Pulsatilla Grandis
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''Pulsatilla grandis'', commonly known as the greater pasque flower, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the genus '' Pulsatilla'' of the family
Ranunculaceae Ranunculaceae (, buttercup or crowfoot family; Latin "little frog", from "frog") is a family (biology), family of over 2,000 known species of flowering plants in 43 genera, distributed worldwide. The largest genera are ''Ranunculus'' (600 spec ...
. This
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
herbaceous plant reaches heights between 5 and 30 centimetres, featuring finely divided basal leaves and erect, violet flowers densely covered in silky hairs. Native to central Europe, it has a distribution spanning from Great Britain and western France eastward to Ukraine, occupying diverse habitats primarily within the broadleaved forest zone. The species grows predominantly on
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...
-like hillsides and
meadow A meadow ( ) is an open habitat or field, vegetated by grasses, herbs, and other non- woody plants. Trees or shrubs may sparsely populate meadows, as long as they maintain an open character. Meadows can occur naturally under favourable con ...
slopes, often favouring southern exposures. In Slovenia, it is rare, protected, and serves as both a regional symbol and a
heraldic Heraldry is a discipline relating to the design, display and study of armorial bearings (known as armory), as well as related disciplines, such as vexillology, together with the study of ceremony, rank and pedigree. Armory, the best-known branc ...
element in local
iconography Iconography, as a branch of art history, studies the identification, description and interpretation of the content of images: the subjects depicted, the particular compositions and details used to do so, and other elements that are distinct fro ...
.


Description

''Pulsatilla grandis'' is a
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
herbaceous plant reaching heights of between 5 and 30 centimetres. It has upright stems emerging from a basal cluster of leaves, which themselves are attached to the stem by long stalks. These basal leaves are finely divided, branching two to four times into narrower with deeply incised (notched) margins. The flowers, which stand erect, have a funnel-like shape with broadly elliptical petals typically coloured violet, and are densely covered in fine, silky hairs. After flowering, the plant produces dry, single-seeded fruits known as
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple fruit, simple dry fruits, dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and Dehi ...
s. Flower colour in this species is quite variable, showing a range of violet shades.


Habitat and distribution

''Pulsatilla grandis'' occupies diverse habitats primarily within central Europe, extending across Great Britain, western France, Belgium, the Netherlands, Sweden, Austria, Switzerland, Germany, Poland, Hungary, Moldova, Romania, and reaching the eastern limit of its range in Ukraine. Within Ukraine, its distribution is largely diffuse, with populations scattered primarily through the regions of
Podilia Podolia or Podillia is a historic region in Eastern Europe located in the west-central and southwestern parts of Ukraine and northeastern Moldova (i.e. northern Transnistria). Podolia is bordered by the Dniester River and Southern Bug, Boh River. ...
, the Pre-
Carpathians The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe and Southeast Europe. Roughly long, it is the third-longest European mountain range after the Urals at and the Scandinavian Mountains ...
, occasionally in the Right-Bank Forest-Steppe, and infrequently in the Left-Bank Steppe. In Ukraine, ''Pulsatilla grandis'' predominantly favours habitats within the zone of broadleaved forests, where the highest concentration of locations—48 distinct sites—has been documented. Within these areas, it occupies primarily southern and south-eastern slopes of
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...
-like hillsides. Specific physical-geographical regions include Male Polissya, Roztochia-Opillia hills, West Podollia Upland, and the
Prut The Prut (also spelled in English as Pruth; , ) is a river in Eastern Europe. It is a left tributary of the Danube, and is long. Part of its course forms Romania's border with Moldova and Ukraine. Characteristics The Prut originates on the eas ...
Dniester The Dniester ( ) is a transboundary river in Eastern Europe. It runs first through Ukraine and then through Moldova (from which it more or less separates the breakaway territory of Transnistria), finally discharging into the Black Sea on Uk ...
Upland. The species has experienced considerable habitat reduction across these territories, with some historically documented sites now extinct, particularly near larger cities or settlements, likely due to
anthropogenic Anthropogenic ("human" + "generating") is an adjective that may refer to: * Anthropogeny, the study of the origins of humanity Anthropogenic may also refer to things that have been generated by humans, as follows: * Human impact on the enviro ...
pressures. In the forest-steppe belt, populations are fragmented, identified across several highland areas such as the North-western and North-eastern Prydniprovia, the Kyiv highland, and central
Transnistria Transnistria, officially known as the Pridnestrovian Moldavian Republic and locally as Pridnestrovie, is a Landlocked country, landlocked Transnistria conflict#International recognition of Transnistria, breakaway state internationally recogn ...
, with plants typically restricted to rocky, steep, and granite-exposed southern slopes. Many locations recorded historically no longer support existing populations, further underscoring the species' vulnerability. The steppe region harbours fewer localities, where it has historically been found in the Oril
Samara Samara, formerly known as Kuybyshev (1935–1991), is the largest city and administrative centre of Samara Oblast in Russia. The city is located at the confluence of the Volga and the Samara (Volga), Samara rivers, with a population of over 1.14 ...
and
Pryazovia Pryazovia (, sometimes spelled , ''Pryozivia''; ) or literally Cis-Azov region is usually used to refer to the geographic area of the north coast of the Sea of Azov, mostly located in south-eastern Ukraine, with a small part in Russia. It is l ...
lowlands. However, current populations here have markedly declined or disappeared altogether due to anthropogenic impact, including
afforestation Afforestation is the establishment of a forest or stand of trees in an area where there was no recent tree cover. There are three types of afforestation: natural Regeneration (biology), regeneration, agroforestry and Tree plantation, tree plan ...
and land-use changes associated with military training grounds. In the Pre-Carpathians, ''Pulsatilla grandis'' has a pronounced affinity for meadow-steppe slopes and forest clearings. The species was first documented in this area from
herbarium A herbarium (plural: herbaria) is a collection of preserved plant biological specimen, specimens and associated data used for scientific study. The specimens may be whole plants or plant parts; these will usually be in dried form mounted on a sh ...
collections made in the 19th century, and though several historic localities have been lost or not recently confirmed, this region continues to support several populations, primarily in the Bukovinian Pre-Carpathians. Anthropogenic factors such as
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to free range (roam around) and consume wild vegetations in order to feed conversion ratio, convert the otherwise indigestible (by human diges ...
, land cultivation, and the collection of flowers have led to observable declines in these areas as well. In
Slovenia Slovenia, officially the Republic of Slovenia, is a country in Central Europe. It borders Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the south and southeast, and a short (46.6 km) coastline within the Adriati ...
, ''Pulsatilla grandis'' (named , the Easter flower) is a rare and protected plant. It has four growing places in the country all of them are situated in the sub-Pannonian phytogeographical area in the northeastern part of Slovenia. The plant is the symbol of the Boč–Donatus Mountain Landscape Park. On a small rise near the Boč growing place, a stone sculpture of the flower, created by the sculptor Franc Tobias from Razvanje, has been put on display in the 2000s. ''Pulsatilla grandis'' is also depicted on the coat of arms of the Diocese of Celje.


References


External links

*
''Pulsatilla grandis'': Species Range
Maps.iucnredlist.org. {{Taxonbar, from=Q145399 grandis Flora of Austria Flora of Bosnia and Herzegovina Flora of Croatia Flora of the Czech Republic Flora of Germany Flora of Hungary Flora of Moldova Flora of Romania Flora of Serbia Flora of Slovakia Flora of Slovenia Flora of Ukraine