three folds. The sharp outer lip is entire, inferiorly sub-perforated and often plicate within with three transverse folds. The shell is usually larger than in ''
Turbonilla ''Turbonilla'' is a large genus of ectoparasitic sea snails, marine gastropod molluscs in the family Pyramidellidae, the pyrams and their allies.Bouchet, P.; Gofas, S. (2011). ''Turbonilla'' Risso, 1826. Accessed through: World Register of Mar ...
.'' The thin operculum is horny, ovate, elongated, with very fine elements and oblique folds and folds oblique. The shell lacks an
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and Subcutaneous tissue, hypodermis. The epidermal layer provides a barrier to infection from environmental pathogens and regulates the ...
.Kiener (1840). General species and iconography of recent shells : comprising the Massena Museum, the collection of Lamarck, the collection of the Museum of Natural History, and the recent discoveries of travellers; Boston :W.D. Ticknor,1837
/ref> The shells are generally quite small, of an elegant form, elongated, marked with more or less deep spots. The general color of the body of these mollusca, is of a dull white. The operculum only, and the edge of the mantle are yellowish. The animal has a head surmounted by broad, rather long tentacles, formed like a pointed horn, and open on the side. The mouth is situated in a flattened, wide, dilated, and quite deeply two-lobed muzzle, separated from the foot by a groove. The respiratory cavity open throughout the whole length of the mantle, bearing on its right margin a long and narrow branchia. The heart has a direction from right to left. The rectum and uterus are attached to each other along the branchiae, and terminate in an ear-shaped gutter, produced by the margin of the mantle. The mucous follicles line the upper wall of the cavity. The foot is round, without a marginal furrow, passing down before in form of an ear-shaped shield, and bearing posteriorly a membranous operculum, with laminas destitute of a spire, which possesses one or two notches to slip upon the folds of the columella.


Species

Species within the genus ''Pyramidella'' include: * '' Pyramidella adamsi'' Carpenter, 1864 * '' Pyramidella africana'' Bartsch, 1915 * '' Pyramidella alfredensis'' Bartsch, 1915 * '' Pyramidella ava'' Bartsch, 1926 * '' Pyramidella bairdi'' Dall & Bartsch, 1909 * '' Pyramidella bicolor'' Menke, 1854 * '' Pyramidella camara'' Bartsch, 1927 * † '' Pyramidella canaliculata'' Gabb, 1873 * '' Pyramidella corrugata'' Lamarck, 1822 * ''
Pyramidella dolabrata ''Pyramidella dolabrata'', common name the "giant Atlantic pyram", is a somewhat variable species of small to medium-sized (max size 36 mm) sea snail, a marine animal, marine gastropod mollusk in the (mostly minute) family Pyramidellidae, th ...
'' (Linnaeus, 1758) * '' Pyramidella elenensis'' Bartsch, 1924 * '' Pyramidella fastigium'' (A. Adams, 1855) * '' Pyramidella gracilis'' A. Adams * '' Pyramidella guardiarioorum'' Poppe, Tagaro & Stahlschmidt, 2015 * '' Pyramidella hancocki'' Strong & Hertlein, 1939 * '' Pyramidella hastata'' (A. Adams in Sowerby, 1854) * '' Pyramidella linearum'' Pilsbry & Lowe, 1932 * '' Pyramidella magdalenensis'' Bartsch, 1917 * '' Pyramidella mexicana'' Dall & Bartsch, 1909 * '' Pyramidella minor'' (E. A. Smith, 1904) * '' Pyramidella moffati'' Dall & Bartsch, 1906 * † '' Pyramidella nanggulanica'' Finlay, 1927 * '' Pyramidella nisoides'' E. von Martens, 1904 * '' Pyramidella ovata'' de Folin, 1873 * '' Pyramidella panamensis'' Dall &Bartsch, 1909 * '' Pyramidella plicata'' (Laseron, 1959) - unreplaced junior homonym of ''Pyramidella plicata'' Lamarck, 1816 * '' Pyramidella pyrrha'' Bartsch, 1915 * '' Pyramidella sanctaehelenae'' (E. A. Smith, 1890) * '' Pyramidella scitula'' (A. Adams, 1854) * '' Pyramidella subdolabrata'' (Mörch, 1854) * '' Pyramidella subglabra'' Odhner, 1919 * '' Pyramidella terebelloides'' (A. Adams, 1854) * '' Pyramidella turrita'' A. Adams, 1855 ;Taxa inquirenda: * '' Pyramidella curtissima'' Locard, 1897 * '' Pyramidella obeliscus'' W. H. Turton, 1932 * '' Pyramidella suavissima'' Preston, 1905 * '' Pyramidella sykesi'' Preston, 1905 (original combination) * '' Pyramidella tarpeia'' Bartsch, 1915 ;Subgenera and species brought into synonymy: * ''Pyramidella (Agatha)'' A. Adams, 1860: synonym of '' Agatha'' A. Adams, 1860 * ''Pyramidella (Longchaeus)'' Mörch, 1875: synonym of '' Longchaeus'' Mörch, 1875 * ''Pyramidella (Syrnola)'' A. Adams, 1860: synonym of '' Syrnola'' A. Adams, 1860 * ''Pyramidella (Tiberia)'' Dall & Bartsch, 1906: synonym of '' Tiberia'' Jeffreys, 1884 * ''Pyramidella (Voluspa)'' Dall & Bartsch, 1904: synonym of '' Longchaeus'' Mörch, 1875 * ''Pyramidella achates'' (Gould, 1853): synonym of '' Longchaeus achates'' (A. Gould, 1853) * ''Pyramidella aclis'' A. Adams, 1853: synonym of '' Rissosyrnola aclis'' (A. Adams, 1853) (original combination) * ''Pyramidella acus'' (Gmelin, 1791) - needle pyram: synonym of '' Longchaeus acus'' (Gmelin, 1791) * ''Pyramidella affinis'' (Laseron, 1959): synonym of '' Chrysallida affinis'' (Laseron, 1959) * ''Pyramidella aganea'' Bartsch, 1915: synonym of ''
Syrnola aganea ''Syrnola aganea'' is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.Rosenberg, G. (2011). ''Syrnola aganea'' (Bartsch, 1915). Accessed through: World Register of Marine Species at ...
'' (Bartsch, 1915) * ''Pyramidella ambigua'' Gould, 1849: synonym of '' Rissoina ambigua'' (Gould, 1849) (original combination) * ''Pyramidella aprustica'' Crema, 1903: synonym of '' Euparthenia bulinea'' (Lowe, 1841) * ''Pyramidella arenosa'' Conrad, 1843: synonym of ''
Longchaeus suturalis ''Longchaeus suturalis'', common name the crenulated pyram, is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.WoRMS (2010). ''Longchaeus suturalis'' (H. C. Lea, 1843). Accessed throu ...
'' (H. C. Lea, 1843) * ''Pyramidella auricoma'' Dall, 1889: synonym of '' Longchaeus auricoma'' (Dall, 1889) * ''Pyramidella auriscati'' (Chemnitz, 1795): synonym of '' Otopleura auriscati'' (Holten, 1802) * ''Pyramidella balteata'' (A. Adams, 1855): synonym of '' Tiberia balteata'' (A. Adams, 1855) * ''Pyramidella canaliculata'' (G.B. Sowerby III, 1874): synonym of ''Pyramidella insularum'' Pilsbry, 1922 * † ''Pyramidella cancellata'' Nyst, 1836: synonym of † '' Sandbergeria cancellata'' (Nyst, 1836) † * ''Pyramidella candida'' Mørch, 1875: synonym of '' Longchaeus candidus'' (Mörch, 1875) * ''Pyramidella capensis'' (Thiele, 1925): synonym of '' Odostomia capensis'' Thiele, 1925 * ''Pyramidella ceylanica'' Preston, 1905: synonym of '' Kleinella ceylanica'' (Preston, 1905) * ''Pyramidella cincta'' Reeve, 1842: synonym of '' Milda cincta'' (Reeve, 1842) * ''Pyramidella conica'' C.B. Adams, 1852: synonym of '' Longchaeus conicus'' (C.B. Adams, 1852) * ''Pyramidella crenulata'' (Holmes, 1859): synonym of ''
Longchaeus suturalis ''Longchaeus suturalis'', common name the crenulated pyram, is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.WoRMS (2010). ''Longchaeus suturalis'' (H. C. Lea, 1843). Accessed throu ...
'' (H. C. Lea, 1843) * ''Pyramidella denticulata'' (G.B. Sowerby II, 1865): synonym of '' Colsyrnola ornata'' (Gould, 1861) * † ''Pyramidella diademata'' Maury, 1917: synonym of ''
Longchaeus suturalis ''Longchaeus suturalis'', common name the crenulated pyram, is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.WoRMS (2010). ''Longchaeus suturalis'' (H. C. Lea, 1843). Accessed throu ...
'' (H. C. Lea, 1843) * † ''Pyramidella exarata'' Michaud, 1838: synonym of † '' Palaeostoa exarata'' (Michaud, 1838) (new combination) * ''Pyramidella folinii'' Dall, 1889: synonym of ''
Longchaeus folinii ''Longchaeus folinii'' is a species of sea snail, a marine gastropod mollusc in the family Pyramidellidae, the pyrams and their allies.Bouchet, P. (2015). Longchaeus folinii (Dall, 1889). Accessed through: World Register of Marine Species at http ...
'' (Dall, 1889) * ''Pyramidella fulva'' G.B. Sowerby II, 1865: synonym of '' Colsyrnola brunnea'' (A. Adams, 1854) * ''Pyramidella fusca'' (C. B. Adams, 1839): synonym of '' Sayella fusca'' (C. B. Adams, 1839) * ''Pyramidella garrettii'' Tryon, 1886: synonym of '' Milda garretti'' (Tryon, 1886) * ''Pyramidella glans'' Reeve, 1842: synonym of '' Otopleura glans'' (Reeve, 1842) (original combination) * ''Pyramidella halaibensis'' (Sturany, 1903): synonym of '' Tropaeas halaibensis'' (Sturany, 1903) * ''Pyramidella hera'' Bartsch, 1915: synonym of '' Syrnola hera'' (Bartsch, 1915) * ''Pyramidella humilis'' Preston, 1905: synonym of '' Quirella humilis'' (Preston, 1905) * ''Pyramidella inopinata'' (Schander, 1994): synonym of '' Longchaeus inopinatus'' (Schander, 1994) * ''Pyramidella insularum'' Pilsbry, 1922: synonym of '' Longchaeus insularum'' (Pilsbry, 1922) * † ''Pyramidella jamaicensis'' Dall, 1896: synonym of † '' Longchaeus jamaicensis'' (Dall, 1896) * † ''Pyramidella kelirensis'' (K. Martin, 1916): synonym of † ''
Longchaeus menadensis ''Longchaeus'' is a genus of sea snails, marine gastropod mollusks in the family Pyramidellidae, the pyrams and their allies.WoRMS (2011). ''Longchaeus'' Mörch, 1875. Accessed through: World Register of Marine Species at http://www.marinespeci ...
'' (Schepman, 1907) * † ''Pyramidella laeviuscula'' S. V. Wood, 1842: synonym of † ''
Longchaeus plicosus ''Longchaeus'' is a genus of sea snails, marine gastropod mollusks in the family Pyramidellidae, the pyrams and their allies.WoRMS (2011). ''Longchaeus'' Mörch, 1875. Accessed through: World Register of Marine Species at http://www.marinespeci ...
'' (Bronn, 1838) * ''Pyramidella lischkei'' Dall & Bartsch, 1906: synonym of '' Styloptygma taeniatum'' (A. Adams, 1863) * ''Pyramidella lissa'' (A. E. Verrill, 1884): synonym of '' Eulimella lissa'' A. E. Verrill, 1884 * ''Pyramidella maculosa'' Lamarck, 1822: synonym of ''
Longchaeus maculosus ''Longchaeus maculosus'', common name the sulcate pyram, is a species of sea snail, a marine (ocean), marine gastropod mollusk in the family (biology), family Pyramidellidae, the pyrams and their allies.MolluscaBase eds. (2022). MolluscaBase. Lon ...
'' (Lamarck, 1822) * ''Pyramidella magnifica'' Adams & Reeve, 1850: synonym of '' Otopleura auriscati'' (Holten, 1802) * ''Pyramidella mazatlanica'' Dall & Bartsch, 1909: synonym of ''
Longchaeus mazatlanicus ''Longchaeus mazatlanicus'' is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.MolluscaBase eds. (2022). MolluscaBase. Longchaeus mazatlanicus (Dall & Bartsch, 1909). Accessed through: ...
'' (Dall & Bartsch, 1909) * ''Pyramidella mediterranea'' Monterosato, 1880: synonym of '' Tiberia minuscula'' (Monterosato, 1880) (dubious synonym) * ''Pyramidella minuscula'' Monterosato, 1880: synonym of '' Tiberia minuscula'' (Monterosato, 1880) * ''Pyramidella mitralis'' A. Adams, 1853: synonym of '' Otopleura mitralis'' (A. Adams, 1855) * ''Pyramidella modica'' (A. Adams, 1860): synonym of '' Syrnola modica''(A. Adams, 1860) * ''Pyramidella nitida'' (A. Adams, 1854): synonym of '' Otopleura nitida'' (A. Adams, 1854) * ''Pyramidella nitidula'' (A Adams, 1860): synonym of '' Tiberia nitidula'' (A. Adams, 1860) * ''Pyramidella nivea'' (Mørch, 1875): synonym of '' Triptychus niveus'' (Mørch, 1875) * ''Pyramidella norna'' Bartsch, 1915: synonym of '' Leucotina elongata'' G. B. Sowerby III, 1892 * † ''Pyramidella obtusior'' Sacco, 1892: synonym of † ''
Longchaeus obtusatus ''Longchaeus'' is a genus of sea snails, marine gastropod mollusks in the family Pyramidellidae, the pyrams and their allies.WoRMS (2011). ''Longchaeus'' Mörch, 1875. Accessed through: World Register of Marine Species at http://www.marinespeci ...
'' (O. Semper, 1861) † (unjustified emendation of obtusatus) * ''Pyramidella octaviana'' Di Geronimo, 1973: synonym of '' Tiberia minuscula'' (Monterosato, 1880) * † ''Pyramidella olssoni'' Maury, 1917: synonym of † ''
Longchaeus olssoni ''Longchaeus'' is a genus of sea snails, marine gastropod mollusks in the family Pyramidellidae, the pyrams and their allies.WoRMS (2011). ''Longchaeus'' Mörch, 1875. Accessed through: World Register of Marine Species at http://www.marinespeci ...
'' (Maury, 1917) * ''Pyramidella ornata'' (Gould, 1861): synonym of '' Colsyrnola ornata'' (Gould, 1861) * ''Pyramidella plicata'' Lamarck, 1816: synonym of '' Otopleura auriscati'' (Holten, H.S., 1802) ( junior subjective synonym) * ''Pyramidella pratii'' Bernardi, 1858: synonym of ''
Longchaeus turritus ''Longchaeus turritus'' is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.Bouchet, P. (2015). Longchaeus turritus. Accessed through: World Register of Marine Species at http://marines ...
'' (A. Adams, 1854) * ''Pyramidella producta'' (C. B. Adams, 1840): synonym of ''
Syrnola producta ''Syrnola producta'' is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.Rosenberg, G. (2011). ''Syrnola producta'' (C. B. Adams, 1840). Accessed through: World Register of Marine Spe ...
'' (C. B. Adams, 1840) * ''Pyramidella propinqua'' A. Adams, 1854: synonym of '' Otopleura mitralis'' (A. Adams, 1854) * ''Pyramidella pulchella'' (A. Adams, 1854): synonym of '' Orinella pulchella'' (A. Adams, 1854) * ''Pyramidella punctata'' Schubert & Wagner, 1829: synonym of '' Longchaeus acus'' (Gmelin, 1791) (invalid: homonym and synonym of ''Pyramidella punctata'' Férussac, 1821) * ''Pyramidella punctata'' Férussac, 1821: synonym of '' Longchaeus acus'' (Gmelin, 1791) * ''Pyramidella pupaeformis'' Souverbie, 1865: synonym of '' Pyrgulina pupaeformis'' (Souverbie, 1865) * ''Pyramidella resticula'' (Dall, 1889): synonym of '' Bacteridium resticulum'' (Dall, 1889) * ''Pyramidella schanderi'' van Aartsen, Gittenberger & Goud, 1998: synonym of '' Pyramidella inopinata'' (Schander, 1994) * † ''Pyramidella semicanaliculata'' Maury, 1917: synonym of ''
Longchaeus suturalis ''Longchaeus suturalis'', common name the crenulated pyram, is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.WoRMS (2010). ''Longchaeus suturalis'' (H. C. Lea, 1843). Accessed throu ...
'' (H. C. Lea, 1843) * ''Pyramidella similis'' Thiele, 1925: synonym of '' Tiberia analoga'' van Aartsen & Corgan, 1996 * ''Pyramidella smithi'' Verrill, 1881: synonym of '' Eulimella unifasciata'' (Forbes, 1844) * ''Pyramidella somaliensis'' Thiele, 1925: synonym of '' Tiberia somaliensis'' (Thiele, 1925) (original combination) * ''Pyramidella sulcata'' (A. Adams, 1854): synonym of ''Pyramidella maculosa'' Lamarck, 1822: synonym of ''
Longchaeus maculosus ''Longchaeus maculosus'', common name the sulcate pyram, is a species of sea snail, a marine (ocean), marine gastropod mollusk in the family (biology), family Pyramidellidae, the pyrams and their allies.MolluscaBase eds. (2022). MolluscaBase. Lon ...
'' (Lamarck, 1822) * ''Pyramidella suturalis'' H. C. Lea, 1843: synonym of ''
Longchaeus suturalis ''Longchaeus suturalis'', common name the crenulated pyram, is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.WoRMS (2010). ''Longchaeus suturalis'' (H. C. Lea, 1843). Accessed throu ...
'' (H. C. Lea, 1843) * ''Pyramidella terebellum'' (O.F. Muller, 1774): synonym of ''Pyramidella dolabrata terebellum'' (Müller, 1774): synonym of ''Pyramidella dolabrata'' (Linnaeus, 1758) * ''Pyramidella teres'' (A. Adams, 1854): synonym of ''Tropaeas teres'' (A. Adams, 1861): synonym of ''
Longchaeus turritus ''Longchaeus turritus'' is a species of sea snail, a marine gastropod mollusk in the family Pyramidellidae, the pyrams and their allies.Bouchet, P. (2015). Longchaeus turritus. Accessed through: World Register of Marine Species at http://marines ...
'' (A. Adams, 1854) * ''Pyramidella tessellata'' (A. Adams, 1854): synonym of ''Pyramidella sulcata'' (A. Adams, 1854): synonym of ''Pyramidella maculosa'' Lamarck, 1822: synonym of ''
Longchaeus maculosus ''Longchaeus maculosus'', common name the sulcate pyram, is a species of sea snail, a marine (ocean), marine gastropod mollusk in the family (biology), family Pyramidellidae, the pyrams and their allies.MolluscaBase eds. (2022). MolluscaBase. Lon ...
'' (Lamarck, 1822) * ''Pyramidella umbilicata'' Thiele, 1925: synonym of '' Tiberia navella'' van Aartsen & Corgan, 1996 * ''Pyramidella unifasciata'' (Forbes, 1844): synonym of '' Tibersyrnola unifasciata'' (Forbes, 1844) * ''Pyramidella variegata'' A. Adams, 1854:WoRMS (2010). ''Pyramidella variegata'' A. Adams. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=208417 on 2011-03-15 synonym of '' Otopleura mitralis'' (A. Adams, 1854) * ''Pyramidella ventricosa'' (Forbes, 1844): synonym of '' Eulimella ventricosa'' (Forbes, 1844) * ''Pyramidella ventricosa'' Guérin, 1831: synonym of '' Milda ventricosa'' (Guérin, 1831)


References

* Lamarck, J.B.P.A. de Monet de 1799. Prodrome d'une nouvelle classification de coquilles, comprenent une réaction appropiée de charactères génériques, et l'établissement d'un grand nombre de genres nouveaux. Mémoires de la Société d'Histoire Naturelle de Paris 1: 63-91 * Vaught, K.C. (1989). ''A classification of the living Mollusca''. American Malacologists: Melbourne, FL (USA). . XII, 195 pp.


External links


GBIF info
{{Taxonbar, from=Q3174924 Pyramidellidae Gastropod genera Taxa named by Jean-Baptiste Lamarck