Pug (moth)
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Eupitheciini is a
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
of
geometer moth The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek (derivative form of or "the earth"), and "measure" in referen ...
s under subfamily
Larentiinae Larentiinae is a subfamily of moths containing roughly 5,800 species that occur mostly in the temperate regions of the world. They are generally considered a subfamily of the geometer moth family (Geometridae) and are divided into a few large or ...
, often referred to as pugs. The tribe was described by Tutt in 1896.


Diversity

The tribe consists of about 47 genera, 15 of which are monotypic.


Recognized genera

* '' Aepylopha'' Turner, 1942 * '' Antimimistis'' Turner, 1922 * ''
Ardonis ''Ardonis'' is a genus of moths in the family Geometridae. Species *''Ardonis chlorophilata'' (Walker, 1863) *''Ardonis dentifera'' Warren, 1906 *''Ardonis filicata'' (Swinhoe, 1892) *''Ardonis malachitis'' (Warren, 1903) *''Ardonis thaumasta'' ...
'' Moore, 1888 * '' Axinoptera'' Hampson, 1893 * '' Bosara'' Walker, 1866 * ''
Calluga ''Calluga'' is a genus of moths in the family Geometridae. Species *''Calluga cissocosma'' (Turner, 1904) *''Calluga costalis'' (Moore, 1887) *''Calluga crassitibia'' (Warren 1901) *''Calluga grammophora'' Prout, 1958 *''Calluga longispinata'' ...
'' Moore,
887 __NOTOC__ Year 887 ( DCCCLXXXVII) was a common year starting on Sunday of the Julian calendar. Events By place Europe * November 17 – East Frankish magnates revolt against the inept emperor Charles III (the Fat) in an assembly ...
/small> * '' Carbia'' Walker, 1866 * '' Casuariclystis'' Holloway, 1997 * '' Celaenaclystis'' Holloway, 1997 * ''
Chloroclystis ''Chloroclystis'' is a genus of moths in the family Geometridae first described by Jacob Hübner Jacob Hübner (20 June 1761 – 13 September 1826, in Augsburg) was a German entomologist. He was the author of ''Sammlung Europäischer Schmetterli ...
'' Hübner,
825 __NOTOC__ Year 825 (Roman numerals, DCCCXXV) was a common year starting on Sunday of the Julian calendar. Events By place India * A group of Persio-Assyrian adherents of the Church of the East, under the leadership of two Persian bis ...
/small> * '' Chrysoclystis'' Warren, 1896 * '' Dasimatia'' Warren, 1898 * '' Dissolophodes'' Warren, 1907 * ''
Eupithecia ''Eupithecia'' is the largest genus of moths of the family Geometridae, and the namesake and type genus of tribe Eupitheciini. Species in the genus are, like those of other genera in the tribe, commonly known as pugs. The genus is highly specios ...
'' Curtis, 1825 * '' Eupithystis'' Holloway, 1997 * '' Eriopithex'' Warren, 1896 * '' Eva'' Vojnits, 1981 * ''
Girida ''Girida'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as ...
'' Mironov & Galsworthy, 2012 * ''
Glaucoclystis ''Glaucoclystis'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classif ...
'' Holloway, 1997 * ''
Gymnoscelis ''Gymnoscelis'', the pugs, is a large genus of moths in the family Geometridae described by Paul Mabille in 1868. Description Palpi porrect (extending forward), where the second joint clothed with hair and reaching beyond the frontal tuft. Third ...
'' Mabille, 1868 * ''
Hybridoneura ''Hybridoneura'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classifi ...
'' Warren, 1898 * ''
Mariaba ''Mariaba'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as ...
'' Walker, 1866 * ''
Mesocolpia ''Mesocolpia'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified ...
'' Warren, 1901 * ''
Mesoptila ''Mesoptila'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified ...
'' Meyrick, 1891 * ''
Microdes ''Microdes'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified a ...
'' Guenée, 1857 * ''
Micromia ''Micromia'' is a monotypic moth genus in the family Geometridae. It contains only one species, ''Micromia fulvipuncta'', which is found in New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historicall ...
'' Warren, 1906 * ''
Micrulia ''Micrulia'' is a genus of moths in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified a ...
'' Warren, 1896 * ''
Mnesiloba ''Mnesiloba'' is a genus of moth in the family Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified a ...
'' Warren, 1901 * '' Nasusina'' Pearsall, 1908 * '' Onagrodes'' Warren, 1896 * '' Otucha'' Warren, 1907 * '' Pareupithecia'' Mironov & Galsworthy, 2012 * ''
Pasiphila ''Pasiphila'' is a genus of moths in the family Geometridae. As of 2005 about 36 species were known, and of these, some 27 are native to New Zealand.Viidalepp, J. and V. Mironov. (2005)''Pasiphila hyrcanica'' sp. n.(Geometridae, Larentiinae) – ...
'' Warren, 1895 * '' Pasiphilodes'' Warren, 1895 * '' Phrissogonus'' Butler, 1882 * ''
Polysphalia ''Polysphalia'' is a monotypic moth genus in the family Geometridae. Its only species, ''Polysphalia cristigera'', is found in New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the Li ...
'' Warren, 1906 * '' Pomasia'' Guenee, 1857 * '' Prorella'' Barnes & McDunnough, 1918 * '' Pseudopolynesia'' Holloway, 1997 * '' Ptychotheca'' Warren, 1906 * '' Pycnoloma'' Warren, 1906 * '' Rhinoprora'' Warren, 1895 * '' Scintillithex'' Holloway, 1997 * '' Sigilliclystis'' Galsworthy, 1999 * '' Spiralisigna'' Holloway, 1997 * '' Symmimetis'' Turner, 1907 * ''
Syncosmia ''Syncosmia'' is a genus of moths in the family Geometridae Species description, first described by Warren in 1897. Species *''Syncosmia craspedozona'' (Prout, 1958) *''Syncosmia discisuffusa'' (Holloway, 1976) *''Syncosmia dissographa'' (Prout, ...
'' Warren, 1897 * '' Tripteridia'' Warren, 1903 * ''
Ziridava Ziridava (''Ziridaua'', ) was a Dacian town located between Apulon and Tibiscum, mentioned by Ptolemy in the area of the Dacian tribe of Biephi (today's Romania, Banat region). Ancient sources Ptolemy's ''Geographia'' Ziridava is ment ...
'' Walker, 1863


Former genera

* ''
Collix ''Collix'' is a genus of moths in the family Geometridae first described by Achille Guenée in 1857. Species * '' Collix adamata'' * '' Collix angustipennis'' * ''Collix astathes ''Collix astathes'' is a moth in the family Geometridae. It w ...
'' Guenée in Boisduval & Guenée, 1857 (now in
Melanthiini Melanthiini is a tribe of geometer moths under subfamily Larentiinae. The tribe was described by Philogène Auguste Joseph Duponchel in 1845. Recognized genera * '' Anticollix'' Prout, 1938 * '' Bundelia'' Viidalepp, 1988 * '' Coenocalpe'' Hü ...
) * ''
Eois ''Eois'' is a genus of tropical moths in the family Geometridae. It was first described by Jacob Hübner in 1818. Caterpillars of ''Eois'' species are most commonly green with darker markings, but species with fully dark caterpillars also exist. ...
'' Hübner, 1818 (now in
Asthenini Asthenini is a tribe of geometer moths under subfamily Larentiinae first described by Warren in 1893. The tribe has been combined with Eupitheciini in the past, most notably by Jeremy Daniel Holloway in his work The Moths of Borneo. Recognized ...
) * '' Parasthena'' Warren, 1902 (now in
Asthenini Asthenini is a tribe of geometer moths under subfamily Larentiinae first described by Warren in 1893. The tribe has been combined with Eupitheciini in the past, most notably by Jeremy Daniel Holloway in his work The Moths of Borneo. Recognized ...
) * ''
Poecilasthena ''Poecilasthena'' is a genus of moths in the family Geometridae. Species *unnamed group ** ''Poecilasthena aedaea'' Turner, 1926 ** ''Poecilasthena balioloma'' (Turner, 1907) ** ''Poecilasthena burmensis'' Prout, 1926 ** ''Poecilasthena characte ...
'' Warren, 1894 (now in
Asthenini Asthenini is a tribe of geometer moths under subfamily Larentiinae first described by Warren in 1893. The tribe has been combined with Eupitheciini in the past, most notably by Jeremy Daniel Holloway in his work The Moths of Borneo. Recognized ...
) * ''
Polynesia Polynesia ( , ) is a subregion of Oceania, made up of more than 1,000 islands scattered over the central and southern Pacific Ocean. The indigenous people who inhabit the islands of Polynesia are called Polynesians. They have many things in ...
'' Swinhoe, 1892 (now in
Asthenini Asthenini is a tribe of geometer moths under subfamily Larentiinae first described by Warren in 1893. The tribe has been combined with Eupitheciini in the past, most notably by Jeremy Daniel Holloway in his work The Moths of Borneo. Recognized ...
) * '' Pseudocollix'' Warren, 1895 (now in
Melanthiini Melanthiini is a tribe of geometer moths under subfamily Larentiinae. The tribe was described by Philogène Auguste Joseph Duponchel in 1845. Recognized genera * '' Anticollix'' Prout, 1938 * '' Bundelia'' Viidalepp, 1988 * '' Coenocalpe'' Hü ...
)


References

* (1999)
"New and revised eupitheciine species (Geometridae, Larentiinae) from Hong Kong and South East Asia"
''Transactions of the Lepidopterological Society of Japan''. 50 (3): 223-234. * (2002). "''Bosara'' Walker: revisional notes on the ''Bosara refusaria'' group of species (Geometridae: Larentiinae)". ''Transactions of the Lepidopterological Society of Japan''. 54 (3): 147-155. * (1996). "''Chloroclystis v-ata relicta'' ssp. n. (Lepidoptera: Geometridae), a new taxon from SW Finland, doomed to go extinct?" ''Entomologica Fennica''. 7 (2): 63-66. * (2011)
"A morphological review of tribes in Larentiinae (Lepidoptera: Geometridae)"
''Zootaxa''. 3136: 1–44. Larentiinae Moth tribes {{Eupitheciini-stub