''Pycnonemosaurus'' (meaning 'dense forest lizard') is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
theropod dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that belonged to the family
Abelisauridae. It was found in the Upper Cretaceous red conglomerate sandstones of the "Cabembe Unit",
Mato Grosso
Mato Grosso ( – lit. "Thick Bush") is one of the states of Brazil, the third largest by area, located in the Central-West region. The state has 1.66% of the Brazilian population and is responsible for 1.9% of the Brazilian GDP.
Neighboring ...
,
Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, and it lived about 70 million years ago during the Late
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
(
Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
stage).
Discovery and naming

The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Pycnonemosaurus nevesi'', was formally described by
Kellner Kellner is a surname. Notable people with the surname include:
* Ádám Kellner (born 1986), Hungarian tennis player
* Alex Kellner (1924–1996), baseball pitcher
* Alexander Kellner (born 1961), Brazilian paleontologist
* Birgit Kellner, Austria ...
and Campos in 2002.
The only known specimen was found in a red conglomeratic sandstone at the Fazenda Roncador locality, in Mato Grosso
Mato Grosso ( – lit. "Thick Bush") is one of the states of Brazil, the third largest by area, located in the Central-West region. The state has 1.66% of the Brazilian population and is responsible for 1.9% of the Brazilian GDP.
Neighboring ...
State that is exposed close to Paulo Creek, which is referred to an unidentified formation of the Upper Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
Bauru Group. During 1952-1953, Llewellyn Ivor Price visited a farm named "Roncador" in the state of Mato Grosso and collected several dinosaur bones. These remains were found by the owner of the farm, Max de Barros Erhart, and his hired workers at the Paulo Creek site. The most important specimen found was the incomplete skeleton of a large abelisaurid theropod, which was found near several titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...
id sauropod bones.[
''Pycnonemosaurus nevesi'' was named from the Greek word ''pycnós'' meaning dense, ''némos'' meaning pastures and woods, and ''saûrus'' meaning reptile or lizard. This naming was an allusion to Mato Grosso State, where the remains were found. The specific name was named after the late Dr. Iedo Batista Neves, who passed in 2000, who encouraged the pursuit of paleontological studies, particularly of Alexander Kellner.][
]
Description
Thus far, the remains of ''Pycnonemosaurus'' have been fragmentary. No elements were well preserved, and the bone surface is well abraded that indicates the elements were partially exposed at the discovery location before being collected.[ The type specimen (DGM 859-R), housed at the ]Earth Sciences Museum
The Earth Sciences Museum (''Museu de Ciências da Terra'') is a geological museum in Rio de Janeiro, Brazil. The building was part of the National Exposition of Brazil in 1908. Its collection includes minerals, fossils, and geological exhibits. ...
, Rio de Janeiro
Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the state of the same name, Brazil's third-most populous state, and the second-most populous city in Brazil, after São Paulo. Listed by the GaWC as a b ...
, consists of five incomplete teeth, parts of seven caudal
Caudal may refer to:
Anatomy
* Caudal (anatomical term) (from Latin ''cauda''; tail), used to describe how close something is to the trailing end of an organism
* Caudal artery, the portion of the dorsal aorta of a vertebrate that passes into the ...
vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e, the distal part of a right pubis, a right tibia
The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
, and the distal articulation of the right fibula
The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity is ...
. The small pubic foot and hatchet-shaped cnemial crest of the tibia distinguishes this species within the abelisaurs. The caudal vertebra has distinct abelisaurid features, such as a fan-shaped transverse process and a cranial projection. However, these awl-like projections are somewhat unlike related abelisaurids, such as ''Aucasaurus
''Aucasaurus'' is a genus of medium-sized abelisaurid theropod dinosaur from Argentina that lived during the Late Cretaceous (Santonian to Campanian stage) of the Anacleto Formation. It was smaller than the related ''Carnotaurus'', although more ...
'', in that they diminish more towards the distal caudals. All remains were found associated and are presently regarded as belonging to the same individual.[
Initial size estimates put the animal at in length and 1.2 tonnes in weight (1.3 short tons),][Paul, G.S., 2010., ''The Princeton Field Guide to Dinosaurs''. Princeton University Press. p. 79] but later analyses have found that it was likely larger, being about long. This new size estimate currently makes ''Pycnonemosaurus'' the largest formally described member of the Abelisauridae thus far.
Phylogeny
The following cladogram follows an analysis of 2021 during the description of ''Llukalkan
''Llukalkan'' (Mapuche for "one who causes fear") is a genus of abelisaurid theropod dinosaur from the Late Cretaceous Bajo de la Carpa Formation of Argentina. The type species is ''Llukalkan aliocranianus''.
Discovery and naming
The holotype ...
''.
Paleoecology
''Pycnonemosaurus'' is the best known abelisaurid from Brazil, where most theropod material is presently rare besides preserved teeth and footprints. Even though only a few species are known from Brazil, it is one of the most informative countries concerning the Lower Cretaceous
Lower may refer to:
*Lower (surname)
*Lower Township, New Jersey
*Lower Receiver (firearms)
*Lower Wick
Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
period. ''Pycnonemosaurus'' was a predator, preying on mid-sized sauropodomorphs, small crocodylomorphs
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction.
During Mesozoic and early Cenozoic times, cr ...
, and dromaeosaurs. Its teeth were small yet sharp, and were used to get a hold of struggling prey as the abelisaur shook and tore them apart.[
]
See also
* Timeline of ceratosaur research
This timeline of ceratosaur research is a chronological listing of events in the history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became the apex predators of ...
References
External links
*
''Pycnonemosaurus'' at Dinosaurier Info
{{Taxonbar, from=Q139189
Brachyrostrans
Late Cretaceous dinosaurs of South America
Cretaceous Brazil
Fossils of Brazil
Fossil taxa described in 2002
Taxa named by Alexander Kellner