Pycnanthus Angolensis
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''Pycnanthus angolensis'' is a species of
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
in the
nutmeg Nutmeg is the seed, or the ground spice derived from the seed, of several tree species of the genus '' Myristica''; fragrant nutmeg or true nutmeg ('' M. fragrans'') is a dark-leaved evergreen tree cultivated for two spices derived from its fru ...
family,
Myristicaceae The Myristicaceae are a family of flowering plants native to Africa, Asia, Pacific islands, and the Americas and has been recognized by most taxonomists. It is sometimes called the "nutmeg family", after its most famous member, '' Myristica fragr ...
. It is native to
Tropical Africa The Afrotropical realm is one of the Earth's eight biogeographic realms. It includes Sub-Saharan Africa, the southern Arabian Peninsula, the island of Madagascar, and the islands of the western Indian Ocean. It was formerly known as the Ethiopi ...
. Its English language common names include African nutmeg, false nutmeg, boxboard, and cardboard.''Pycnanthus angolensis''.
AgroForestryTree Database. World Agroforestry Centre.
In Africa it is widely known as ''ilomba''.Richter, H. G. and M. J. Dallwitz. 2000 onwards

Commercial timbers: descriptions, illustrations, identification, and information retrieval. DELTA – DEscription Language for TAxonomy. June, 2009.


Description

This
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
tree grows up to 40 meters tall and usually up to a meter wide, sometimes up to 1.5 meters or more. The
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure, and the stem of woody plants * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Comput ...
is straight and cylindrical with fissures and flaking
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
. The
sap Sap is a fluid transported in the xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a s ...
is honey-colored and turns red in time. The branches are in whorls. The leathery leaves are up to 31 centimeters long by 9 wide. The blades have pointed tips, heart-shaped bases, and thick midribs. They are hairless on top and coated with
rusty Rusty may refer to something covered with rust or with a rust (color). Rusty is also a nickname for people who have red hair, have a rust-hued skin tone, or have the given name Russell (given name), Russell. Rusty may also refer to: People *Rust ...
,
felt Felt is a textile that is produced by matting, condensing, and pressing fibers together. Felt can be made of natural fibers such as wool or animal fur, or from synthetic fibers such as petroleum-based acrylic fiber, acrylic or acrylonitrile or ...
like hairs on the undersides. The leaves usually bear signs of insect damage, a feature so common it is considered characteristic of the species. The flowers are arranged in dense, rusty
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
s up to 15 centimeters long. The individual flowers are difficult to see in the tight panicle until the
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s develop, being only about a millimeter long. The flowers are hairy and fragrant.''Pycnanthus angolensis'' (Welw.) Warb.
In: Stannard, B. L. ''Flora Zambesiaca'' Vol. 9, Part 2: Myristicaceae. 1997. Royal Botanic Gardens, Kew.
The fruit is a rounded
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
reaching over 3 centimeters long and wide, borne in clusters. It is hairy brown when new, turning yellow-orange, and has cartilaginous flesh that dries woody. It contains a black seed with a red
aril An aril (), also called arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode, or false aril, is sometimes distinguished: whereas an aril grows from the attachment point of the seed to the ova ...
which resembles that of
nutmeg Nutmeg is the seed, or the ground spice derived from the seed, of several tree species of the genus '' Myristica''; fragrant nutmeg or true nutmeg ('' M. fragrans'') is a dark-leaved evergreen tree cultivated for two spices derived from its fru ...
.''Pycnanthus angolensis''.
PROTA.
The fruit ripens over a long period continuing into the next flowering season, which begins around October.


Ecology

The tree grows in moist
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s up to about 1200 meters in elevation among other evergreens and semideciduous trees. It occurs in
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
, sometimes taking hold in new canopy gaps or clearings. It thrives in sunny locations. It grows in
riparian A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripar ...
forests such as
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
s, and in some regions it can be found in
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s. It occurs in areas receiving between about 1300 and 1800 millimeters of rain per year, but the optimal seems to be about 2000 millimeters. Many animals feed on the fruits, such as
hornbill Hornbills are birds found in tropical and subtropical Africa, Asia and Melanesia of the family Bucerotidae. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a horny casque on the upper ...
s.''Pycnanthus angolensis''.
An Introduction to the Trees from the North of the Republic of Congo. Royal Botanic Garden, Edinburgh.


Uses

This species has a wide variety of human uses. It is harvested for its
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
, which is light, soft, and whitish gray or pink-tinged in color. Its popularity rose after
World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
when
plywood Plywood is a composite material manufactured from thin layers, or "plies", of wood veneer that have been stacked and glued together. It is an engineered wood from the family of manufactured boards, which include plywood, medium-density fibreboa ...
was in demand, and during the mid-20th century it was one of the more valuable timbers in Central Africa. It is not very durable and tends to
warp Warp, warped or warping may refer to: Arts and entertainment Books and comics * WaRP Graphics, an alternative comics publisher * ''Warp'' (First Comics), comic book series published by First Comics based on the play ''Warp!'' * Warp (comics), a D ...
, but it is easy to cut and work and can be used for many purposes. It is suitable for
furniture Furniture refers to objects intended to support various human activities such as seating (e.g., Stool (seat), stools, chairs, and sofas), eating (table (furniture), tables), storing items, working, and sleeping (e.g., beds and hammocks). Furnitur ...
, and in house construction as
panelling Panelling (or paneling in the United States) is a millwork wall covering constructed from rigid or semi-rigid components. These are traditionally interlocking wood, but could be plastic or other materials. Panelling was developed in antiquity ...
,
siding Siding may refer to: * Siding (construction), the outer covering or cladding of a house * Siding (rail) In rail terminology, a siding is a low-speed track section distinct from a running line or through route such as a main line, branch lin ...
,
roof shingle A roof’s shingles are a roof covering consisting of individual overlapping elements. These elements are typically flat, rectangular shapes laid in courses from the bottom edge of the roof up, with each successive course overlapping the joint ...
s, and framing. It is used for
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work (physics), work. The concept was originally applied solely to those materials capable of releasing chem ...
and
paper pulp Pulp is a fibrous lignocellulosic material prepared by chemically, semi-chemically, or mechanically isolating the cellulosic fibers of wood, fiber crops, waste paper, or rags. Mixed with water and other chemicals or plant-based additives, pul ...
. The wood can be vulnerable to
termites Termites are a group of detritophagous eusocial cockroaches which consume a variety of decaying plant material, generally in the form of wood, leaf litter, and soil humus. They are distinguished by their moniliform antennae and the sof ...
,
powderpost beetle Powderpost beetles are a group of seventy species of woodboring beetles classified in the insect subfamily Lyctinae. These beetles, along with spider beetles, death watch beetles, common furniture beetles, skin beetles, and others, make u ...
s, and other
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
. The yellowish or reddish
fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers specif ...
from the aromatic seed is called "kombo butter" or "Angola tallow". A seed can be up to 70% fat. It is used as a
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work (physics), work. The concept was originally applied solely to those materials capable of releasing chem ...
for lighting and is made into
soap Soap is a salt (chemistry), salt of a fatty acid (sometimes other carboxylic acids) used for cleaning and lubricating products as well as other applications. In a domestic setting, soaps, specifically "toilet soaps", are surfactants usually u ...
. Seed remnants are used in
compost Compost is a mixture of ingredients used as plant fertilizer and to improve soil's physical, chemical, and biological properties. It is commonly prepared by Decomposition, decomposing plant and food waste, recycling organic materials, and man ...
. When ignited, the oily seed burns slowly and can be used as a
candle A candle is an ignitable candle wick, wick embedded in wax, or another flammable solid substance such as tallow, that provides light, and in some cases, a Aroma compound, fragrance. A candle can also provide heat or a method of keeping time. ...
. In
Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
the tree is grown in
banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large treelike herbaceous flowering plants in the genus '' Musa''. In some countries, cooking bananas are called plantains, distinguishing the ...
,
coffee Coffee is a beverage brewed from roasted, ground coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content, but decaffeinated coffee is also commercially a ...
, and
cocoa Cocoa may refer to: Chocolate * Chocolate * ''Theobroma cacao'', the cocoa tree * Cocoa bean, seed of ''Theobroma cacao'' * Chocolate liquor, or cocoa liquor, pure, liquid chocolate extracted from the cocoa bean, including both cocoa butter and ...
plantations to shade the crops. Most parts of the tree have been used in
traditional African medicine Traditional African medicine is a range of traditional medicine disciplines involving indigenous herbalism and African spirituality, typically including diviners, midwives, and herbalists. Practitioners of traditional African medicine claim, ...
. The sap has been used to control bleeding. The bark has been used as a poison
antidote An antidote is a substance that can counteract a form of poisoning. The term ultimately derives from the Greek term φάρμακον ἀντίδοτον ''(pharmakon antidoton)'', "(medicine) given as a remedy". An older term in English which is ...
and a treatment for
leprosy Leprosy, also known as Hansen's disease (HD), is a Chronic condition, long-term infection by the bacteria ''Mycobacterium leprae'' or ''Mycobacterium lepromatosis''. Infection can lead to damage of the Peripheral nervous system, nerves, respir ...
,
anemia Anemia (also spelt anaemia in British English) is a blood disorder in which the blood has a reduced ability to carry oxygen. This can be due to a lower than normal number of red blood cells, a reduction in the amount of hemoglobin availabl ...
,
infertility In biology, infertility is the inability of a male and female organism to Sexual reproduction, reproduce. It is usually not the natural state of a healthy organism that has reached sexual maturity, so children who have not undergone puberty, whi ...
,
gonorrhea Gonorrhoea or gonorrhea, colloquially known as the clap, is a sexually transmitted infection (STI) caused by the bacterium ''Neisseria gonorrhoeae''. Infection may involve the genitals, mouth, or rectum. Gonorrhea is spread through sexual c ...
, and
malaria Malaria is a Mosquito-borne disease, mosquito-borne infectious disease that affects vertebrates and ''Anopheles'' mosquitoes. Human malaria causes Signs and symptoms, symptoms that typically include fever, Fatigue (medical), fatigue, vomitin ...
. Leaf extracts are consumed or used in an
enema An enema, also known as a clyster, is the rectal administration of a fluid by injection into the Large intestine, lower bowel via the anus.Cullingworth, ''A Manual of Nursing, Medical and Surgical'':155 The word ''enema'' can also refer to the ...
to treat
edema Edema (American English), also spelled oedema (British English), and also known as fluid retention, swelling, dropsy and hydropsy, is the build-up of fluid in the body's tissue (biology), tissue. Most commonly, the legs or arms are affected. S ...
. Root extracts are used to treat parasitic infections, such as
schistosomiasis Schistosomiasis, also known as snail fever, bilharzia, and Katayama fever is a neglected tropical helminthiasis, disease caused by parasitism, parasitic Schistosoma, flatworms called schistosomes. It affects both humans and animals. It affects ...
. The seed oil is used to treat
thrush Thrush may refer to: Birds * Thrush (bird), any of the birds in the family Turdidae ** List of thrush species * Antthrushes, the Formicariidae family of birds * Dohrn's warbler, or Dohrn's thrush-babbler, a species ''Sylvia dohrni'' in the famil ...
. Like the fat of nutmeg, kombo butter is mostly
myristic acid Myristic acid (IUPAC name: tetradecanoic acid) is a common saturated fatty acid with the molecular formula . Its salts and esters are commonly referred to as myristates or tetradecanoates. The name of the acyl group derived from myristic acid is m ...
, with a high amount of myristoleic acid, as well. It contains the unique compound kombic acid, which was named for the tree under its nomenclatural synonym, ''P. kombo''.


Cultivation

This tree is cultivated for its products. Seeds are sown in the field, and seedlings are grown in
plant nurseries A nursery is a place where plants are propagated and grown to a desired size. Mostly the plants concerned are for gardening, forestry, or conservation biology, rather than agriculture. They include retail nurseries, which sell to the general ...
until they have large
taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproot ...
s. They cannot tolerate
drought A drought is a period of drier-than-normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, ...
. After a year the tree is about half a meter tall, and within 4 years it can reach 4 meters. A 20-year-old tree can be 25 meters tall. Stands of cultivated trees are pruned and thinned periodically. Pest insects such as '' Monochamus scabiosus'' and '' Mallodon downesi'' and
fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
such as '' Ophiostoma'' sp. have been observed, but they are not severe problems. The freshly harvested wood may become discolored due to
bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
l growth, the light-colored wood developing dark brown staining. Trees can be cut when they reach a diameter of 50 centimeters, at around 30 years old.
Agricultural cooperative An agricultural cooperative, also known as a farmers' co-op, is a producer cooperative in which farmers pool their resources in certain areas of activities. A broad typology of agricultural cooperatives distinguishes between agricultural servic ...
s in Ghana cultivate the trees for the kombo butter.


Common names

There are
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
s for the plant in many languages. It is called ''mkungu mwitu'' in
Swahili Swahili may refer to: * Swahili language, a Bantu language officially used in Tanzania, Kenya and Uganda and widely spoken in the African Great Lakes. * Swahili people, an ethnic group in East Africa. * Swahili culture, the culture of the Swahili p ...
, ''akwa-mili'' and ''oje'' in
Igbo Igbo may refer to: * Igbo people, an ethnic group of Nigeria * Igbo language, their language * anything related to Igboland, a cultural region in Nigeria See also * Ibo (disambiguation) * Igbo mythology * Igbo music * Igbo art * * Igbo-Ukwu, a t ...
, ''lunaba'' and ''munaba'' in
Luganda Ganda or Luganda ( ; ) is a Bantu language spoken in the African Great Lakes region. It is one of the major languages in Uganda and is spoken by more than 5.56 million Ganda people, Baganda and other people principally in central Uganda, includ ...
, ''akomu'' in Yoruba, ''calabo'' in Spanish, and ''arbre à suif'' and ''faux muscadier'' in French. Local names include ''pó casson'' in
São Tomé and Príncipe São Tomé and Príncipe, officially the Democratic Republic of São Tomé and Príncipe, is an island country in the Gulf of Guinea, off the western equatorial coast of Central Africa. It consists of two archipelagos around the two main isla ...
, ''gboyei'' in
Sierra Leone Sierra Leone, officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered to the southeast by Liberia and by Guinea to the north. Sierra Leone's land area is . It has a tropical climate and envi ...
and
Liberia Liberia, officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to Guinea–Liberia border, its north, Ivory Coast to Ivory Coast–Lib ...
, ''otie'' in
Ghana Ghana, officially the Republic of Ghana, is a country in West Africa. It is situated along the Gulf of Guinea and the Atlantic Ocean to the south, and shares borders with Côte d’Ivoire to the west, Burkina Faso to the north, and Togo to t ...
, ''eteng'' in
Cameroon Cameroon, officially the Republic of Cameroon, is a country in Central Africa. It shares boundaries with Nigeria to the west and north, Chad to the northeast, the Central African Republic to the east, and Equatorial Guinea, Gabon, and the R ...
, ''lolako'' in
Zaire Zaire, officially the Republic of Zaire, was the name of the Democratic Republic of the Congo from 1971 to 18 May 1997. Located in Central Africa, it was, by area, the third-largest country in Africa after Sudan and Algeria, and the 11th-la ...
,''Pycnanthus angolensis''.
Wood Technical Fact Sheet. Forest Products Laboratory. Forest Service, USDA.
and ''adria'', ''effoi'', ''hétéré'', ''qualélé'', and ''walélé'' in
Ivory Coast Ivory Coast, also known as Côte d'Ivoire and officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital city of Yamoussoukro is located in the centre of the country, while its largest List of ci ...
.


References

{{Taxonbar, from=Q6093520 Myristicaceae Trees of Africa Plants used in traditional African medicine Wood