Punpun Foragers
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West African hunter-gatherers, West African foragers, or West African pygmies dwelled in western
Central Africa Central Africa (French language, French: ''Afrique centrale''; Spanish language, Spanish: ''África central''; Portuguese language, Portuguese: ''África Central'') is a subregion of the African continent comprising various countries accordin ...
earlier than 32,000 BP and dwelled in
West Africa West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations geoscheme for Africa#Western Africa, United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Gha ...
between 16,000 BP and 12,000 BP until as late as 1000 BP or some period of time after 1500 CE. West African hunter-gatherers are
archaeologically Archaeology or archeology is the study of human activity through the recovery and analysis of material culture. The archaeological record consists of artifacts, architecture, biofacts or ecofacts, sites, and cultural landscapes. Archaeology ...
associated with the West African Microlithic Technocomplex. Despite its significance in the prehistory of West Africa, the peopling of various parts of Western Africa from the
Sub-Saharan Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...
regions of
coastal A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...
West Africa and the
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
of western Central Africa often goes overlooked. Prior to West African hunter-gatherers, there may have been various peoples (e.g., Iwo Eleru people, possibly Aterians) who continuously occupied West Africa amid the
Middle Stone Age The Middle Stone Age (or MSA) was a period of African prehistory between the Early Stone Age and the Late Stone Age. It is generally considered to have begun around 280,000 years ago and ended around 50–25,000 years ago. The beginnings of ...
. Macrolith-using late Middle Stone Age peoples (e.g., the possibly archaic human admixed or late-persisting
early modern human Early modern human (EMH), or anatomically modern human (AMH), are terms used to distinguish ''Homo sapiens'' (Homo sapiens sapiens, sometimes ''Homo sapiens sapiens'') that are Human anatomy, anatomically consistent with the Human variability, r ...
Iwo Eleru fossils of the late Middle Stone Age), who dwelled in Central Africa, western Central Africa, and West Africa, were displaced by microlith-using Late Stone Age Africans (e.g., non-archaic human admixed Late Stone Age Shum Laka fossils dated between 7000 BP and 3000 BP) as they migrated from Central Africa, to western Central Africa, into West Africa. Earlier than 32,000 BP, or by 30,000 BP, Late Stone Age West African hunter-gatherers were dwelling in the
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
of western Central Africa (e.g., earlier than 32,000 BP at de Maret in
Shum Laka The archaeological site of Shum Laka is the most prominent rockshelter site in the Grasslands region of the Laka Valley, northwest Cameroon. Occupations at this rockshelter date to the Later Stone Age. This region is important to investigations ...
, 12,000 BP at Mbi Crater). Between 16,000 BP and 12,000 BP, Late Stone Age West Africans began dwelling in the eastern and central forested regions (e.g., Ghana, Ivory Coast, Nigeria; between 18,000 BP and 13,000 BP at Temet West and Asokrochona in the southern region of Ghana, 13,050 ± 230 BP at
Bingerville Bingerville is a town in south-eastern Ivory Coast. It is a suburb of Abidjan city and one of four sub-prefectures of Abidjan Autonomous District. Bingerville is also a commune. The town is located about 10 kilometres east of Abidjan and lies ...
in the southern region of Ivory Coast, 11,200 ± 200 BP at Iwo Eleru in Nigeria) of West Africa. West African hunter-gatherers resided at the Nigerian sites of Iwo Eleru and Rop, at the Ivorian site of Bingerville, at the Cameroonian site of Shum Laka, at the Malian site of
Ounjougou Ounjougou is the name of a lieu-dit found in the middle of an important complex of archaeological sites in the Upper Yamé Valley on the Bandiagara Plateau, in Dogon Country, Mali. The Ounjougou archaeological complex consists of over a hundred ...
, and at the Senegalese sites of Fatandi and Toumboura. Prior to the Holocene era, interaction between West Africans migrating from the
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
and West African hunter-gatherers of the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
and
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
regions were limited, as evidenced by West African hunter-gatherer microlithic cultural continuity. West African hunter-gatherers likely were the sole occupants of the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
and
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
regions of West Africa. Unlike Central African hunter-gatherers, who dwell in more secluded areas in the
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
of
Central Africa Central Africa (French language, French: ''Afrique centrale''; Spanish language, Spanish: ''África central''; Portuguese language, Portuguese: ''África Central'') is a subregion of the African continent comprising various countries accordin ...
, West African hunter-gatherers likely dwelt in more open areas of West Africa. Migration of Saharan peoples south of the Sahelian region resulted in seasonal interaction and gradual absorption of West African hunter-gatherers, who primarily dwelt in the
savannas A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient li ...
and
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
of
West Africa West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations geoscheme for Africa#Western Africa, United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Gha ...
. After having persisted as late as 1000 BP, or some period of time after 1500 CE, remaining West African hunter-gatherers, many of whom dwelt in the forest-savanna region, were ultimately acculturated and admixed into larger groups of West African agriculturalists, akin to the migratory Bantu-speaking agriculturalists and their encounters with Central African hunter-gatherers.


Prehistory


Pleistocene

Middle Stone Age West Africans may have dwelled at
Ounjougou Ounjougou is the name of a lieu-dit found in the middle of an important complex of archaeological sites in the Upper Yamé Valley on the Bandiagara Plateau, in Dogon Country, Mali. The Ounjougou archaeological complex consists of over a hundred ...
, Mali (71,000 BP – 59,000 BP, 59,000 BP – 28,000 BP), Faleme Valley, Senegal (Late MIS 5), Tiemassas, Senegal (62,000 BP – 25,000 BP), Birimi, Ghana (50,000 BP – 20,000 BP), Missira ( MIS 4), Toumboura, Senegal (33,000 BP), Laminia, Gambia (24,000 BP – 21,000 BP), Ndiayène Pendao, Senegal (11,600 BP), and Saxonomunya (11,000 BP), near
Falémé Falémé is a commune in the Cercle of Kayes in the Kayes Region of south-western Mali. The main town (''chef-lieu An administrative centre is a seat of regional administration or local government, or a county town, or the place where the ce ...
, Mali. There is also scant evidence of Middle Stone Age dwelling at Ounjougou, Mali between 191,000 BP – 130,000 BP. Aside the scant evidence, Middle Stone Age West Africans likely dwelled continuously in West Africa between MIS 4 and MIS 2, and likely were not present in West Africa before MIS 5. Amid MIS 5, Middle Stone Age West Africans may have migrated across the West Sudanian savanna and continued to reside in the region (e.g.,
West Sudanian savanna The West Sudanian savanna is a tropical savanna ecoregion that extends across West Africa. Geography The ecoregion stretches east and west across West Africa, from the Atlantic coast of Senegal to the Mandara Mountains on Nigeria's eastern bor ...
, West African
Sahel The Sahel region (; ), or Sahelian acacia savanna, is a Biogeography, biogeographical region in Africa. It is the Ecotone, transition zone between the more humid Sudanian savannas to its south and the drier Sahara to the north. The Sahel has a ...
). In the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
, Middle Stone Age West Africans began to dwell along parts of the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
and
coastal A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...
region of West Africa (e.g., Tiemassas, Senegal). More specifically, by at least 61,000 BP, Middle Stone Age West Africans may have begun to migrate south of the West Sudanian savanna, and, by at least 25,000 BP, may have begun to dwell near the coast of West Africa. Amid aridification in MIS 5 and regional change of climate in MIS 4, in the
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
and the Sahel, Aterians may have migrated southward into West Africa (e.g., Baie du Levrier, Mauritania; Tiemassas, Senegal; Lower
Senegal River The Senegal River ( or "Senegal" - compound of the  Serer term "Seen" or "Sene" or "Sen" (from  Roog Seen, Supreme Deity in Serer religion) and "O Gal" (meaning "body of water")); , , , ) is a river in West Africa; much of its length mark ...
Valley). In 35,000 BP, Middle Stone Age West Africans and West African
archaic humans ''Homo'' () is a genus of great ape (family Hominidae) that emerged from the genus ''Australopithecus'' and encompasses only a single extant species, ''Homo sapiens'' (modern humans), along with a number of extinct species (collectively calle ...
are presumed to have admixed with one another, resulting in the development of the Iwo Eleru people (e.g., Iwo Eleru skull), who may have remained isolated in West Africa, and thus distinct from both contemporaneous Africans in the Sahara and from any other African populations amid the transitory period between the Pleistocene and Holocene. Earlier than 32,000 BP, or by 30,000 BP, Late Stone Age West African hunter-gatherers were living in the
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
of western Central Africa (e.g., earlier than 32,000 BP at de Maret in
Shum Laka The archaeological site of Shum Laka is the most prominent rockshelter site in the Grasslands region of the Laka Valley, northwest Cameroon. Occupations at this rockshelter date to the Later Stone Age. This region is important to investigations ...
, 12,000 BP at Mbi Crater). An excessively dry Ogolian period occurred, spanning from 20,000 BP to 12,000 BP. By 15,000 BP, the number of settlements made by Middle Stone Age West Africans decreased due to the increasingly humid conditions, expansion of the West African
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
, and greater the number of settlements made by Late Stone Age West African hunter-gatherers. Macrolith-using late Middle Stone Age peoples (e.g., the possibly archaic human admixed or late-persisting
early modern human Early modern human (EMH), or anatomically modern human (AMH), are terms used to distinguish ''Homo sapiens'' (Homo sapiens sapiens, sometimes ''Homo sapiens sapiens'') that are Human anatomy, anatomically consistent with the Human variability, r ...
Iwo Eleru fossils of the late Middle Stone Age), who dwelled in Central Africa, western Central Africa, and West Africa, were displaced by microlith-using Late Stone Age Africans (e.g., non-archaic human admixed Late Stone Age Shum Laka fossils dated between 7000 BP and 3000 BP) as they migrated from Central Africa, to western Central Africa, into West Africa. Between 16,000 BP and 12,000 BP, Late Stone Age West Africans began dwelling in the eastern and central forested regions (e.g., Ghana, Ivory Coast, Nigeria; between 18,000 BP and 13,000 BP at Temet West and Asokrochona in the southern region of Ghana, 13,050 ± 230 BP at
Bingerville Bingerville is a town in south-eastern Ivory Coast. It is a suburb of Abidjan city and one of four sub-prefectures of Abidjan Autonomous District. Bingerville is also a commune. The town is located about 10 kilometres east of Abidjan and lies ...
in the southern region of Ivory Coast, 11,200 ± 200 BP at Iwo Eleru in Nigeria) of West Africa. By 11,000 BP, the late settlement made by Middle Stone Age West Africans and earliest settlement made by Late Stone Age West African hunter-gatherers emerged in the westernmost region (e.g., Falémé Valley, Senegal) of West Africa. Middle Stone Age West Africans and Late Stone Age West African hunter-gatherers probably did not become admixed with one another and were culturally and ecologically distinct from one another.


Holocene

Between 12,000 BP and 8000 BP, West African hunter-gatherers then likely migrated from coastal West Africa, toward the north in West Africa (as far as Mali, Burkina Faso, and Mauritania), as evidenced by their
microlithic industries Stone tools have been used throughout human history but are most closely associated with prehistoric cultures and in particular those of the Stone Age. Stone tools may be made of either ground stone or knapped stone, the latter fashioned by a cr ...
(e.g., quartz, sandstone). Amid the early period of the Holocene, West African hunter-gatherers may have had Sahelian stone industries, from Senegal to Niger, which derived either from a distinct
Sub-Saharan African Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...
stone tradition, or from the
Shum Laka The archaeological site of Shum Laka is the most prominent rockshelter site in the Grasslands region of the Laka Valley, northwest Cameroon. Occupations at this rockshelter date to the Later Stone Age. This region is important to investigations ...
stone tradition of Cameroon. In the 10th millennium BCE, Niger-Congo speakers developed pyrotechnology and employed subsistence strategy at
Ounjougou Ounjougou is the name of a lieu-dit found in the middle of an important complex of archaeological sites in the Upper Yamé Valley on the Bandiagara Plateau, in Dogon Country, Mali. The Ounjougou archaeological complex consists of over a hundred ...
, Mali. Prior to 9400 BCE, Niger-Congo speakers independently created and used matured ceramic technology (e.g.,
pottery Pottery is the process and the products of forming vessels and other objects with clay and other raw materials, which are fired at high temperatures to give them a hard and durable form. The place where such wares are made by a ''potter'' is al ...
, pots) to contain and
cook Cook or The Cook may refer to: Food preparation * Cooking, the preparation of food * Cook (domestic worker), a household staff member who prepares food * Cook (profession), an individual who prepares food for consumption in the food industry * C ...
grains (e.g.,
Digitaria exilis ''Digitaria exilis'', referred to as findi or fundi in areas of Africa, such as The Gambia, with English common names white fonio, fonio millet, and hungry rice or acha rice, is a grass species. It is the most important of a diverse group of wil ...
,
pearl millet Pearl millet (''Cenchrus americanus'', commonly known as the synonym ''Pennisetum glaucum'') is the most widely grown type of millet. It has been grown in Africa and the Indian subcontinent since prehistoric times. The center of diversity, and ...
); ethnographically and historically, West African women have been the creators of pottery in most West African
ceramic A ceramic is any of the various hard, brittle, heat-resistant, and corrosion-resistant materials made by shaping and then firing an inorganic, nonmetallic material, such as clay, at a high temperature. Common examples are earthenware, porcela ...
traditions and their production of ceramics is closely associated with
creativity Creativity is the ability to form novel and valuable Idea, ideas or works using one's imagination. Products of creativity may be intangible (e.g. an idea, scientific theory, Literature, literary work, musical composition, or joke), or a physica ...
and
fertility Fertility in colloquial terms refers the ability to have offspring. In demographic contexts, fertility refers to the actual production of offspring, rather than the physical capability to reproduce, which is termed fecundity. The fertility rate ...
. Amid the tenth millennium BCE, microlith-using West Africans migrated into and dwelt in Ounjougou alongside earlier residing West Africans in Ounjougou. Among two existing cultural areas, earlier residing West Africans in Ounjougou were of a cultural area encompassing the
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
region (e.g., Tenere, Niger/Chad;
Aïr Mountains The Aïr Mountains or Aïr Massif (Air Tamajeq language, Tamajăq: ''Ayǝr''; Hausa language, Hausa: Eastern ''Azbin'', Western ''Abzin'') is a triangular massif, located in northern Niger, within the Sahara. Part of the West Sa ...
, Niger;
Acacus Mountains The Acacus Mountains or Tadrart Akakus ( / ALA-LC: ''Tadrārt Akākūs'') form a mountain range in the desert of the Ghat District in western Libya, part of the Sahara. They are situated east of the city of Ghat, Libya, and stretch north from the ...
, Libya/Algeria; Tagalagal, Niger; Temet, Niger) of
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
and microlith-using West Africans were of a cultural area encompassing the forest region of West Africa. Following northward expansion from coastal West Africa refugia, West African hunter-gatherers arrived and began dwelling at Korounkorokale, in Pays Mande, Mali, where they engaged in hunting and fishing. By 4000 BCE, red
ocher Ochre ( ; , ), iron ochre, or ocher in American English, is a natural clay earth pigment, a mixture of ferric oxide and varying amounts of clay and sand. It ranges in colour from yellow to deep orange or brown. It is also the name of the col ...
, used to paint
pottery Pottery is the process and the products of forming vessels and other objects with clay and other raw materials, which are fired at high temperatures to give them a hard and durable form. The place where such wares are made by a ''potter'' is al ...
,
jewelry Jewellery (or jewelry in American English) consists of decorative items worn for personal adornment such as brooches, ring (jewellery), rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the ...
, or
pictographs A pictogram (also pictogramme, pictograph, or simply picto) is a graphical symbol that conveys meaning through its visual resemblance to a physical object. Pictograms are used in systems of writing and visual communication. A pictography is a wri ...
, was developed by West African hunter-gatherers, which may have developed as a result of interaction with populations from lake areas to the northeast. With the increased use of grinded stones, and thus, cultural development of utilizing vegetation for food, this resulted in a decreased use of stone projectiles, and thus, decreased hunting cultural practices. By 700 CE, along with Niani having been established, Korounkorokale was embedded within the Kingdom of
Kangaba Kangaba is a town, commune, and seat of the Kangaba Cercle in the Koulikoro Region of south-western Mali. Kangaba is believed to have been founded in 1050 by Mandinka tribesmen and was a vassal state of the Ghana Empire. Once known as Kaba, the ...
. West African hunter-gatherers and their ancient cultural traditions may have persisted shortly thereafter, as West African hunter-gatherers became fully acculturated, and
Malinke Maninka (also known as Malinke), or more precisely Eastern Maninka, is the name of several closely related languages and dialects of the southeastern Manding subgroup of the Mande language family (itself, possibly linked to the Niger–Congo ...
metallurgy and pottery traditions became predominant. By 4000 BP, interaction between Saharan occupants and Sub-Saharan West African hunter-gatherers increased as Saharan occupants increasingly migrated southward into Sub-Saharan West Africa. As desertification was underway, West African hunter-gatherers of the Middle Niger were likely the first to encounter southward migrating Saharan occupants. Increased interaction may have resulted in the adoption of pottery and polished stone production, which, subsequently, may have led to these cultural practices being further diffused unto other West African hunter-gatherers. Additionally,
pastoralism Pastoralism is a form of animal husbandry where domesticated animals (known as "livestock") are released onto large vegetated outdoor lands (pastures) for grazing, historically by nomadic people who moved around with their herds. The anim ...
may have been adopted by some West African hunter-gatherers. As West African hunter-gatherers of the Middle Niger became increasingly acculturated and eventually admixed into more numerous, surrounding southward migrating Saharan occupants, some West African hunter-gatherers, further south, may have continued their
hunting Hunting is the Human activity, human practice of seeking, pursuing, capturing, and killing wildlife or feral animals. The most common reasons for humans to hunt are to obtain the animal's body for meat and useful animal products (fur/hide (sk ...
-
gathering Gather, gatherer, or gathering may refer to: Anthropology and sociology *Hunter-gatherer, a person or a society whose subsistence depends on hunting and gathering of wild foods * Intensive gathering, the practice of cultivating wild plants as a s ...
and/or basic vegetable cultivation cultures. Eventually, even these socially organized West African hunter-gatherers, were likely acculturated and admixed into the more numerous, surrounding West Africans from the Sahara. Amid the middle of the Holocene, West African hunter-gatherers continued to dwell along the rivers and within the
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
of coastal West Africa. West African hunter-gatherer stone industries had little presence to the north of the
West Sudanian savanna The West Sudanian savanna is a tropical savanna ecoregion that extends across West Africa. Geography The ecoregion stretches east and west across West Africa, from the Atlantic coast of Senegal to the Mandara Mountains on Nigeria's eastern bor ...
and
Sahel The Sahel region (; ), or Sahelian acacia savanna, is a Biogeography, biogeographical region in Africa. It is the Ecotone, transition zone between the more humid Sudanian savannas to its south and the drier Sahara to the north. The Sahel has a ...
boundary, which may indicate that it served as a type of natural environmental barricade to their greatly mobile hunter-gatherer lifestyle. Increased use of ceramics among West African hunter-gatherers also occurred, as evidenced by ceramics dated to 5370 ± 100 BP in
Bosumpra Cave Bosumpra Cave is an Archaeology, archaeological site situated on Kwahu Plateau, the Kwahu plateau, which forms part of the easternmost section of the Ashanti Region, Ashanti uplands. The plateau and uplands lie just north of the Akan people, Akan ...
, Ghana and ceramics dated to 4180 ± 160 BP in Mbi Crater, Cameroon. While likely still maintaining their hunter-gatherer culture, West African hunter-gatherers may have increasingly utilized local flora (e.g.,
palm oil Palm oil is an edible vegetable oil derived from the mesocarp (reddish pulp) of the fruit of oil palms. The oil is used in food manufacturing, in beauty products, and as biofuel. Palm oil accounted for about 36% of global oils produced from o ...
,
tubers Tubers are a type of enlarged structure that plants use as storage organs for nutrients, derived from stems or roots. Tubers help plants perennate (survive winter or dry months), provide energy and nutrients, and are a means of asexual reprod ...
). Desertification of the
Green Sahara The African humid period (AHP; also known by other names) was a climate period in Africa during the late Pleistocene and Holocene geologic epochs, when northern Africa was wetter than today. The covering of much of the Sahara desert by grass ...
(4000 BP – 3500 BP) resulted in the migration of Saharan pastoralists and agropastoralists south of the Sahelian region. Consequently, seasonal interaction likely occurred between Saharan pastoralists and agropastoralists and West African hunter-gatherers, who also practiced basic agriculture via vegetable cultivation. Sites in Ghana (e.g., Ntereso, Kintampo, Daboya) provide an example of group contact in 3500 BP, as evidenced by Punpun microlithic industries that appear in close proximity to Saharan projectile points, beads, stone innovations (e.g., stone arm rings, small stone axes), and livestock. Rather than Saharan pastoralists and agropastoralists replacing West African hunter-gatherers, there apparently was a merger of groups, as at Kintampo, there was evidence of adaptation to the subsistence conditions of the forest-savanna region of West Africa. Domesticated crops (e.g.,
pearl millet Pearl millet (''Cenchrus americanus'', commonly known as the synonym ''Pennisetum glaucum'') is the most widely grown type of millet. It has been grown in Africa and the Indian subcontinent since prehistoric times. The center of diversity, and ...
,
cowpea The cowpea (''Vigna unguiculata'') is an annual herbaceous legume from the genus '' Vigna''. Its tolerance for sandy soil and low rainfall have made it an important crop in the semiarid regions across Africa and Asia. It requires very few inpu ...
, large amounts of
oil palm ''Elaeis'' () is a genus of palms, called oil palms, containing two species, native to Africa and the Americas. They are used in commercial agriculture in the production of palm oil. Description Mature palms are single-stemmed, and can gro ...
) and undomesticated flora were availed in rockshelters (e.g., B-sites, K6), near the
Guinean forest-savanna mosaic Demographics of Guinea describes the condition and overview of Guinea's peoples. Demographic topics include basic education, health, and population statistics as well as identified racial and religious affiliations. Population According to t ...
, in the southern region of central Ghana. West African agriculturalists of Kintampo and West African hunter-gatherers of Punpun were migratory peoples, who settled at the sites seasonally for various reasons (e.g., oil palm production); this is evidenced by the varied way in which flora are situated at the rockshelters. West African hunter-gatherers may have migrated southward near the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
region or scattered into smaller groups amid arid seasons. Various activities (e.g., production of local resources) occurred in partially settled areas of the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
and
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
regions. After 4500 BP, desertification may have resulted in Saharan peoples migrating toward the south. The southern parts of the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
region, near Kintampo, may have been unfit for the subsistence techniques of farming
domesticated crops Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of resou ...
(e.g., pearl millet) from the region of northern Africa. As a result, subsistence techniques were adapted to the natural environment of the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
region, and local crops (e.g., oil palm, yams), may have been introduced into what was usually farmed. Successful adaptation to the local ecology seems to have occurred, from the southern part of the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
region to the
coastal region A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...
of West Africa. West African agriculturalists likely formed mutual relations with the West African hunter-gatherers. As a result of these relations, West African hunter-gatherers likely provided West African agriculturalists with oil-rich and Vitamin A-rich nuts as part of their local food source. Additionally, West African agriculturalists may have acquired forest
subsistence A subsistence economy is an economy directed to basic subsistence (the provision of food, clothing and shelter) rather than to the market. Definition "Subsistence" is understood as supporting oneself and family at a minimum level. Basic subsiste ...
knowledge and strategies from West African hunter-gatherers. With exception to some parts of West Africa (e.g., Ntereso, Kintampo), prior to late first millennium BCE, West African hunter-gatherers, who were the most widely spread cultural group of socially organized populations, were likely the only group to populate the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
and
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
regions of West Africa. The expansion of West African hunter-gatherers north, toward the Sahelian region of the Middle Niger, led to interaction with populations from further north. Prior to initial encounter with migrating populations from further north, West African hunter-gatherers may have already engaged in basic agricultural production of
tubers Tubers are a type of enlarged structure that plants use as storage organs for nutrients, derived from stems or roots. Tubers help plants perennate (survive winter or dry months), provide energy and nutrients, and are a means of asexual reprod ...
as well as utilizing
Elaeis guineensis ''Elaeis guineensis'' is a species of Arecaceae, palm commonly just called oil palm but also sometimes African oil palm or macaw-fat. The first Western world, Western person to describe it and bring back seeds was the French naturalist Michel Ad ...
and
Canarium schweinfurthii ''Canarium schweinfurthii'' (commonly known as the bush candle, African olive, African elemi, Empafu, or canarium), is a species of large tree native to tropical Africa.ICRAF Names in many African languages are variations of ''mupafu''. Descrip ...
. After interaction began, some West African hunter-gatherers may have acquired knowledge of pottery and polished stone production, which then spread further southward onto other West African hunter-gatherers, while others may have acquired knowledge of pastoralism. Continued interaction may have resulted in further acculturation (e.g., loss of West African hunter-gatherer languages). Isolated groups of West African hunter-gatherers may have continually dwelled throughout the region of the Pays Mande mountains after the development of metallurgy. West African hunter-gatherers may have even adopted, culturally adapted metallurgical practices, while still maintaining their ancient stone industrial traditions. Cultural continuity, via stone industries of isolated West African hunter-gatherers from the forest-savanna region, has been found throughout West Africa as late as the end of first millennium CE. Kamabai Shelter, in Sierra Leone, had quartz microliths dated to 1190 ± 95 BP. In Mali, quartz microliths were dated to 1430 ± 80 BP in Nyamanko and dated to 1020 ± 105 BP in Korounkorokale. Kariya Wuro, in Nigeria, had quartz microliths dated to 950 ± 30 BP. After having persisted as late as the end of first millennium CE, or 1000 BP, many of the remaining West African hunter-gatherers were likely ultimately acculturated and admixed into larger groups of West African agriculturalists, akin to the migratory Bantu-speaking agriculturalists and their encounters with Central African hunter-gatherers.


History

According to early European literature of the 16th century CE, West African pygmies dwelled throughout West Africa (e.g., Cameroon, Sierra Leone, Liberia). In 1500 CE, when the Dogon people entered the Bandiagara Cliffs, they encountered West African pygmies known as the Tellem.


Oral Traditions About West African Hunter-Gatherers

Mande Mande may refer to: * Mandé peoples of western Africa * Mande languages, their Niger-Congo languages * Manding languages, Manding, a term covering a subgroup of Mande peoples, and sometimes used for one of them, Mandinka people, Mandinka * Garo p ...
(e.g., Soninke,
Malinke Maninka (also known as Malinke), or more precisely Eastern Maninka, is the name of several closely related languages and dialects of the southeastern Manding subgroup of the Mande language family (itself, possibly linked to the Niger–Congo ...
) peoples and
Dogon people The Dogon are an ethnic group indigenous to the central plateau region of Mali, in West Africa, south of the Niger bend, near the city of Bandiagara, and in Burkina Faso. The population numbers between 400,000 and 800,000. They speak the Dogo ...
had oral traditions of West African pygmies (e.g.,
Tellem The Tellem (meaning: "those who were before us" or "We found them" in the Dogon languageBedaux, Rogier M. A., ''Tellem, reconnaissance archéologique d'une culture de l'Ouest africain au Moyen Age : recherches architectoniques'', Journal de la ...
). For the Dogon, even before the Tellem, there were groups (e.g., Yeban, Andoumboulou) that were even more ancient. Water-based economic (e.g., fishing) peoples (e.g., Bozo, Sorkawa), who are reputed to be one of the Niger River's first settlers, recognized that there were even earlier settled peoples – “red men.” The oral history among numerous modern West Africans is that their ancestors were West African pygmies. Among the Sousou, in Guinea, West African pygmies were known as the Doki. Among the
Wolof Wolof or Wollof may refer to: * Wolof people, an ethnic group found in Senegal, Gambia, and Mauritania * Wolof language, a language spoken in Senegal, Gambia, and Mauritania * The Wolof or Jolof Empire, a medieval West African successor of the Mal ...
, West African pygmies were known as Kondrong, who lived in the forest region. Among the Malinke, West African pygmies were known as Komo Koudoumi. Among peoples in Liberia, West African pygmies were known as Jinna. Among modern West Africans (e.g., Mende of Sierra Leone, Guere of Ivory Coast), there is oral history of their ancestors encountering West African pygmies. Given the varying heights among modern West Africans who dwell within the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
region, this may indicate that admixing had occurred between West African pygmies and the southward migrating ancestors of modern West Africans incoming from the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
region.


Languages

West African hunter-gatherers may have spoken a set of presently
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
Sub-Saharan
West African languages West is one of the four cardinal directions or points of the compass. It is the opposite direction from east and is the direction in which the Sun sets on the Earth. Etymology The word "west" is a Germanic word passed into some Romance langu ...
. In the northeastern region of Nigeria,
Jalaa Jalaa (autonym: ), also known as Cèntûm, Centúúm or Cen Tuum, is an extinct language of northeastern Nigeria (Loojaa settlement in Balanga Local Government Area, Gombe State), of uncertain origins, apparently a language isolate. The Jalabe ( ...
, a
language isolate A language isolate is a language that has no demonstrable genetic relationship with any other languages. Basque in Europe, Ainu and Burushaski in Asia, Sandawe in Africa, Haida and Zuni in North America, Kanoê in South America, and Tiwi ...
, may have been a
descendant language In historical linguistics, a daughter language, also known as descendant language, is a language descended from another language, its mother language, through a process of genetic descent. If more than one language has developed from the same pro ...
from the original set(s) of languages spoken by West African pygmies.


Ancient DNA


Cameroon

Ancient DNA was able to be obtained from two
Shum Laka The archaeological site of Shum Laka is the most prominent rockshelter site in the Grasslands region of the Laka Valley, northwest Cameroon. Occupations at this rockshelter date to the Later Stone Age. This region is important to investigations ...
foragers from the early period of the Stone to Metal Age, in 8000 BP, and two Shum Laka foragers from the late period of the Stone to Metal Age, in 3000 BP. The mitochondrial DNA and Y-Chromosome haplogroups found in the ancient Shum Laka foragers were Sub-Saharan African haplogroups. Two earlier Shum Laka foragers were of haplogroup L0a2a1 – broadly distributed throughout modern African populations – and two later Shum Laka foragers were of haplogroup L1c2a1b – distributed among both modern West and Central African agriculturalists and hunter-gatherers. One earlier Shum Laka forager was of haplogroup B and one later Shum Laka forager haplogroup B2b, which, together, as macrohaplogroup B, is distributed among modern Central African hunter-gatherers (e.g.,
Baka Baka, baká or BAKA may refer to: Ethnicities and languages * Baka people (Cameroon and Gabon), an African ethnic group * Baka people (Congo and South Sudan), an African ethnic group * Baka language, a dialect cluster of Cameroon and Gabon * Baka ...
, Bakola, Biaka, Bedzan). The autosomal admixture of the four ancient Shum Laka forager children was ~35% Western Central African hunter-gatherer and ~65%
Basal West African Basal West African is a hypothetical line of descent that is no longer extant. History Utilizing Western Africans (e.g., Esan of Nigeria, Mende of Sierra Leone, western Gambians), the divergence of a set of early modern or archaic humans, nu ...
– or, an admixture composed of a modern western Central African hunter-gatherer component, a modern West African component, existing locally before 8000 BP, and a modern East African/
West African West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Ma ...
component likely from further north in the regions of the
Sahel The Sahel region (; ), or Sahelian acacia savanna, is a Biogeography, biogeographical region in Africa. It is the Ecotone, transition zone between the more humid Sudanian savannas to its south and the drier Sahara to the north. The Sahel has a ...
and
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
. The two earlier Shum Laka foragers from 8000 BP and two later Shum Laka foragers from 3000 BP show 5000 years of population continuity in region. Yet, modern peoples of Cameroon are more closely related to modern West Africans than to the ancient Shum Laka foragers. Modern Cameroonian hunter-gatherers, while partly descended, are not largely descended from the Shum Laka foragers, due to the apparent absence of descent from Basal West Africans. The
Bantu expansion Bantu may refer to: * Bantu languages, constitute the largest sub-branch of the Niger–Congo languages * Bantu peoples, over 400 peoples of Africa speaking a Bantu language * Bantu knots, a type of African hairstyle * Black Association for Natio ...
is hypothesized to have originated in a homeland of Bantu-speaking peoples located around western Cameroon, a part of which Shum Laka is viewed as being of importance in the early period of this expansion. By 3000 BP, the Bantu expansion is hypothesized to have already begun. Yet, the sampled ancient Shum Laka foragers – two from 8000 BP and two from 3000 BP – show that most modern Niger-Congo speakers are greatly distinct from the ancient Shum Laka foragers, thus, showing that the ancient Shum Laka foragers were not the ancestral source population for modern Bantu-speaking peoples. While
Southern African hunter-gatherers Khoisan ( ) or () is an Hypernymy and hyponymy, umbrella term for the various Indigenous peoples of Africa, indigenous peoples of Southern Africa who traditionally speak non-Bantu languages, combining the Khoekhoen and the San people, Sān peo ...
are generally recognized as being the earliest divergent modern human group, having diverged from other groups around 250,000 BP - 200,000 BP, as a result of the sampling of the ancient Shum Laka foragers, Central African hunter-gatherers are shown to have likely diverged at a similar time, if not even earlier.


References


Bibliography

* * {{cite journal , last1=MacDonald , first1=Kevin , title=Korounkorokalé revisited: The Pays Mande and the West African microlithic technocomplex , url=https://link.springer.com/article/10.1007%2FBF02968406 , journal=African Archaeological Review , year=1997 , volume=14 , issue=3 , pages=192–196 , doi=10.1007/BF02968406 , jstor=25130625 , issn=0263-0338 , oclc=5547019636 , s2cid=161691927, url-access=subscription History of West Africa Prehistoric Africa Stone Age Africa Archaeology of West Africa Ancient peoples of Africa African Pygmies