''Pumiliornis tessellatus'' is an ancient

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

from the Middle Eocene of

Messel Messel is a municipality in the district of Darmstadt-Dieburg in Hesse near Frankfurt am Main in Germany. The village is first mentioned, as ''Masilla'', in the Lorsch codex. Messel was the property of the lords of Groschlag from ca. 1400 to 1799 ...

Hesse, Germany Hesse or Hessen ( ), officially the State of Hesse (), is a States of Germany, state in Germany. Its capital city is Wiesbaden, and the largest urban area is Frankfurt, which is also the country's principal financial centre. Two other major hist ...

. It is described as a

wren Wrens are a family, Troglodytidae, of small brown passerine birds. The family includes 96 species and is divided into 19 genera. All species are restricted to the New World except for the Eurasian wren that is widely distributed in the Old Worl ...

-sized

anisodactyl In biology, dactyly is the arrangement of digits (fingers and toes) on the hands, feet, or sometimes wings of a tetrapod animal. The term is derived from the Greek word () meaning "finger." Sometimes the suffix "-dactylia" is used. The derive ...

bird with a long, slender bill and strong

hallux Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being ''digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being ''plantigrade''; ...

. Its species name ''tessellatus'', meaning "mosaic" in Latin, is a reference to its unusual distribution of characters and uncertain phylogenetic placement. It has some anatomical affinities with Cuculiformes, but similar fossils that might be related to this taxon do not. In 2014, a new specimen of ''Pumiliornis'' was described that showed preserved stomach contents of

pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...

grains from a eudicotyledonous

angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit ...

, making it the earliest fossil evidence of flower-visiting behavior in birds.

References

{{Taxonbar, from=Q16688836 Eocene birds Eocene birds of Europe Fossil taxa described in 1999 Prehistoric bird genera Psittacopedidae