''Pterostichus'' is a very large genus of

ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it ...

s with a

Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...

distribution in the subfamily

Harpalinae Harpalinae is the largest subfamily of ground beetles, containing more than 19,000 species worldwide. Biology This subfamily contains the most apomorphic ground beetles, displaying a wide range of forms and behaviors. The morphology of spe ...

. It has over 1,200 species. The beetles are

predatory Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...

, but sometimes feed on

strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...

. They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species.

Species

These species and subgenera are included in the genus ''Pterostichus''. Subgenus Abea Morita, 1992 * '' Pterostichus yamauchii'' Morita, 1992 Subgenus Adelopterus Reitter, 1886 * '' Pterostichus ambiguus'' (Fairmaire, 1858) Subgenus Adelosia Stephens, 1835 * '' Pterostichus macer'' (Marsham, 1802) Subgenus Agastillus Reitter, 1892 * '' Pterostichus capitatus'' (Chaudoir, 1850) * '' Pterostichus cratocephalus'' (Tschitscherine, 1897) * '' Pterostichus cucujinus'' Reitter, 1892 * '' Pterostichus eriwanicus'' (Tschitscherine, 1897) Subgenus Anilloferonia Van Dyke, 1926 * '' Pterostichus diana'' LaBonte, 2013 * '' Pterostichus malkini'' (Hatch, 1953) * ''

Pterostichus testaceus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Van Dyke, 1926) Subgenus Anomostichus Sciaky, 1995 * '' Pterostichus anomostriatus'' Sciaky, 1995 Subgenus Aphaonus Reitter, 1887 * ''

Pterostichus arcanoides ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries The garden strawberry (or simply ...

'' (Lorenz, 1998) * '' Pterostichus compressus'' (Rost, 1892) * '' Pterostichus cylindriformis'' (Reitter, 1887) * '' Pterostichus koenigianus'' (Tschitscherine, 1891) * ''

Pterostichus mariamae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Zamotajlov; Solodovnikov & Fominykh, 2015 * ''

Pterostichus miroshnikovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Zamotajlov, 1991) * ''

Pterostichus neilgaimani ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries The garden strawberry (or simply ...

'' Chaladze & Kalatozishvili, 2017 * ''

Pterostichus pseudopercus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Reitter, 1889) * ''

Pterostichus starckianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Reitter, 1887) * ''

Pterostichus svetlanae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Zamotajlov; Solodovnikov & Fominykh, 2015 * ''

Pterostichus trubilini ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Zamotajlov, 1997 Subgenus Argutor Dejean, 1821 * ''

Pterostichus aanistschenkoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O. & E.Berlov, 1999 * ''

Pterostichus akozyrevi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O. & E.Berlov, 1999 * ''

Pterostichus bhadravati ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1965 * '' Pterostichus chameleon'' (Motschulsky, 1866) * '' Pterostichus commutabilis'' (Motschulsky, 1866) * '' Pterostichus cryobioides'' (Chaudoir, 1868) * '' Pterostichus cursor'' (Dejean, 1828) * '' Pterostichus dulcis'' (Bates, 1883) * '' Pterostichus kerzhneri'' Lafer, 1983 * '' Pterostichus leonisi'' Apfelbeck, 1904 * ''

Pterostichus praetermissus ''Pterostichus praetermissus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' (Chaudoir, 1868) * ''

Pterostichus sulcitarsis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' A.Morawitz, 1862 * '' Pterostichus vernalis'' (Panzer, 1796) Subgenus Asioplatysma Kryzhanovskij, 1968 * '' Pterostichus capito'' (Tschitscherine, 1900) * '' Pterostichus darvazicus'' Kryzhanovskij, 1968 * '' Pterostichus karateghinicus'' Mikhailov, 1972 * '' Pterostichus klapperichi'' Jedlicka, 1956 * '' Pterostichus medvedevi'' Kryzhanovskij & Mikhailov, 1972 * ''

Pterostichus mujahedeeni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Savich, 1999 * ''

Pterostichus rufopiceus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Heyden, 1890 * '' Pterostichus talibani'' Savich, 1999 Subgenus Badistrinus Motschulsky, 1866 * '' Pterostichus arrowi'' Jedlicka, 1936 * '' Pterostichus arrowianus'' Jedlicka, 1938 * '' Pterostichus bandotaro'' Tanaka, 1958 * '' Pterostichus doris'' Jedlicka, 1962 * '' Pterostichus haptoderoides'' Tschitscherine, 1889 * '' Pterostichus kajimurai'' Habu & Tanaka, 1957 * '' Pterostichus laticollis'' (Motschulsky, 1844) * ''

Pterostichus procephalus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Bates, 1873 Subgenus Bothriopterus Chaudoir, 1835 * '' Pterostichus ademidovae'' O.Berlov, 1996 * '' Pterostichus adstrictus'' Eschscholtz, 1823 * '' Pterostichus aeneocupreus'' (Fairmaire, 1887) * '' Pterostichus bhunetansis'' Davies, 2004 * '' Pterostichus commixtiformis'' Roubal, 1920 * '' Pterostichus ferghanicus'' Kabak, 2001 * '' Pterostichus lustrans'' LeConte, 1851 * '' Pterostichus mariae'' (Lutshnik, 1921) * ''

Pterostichus mutus ''Pterostichus mutus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in Nor ...

'' (Say, 1823) * '' Pterostichus oblongopunctatus'' (Fabricius, 1787) * ''

Pterostichus oregonus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1861 * ''

Pterostichus pensylvanicus ''Pterostichus pensylvanicus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' LeConte, 1873 * '' Pterostichus quadrifoveolatus'' Letzner, 1852 * ''

Pterostichus subovatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1861) * ''

Pterostichus trinarius ''Pterostichus trinarius'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Casey, 1918) * '' Pterostichus tropicalis'' (Bates, 1882) * ''

Pterostichus zyzzovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O. & E.Berlov, 1996 Subgenus Calopterus Chaudoir, 1838 * ''

Pterostichus pilosus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Host, 1790) * '' Pterostichus selmanni'' (Duftschmid, 1812) Subgenus Carllindrothius Habu, 1984 * '' Pterostichus colonus'' (Bates, 1883) Subgenus Cheporus Latreille, 1829 * '' Pterostichus burmeisteri'' Heer, 1837 * '' Pterostichus dissimilis'' (A. & G.B.Villa, 1833) * ''

Pterostichus muehlfeldii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Duftschmid, 1812) * '' Pterostichus transversalis'' (Duftschmid, 1812) Subgenus Chinapterus O.Berlov, 1998 * ''

Pterostichus lianhuaensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Dorjerem; Shi & Liang, 2020 * ''

Pterostichus liupanensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Dorjerem; Shi & Liang, 2020 * '' Pterostichus przewalskyi'' Tschitscherine, 1888 * ''

Pterostichus singularis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tschitscherine, 1889 Subgenus Circinatus Sciaky, 1996 * ''

Pterostichus adelphus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * '' Pterostichus agilis'' Allegro & Sciaky, 2010 * ''

Pterostichus ailaoicus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * '' Pterostichus baenningeri'' Jedlicka, 1931 * ''

Pterostichus batxatensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * '' Pterostichus beneshi'' Sciaky, 1996 * '' Pterostichus bullatus'' Allegro & Sciaky, 2010 * ''

Pterostichus camelus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus caobang ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * '' Pterostichus cavazzutianus'' Shi & Liang, 2015 * '' Pterostichus dentifer'' Allegro & Sciaky, 2010 * '' Pterostichus dimorphus'' Shi & Liang, 2015 * '' Pterostichus expolitus'' Fedorenko, 2019 * ''

Pterostichus hoanglien ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * '' Pterostichus laocaiensis'' Fedorenko, 2019 * '' Pterostichus liciniformis'' Csiki, 1930 * ''

Pterostichus maitreya ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus miao ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus phangxipang ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * '' Pterostichus phiaoc'' Fedorenko, 2019 * '' Pterostichus pohnerti'' Jedlicka, 1934 * '' Pterostichus subtilissimus'' Sciaky, 1996 * ''

Pterostichus tumulus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus wangjiani ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus xilingensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Allegro & Sciaky, 2010 * '' Pterostichus yan'' Shi & Liang, 2015 * ''

Pterostichus yuxiaodongi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus zhygealu ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus zoiai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sciaky, 1996 Subgenus Cophosus Dejean, 1821 * '' Pterostichus cylindricus'' (Herbst, 1784) Subgenus Cryobius Chaudoir, 1838 * '' Pterostichus abaxoides'' (Dejean, 1828) * '' Pterostichus acrogonus'' (Chaudoir, 1843) * '' Pterostichus akkusianus'' Kirschenhofer, 1981 * '' Pterostichus altaiensis'' (Poppius, 1906) * '' Pterostichus amaroides'' (Dejean, 1828) * '' Pterostichus amblypterus'' (Chaudoir, 1868) * '' Pterostichus amoenus'' (Dejean, 1828) * '' Pterostichus amurensis'' (Poppius, 1906) * '' Pterostichus anatolicus'' Jedlicka, 1963 * '' Pterostichus apenninus'' (Dejean, 1831) * '' Pterostichus aralarensis'' (Mateu, 1945) * '' Pterostichus archangaicus'' Shilenkov, 2000 * '' Pterostichus arcticola'' (Chaudoir, 1868) * '' Pterostichus argutoriformis'' (Poppius, 1906) * '' Pterostichus auriga'' Ball, 1962 * '' Pterostichus bargusinicus'' Shilenkov, 2000 * '' Pterostichus barlensis'' Straneo, 1935 * '' Pterostichus barryorum'' Ball, 1962 * '' Pterostichus biocryus'' Ball, 1962 * '' Pterostichus blandulus'' L.Miller, 1859 * '' Pterostichus blumenthali'' Heinz, 1965 * '' Pterostichus brevicornis'' (Kirby, 1837) * '' Pterostichus breviusculus'' (R.F.Sahlberg, 1844) * '' Pterostichus bryanti'' (Van Dyke, 1951) * '' Pterostichus bryantoides'' Ball, 1962 * '' Pterostichus burjaticus'' (Poppius, 1906) * '' Pterostichus cacumensis'' Ball, 1966 * '' Pterostichus cantabricus'' (L.Schaufuss, 1862) * '' Pterostichus caribou'' Ball, 1962 * '' Pterostichus carradei'' (Gautier des Cottes, 1866) * '' Pterostichus champenoisi'' (Croissandeau, 1893) * '' Pterostichus chipewyan'' Ball, 1962 * '' Pterostichus colasi'' (Jeannel, 1937) * '' Pterostichus convexus'' (Gebler, 1847) * '' Pterostichus coreicus'' Jedlicka, 1962 * '' Pterostichus davshensis'' Shilenkov, 2000 * '' Pterostichus dubiosus'' (Tschitscherine, 1894) * '' Pterostichus ehlersi'' (Heyden, 1881) * '' Pterostichus empetricola'' (Dejean, 1828) * '' Pterostichus espanoli'' (J. & E.Vives, 1977) * '' Pterostichus euxinus'' Straneo, 1935 * '' Pterostichus exceptus'' (J.Sahlberg, 1885) * '' Pterostichus firmus'' (Lutshnik, 1933) * '' Pterostichus fulvescens'' (Motschulsky, 1844) * '' Pterostichus gerstlensis'' Ball, 1962 * '' Pterostichus glacialis'' (Brisout de Barneville, 1863) * '' Pterostichus glukhomanka'' Sundukov, 2013 * '' Pterostichus haftorni'' Lindroth, 1969 * '' Pterostichus herzi'' (Poppius, 1906) * '' Pterostichus homalonotus'' (Tschitscherine, 1894) * '' Pterostichus horvatovichi'' Kirschenhofer, 1991 * '' Pterostichus hudsonicus'' LeConte, 1863 * '' Pterostichus infimus'' (Chaudoir, 1868) * '' Pterostichus inopinatus'' (Lutshnik, 1933) * '' Pterostichus iripennis'' (Chaudoir, 1868) * '' Pterostichus ishiharai'' Ishida & Shibata, 1961 * '' Pterostichus jacobsoni'' (Poppius, 1906) * '' Pterostichus jaechi'' Kirschenhofer, 1997 * '' Pterostichus josephi'' Csiki, 1930 * '' Pterostichus kadjaranze'' (Jedlicka, 1947) * '' Pterostichus kaninensis'' (Poppius, 1906) * '' Pterostichus kavanaughi'' Shilenkov, 2000 * '' Pterostichus kaynashensis'' Schweiger, 1967 * '' Pterostichus kolymensis'' Erjiomin, 1998 * '' Pterostichus korgei'' Jedlicka, 1964 * '' Pterostichus kotzebuei'' Ball, 1962 * '' Pterostichus kultianus'' Jedlicka, 1946 * '' Pterostichus kuraicus'' Shilenkov, 2000 * '' Pterostichus kurosai'' Tanaka, 1958 * '' Pterostichus kurosawai'' Tanaka, 1958 * '' Pterostichus lamuticus'' (Poppius, 1906) * '' Pterostichus latiusculus'' (Chaudoir, 1868) * '' Pterostichus longipes'' (Poppius, 1906) * '' Pterostichus lucidus'' (Motschulsky, 1844) * '' Pterostichus macedonicus'' Apfelbeck, 1918 * '' Pterostichus macrothorax'' (Poppius, 1906) * '' Pterostichus maeklini'' (Poppius, 1906) * '' Pterostichus mandibularoides'' Ball, 1966 * '' Pterostichus marani'' Jedlicka, 1967 * '' Pterostichus middendorffi'' (J.Sahlberg, 1875) * ''

Pterostichus negligens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Sturm, 1824) * '' Pterostichus nemoralis'' (Graells, 1851) * '' Pterostichus nigripalpis'' (Poppius, 1906) * ''

Pterostichus nivalis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (R.F.Sahlberg, 1844) * '' Pterostichus nordqvisti'' (J.Sahlberg, 1885) * ''

Pterostichus oblongiusculus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1850) * '' Pterostichus ochoticus'' (R.F.Sahlberg, 1844) * ''

Pterostichus palekhai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' E. & O.Berlov, 1996 * '' Pterostichus parasimilis'' Ball, 1962 * '' Pterostichus parviceps'' (Poppius, 1906) * '' Pterostichus pinguedineus'' (Eschscholtz, 1823) * '' Pterostichus pithyusicus'' (Lutshnik, 1933) * ''

Pterostichus planus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (J.Sahlberg, 1885) * '' Pterostichus poppiusianus'' (Jakobson, 1907) * '' Pterostichus procerulus'' (Heyden, 1880) * '' Pterostichus properans'' (Chaudoir, 1868) * '' Pterostichus pseudostuxbergi'' (Poppius, 1906) * ''

Pterostichus pumilio ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus punctiger ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (J.Sahlberg, 1880) * '' Pterostichus pusillus'' (Dejean, 1828) * ''

Pterostichus quadraticollis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1846) * ''

Pterostichus riparius ''Pterostichus riparius'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Dejean, 1828) * '' Pterostichus rodiovoni'' Shilenkov, 2000 * '' Pterostichus rufonitens'' (Fairmaire, 1866) * ''

Pterostichus safonovae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Budarin, 1995 * ''

Pterostichus sahlbergi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1894) * '' Pterostichus scitus'' (Mäklin, 1878) * ''

Pterostichus shilenkovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Erjiomin & Kabak, 1991 * '' Pterostichus similis'' Mannerheim, 1852 * '' Pterostichus sojot'' Shilenkov, 2000 * '' Pterostichus soperi'' Ball, 1966 * ''

Pterostichus splendidus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Park, 2013 * ''

Pterostichus spornyi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sundukov, 2013 * ''

Pterostichus stantonensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1966 * ''

Pterostichus subgibbus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1860) * ''

Pterostichus subiasi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Ortuño & Zaballos, 1992) * ''

Pterostichus subsinuatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * '' Pterostichus surgens'' LeConte, 1878 * ''

Pterostichus tareumiut ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1962 * ''

Pterostichus taskylensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus tatricus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kult, 1947 * '' Pterostichus theeli'' (Mäklin, 1878) * '' Pterostichus tichomirovi'' Erjiomin, 1990 * ''

Pterostichus tiliaceoradix ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1962 * '' Pterostichus tokmakovae'' Sundukov, 2013 * ''

Pterostichus tunkinensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus udokanensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * '' Pterostichus ulubeyensis'' Kirschenhofer, 1981 * ''

Pterostichus unctulatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Duftschmid, 1812) * '' Pterostichus ventricosus'' (Eschscholtz, 1823) * ''

Pterostichus woodi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball & Currie, 1997 Subgenus Cylindrocharis Casey, 1918 * '' Pterostichus acutipes'' Barr, 1971 * '' Pterostichus hypogeus'' Barr, 1971 * '' Pterostichus rostratus'' (Newman, 1838) Subgenus Eosteropus Tschitscherine, 1902 * '' Pterostichus abensis'' Morita, 2010 * '' Pterostichus abudarini'' O. & E.Berlov, 1997 * '' Pterostichus aenescens'' (Chaudoir, 1850) * '' Pterostichus aereipennis'' (Solsky, 1872) * '' Pterostichus aethiops'' (Panzer, 1796) * '' Pterostichus ainus'' Plutenko, 2005 * '' Pterostichus alacer'' A.Morawitz, 1862 * '' Pterostichus bituberculatus'' (Tschitscherine, 1899) * '' Pterostichus circulosus'' Lindroth, 1966 * '' Pterostichus coruscus'' (Tschitscherine, 1895) * '' Pterostichus creper'' (Tschitscherine, 1902) * '' Pterostichus discrepans'' A.Morawitz, 1862 * '' Pterostichus dudkoi'' Sundukov, 2013 * '' Pterostichus faldermanni'' Schatzmayr, 1929 * '' Pterostichus funakoshii'' Morita, 2007 * '' Pterostichus hidanus'' Morita, 2010 * '' Pterostichus hiramatsui'' Morita, 2010 * '' Pterostichus hirasawai'' Morita, 2010 * '' Pterostichus imurai'' Morita, 2007 * '' Pterostichus japonicus'' (Motschulsky, 1861) * '' Pterostichus karasawai'' Tanaka, 1958 * ''

Pterostichus kimioi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2011 * '' Pterostichus makolskii'' G.Müller, 1933 * '' Pterostichus mandzhuricus'' (Lutshnik, 1916) * '' Pterostichus mannerheimii'' (Dejean, 1831) * '' Pterostichus matsunagai'' Morita, 2011 * '' Pterostichus maurusiacus'' (Mannerheim, 1825) * '' Pterostichus mizunoyai'' Morita, 2007 * '' Pterostichus moestus'' (Say, 1823) * ''

Pterostichus nasuensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2007 * ''

Pterostichus noburai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2008 * '' Pterostichus ohkawai'' Morita, 2010 * ''

Pterostichus orientalis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1844) * '' Pterostichus rengensis'' Morita, 2010 * ''

Pterostichus rufitarsis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus sakuraii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2008 * ''

Pterostichus shimizui ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2010 * ''

Pterostichus sudai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 1991 * ''

Pterostichus superciliosus ''Pterostichus superciliosus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' (Say, 1823) * ''

Pterostichus tenuimarginatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1868) * '' Pterostichus tokui'' Morita, 2007 * ''

Pterostichus tuberculiger ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1897) * '' Pterostichus virescens'' (Gebler, 1833) * ''

Pterostichus yoshizawai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2010 Subgenus Ethira Andrewes, 1936 * '' Pterostichus cometes'' (Andrewes, 1936) * '' Pterostichus depilatus'' (Bates, 1889) * '' Pterostichus heinzianus'' Sciaky, 1996 * '' Pterostichus multiseta'' (Straneo, 1984) * '' Pterostichus piliferus'' (Bates, 1878) * '' Pterostichus pseudopilifer'' (Straneo, 1957) * ''

Pterostichus rugiceps ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * '' Pterostichus seticeps'' (Straneo, 1984) * '' Pterostichus sharanus'' (Straneo, 1984) * ''

Pterostichus variseta ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * '' Pterostichus viridellus'' Straneo, 1984 Subgenus Falsargutor Kryzhanovskij, 1983 * '' Pterostichus kataevi'' Kryzhanovskij, 1989 * ''

Pterostichus ponticus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kirschenhofer, 1987 * ''

Pterostichus pseudopedius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Reitter, 1887 Subgenus Feronidius Jeannel, 1942 * '' Pterostichus brevipennis'' (Chevrolat, 1840) * '' Pterostichus fornicatus'' (Kolenati, 1845) * '' Pterostichus hungaricus'' (Dejean, 1828) * '' Pterostichus incommodus'' Schaum, 1858 * '' Pterostichus melas'' (Creutzer, 1799) Subgenus Feronina Casey, 1918 * '' Pterostichus barri'' Bousquet, 2006 * ''

Pterostichus palmi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Schaeffer, 1910 Subgenus Fukienostichus Lafer, 1983 * ''

Pterostichus shaovuensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1956 Subgenus Gastrosticta Casey, 1918 * '' Pterostichus enodis'' Bousquet, 1992 * ''

Pterostichus mutoides ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Bousquet, 1992 * ''

Pterostichus obesulus ''Pterostichus obesulus'' is a species of beetle first discovered in 1873. No sub-species are listed in the Catalogue of Life The Catalogue of Life (CoL) is an online database that provides an index of known species of animals, plants, fungi, ...

'' LeConte, 1873 * '' Pterostichus ophryoderus'' (Chaudoir, 1878) * '' Pterostichus punctiventris'' (Chaudoir, 1878) * ''

Pterostichus putus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus sayanus ''Pterostichus sayanus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in N ...

'' Csiki, 1930 * '' Pterostichus subacutus'' (Casey, 1918) * '' Pterostichus tumescens'' LeConte, 1863 * ''

Pterostichus ventralis ''Pterostichus ventralis'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Say, 1823) Subgenus Georgeballius Habu, 1984 * '' Pterostichus hoplites'' (Bates, 1883) Subgenus Gutta Wrase & J.Schmidt, 2006 * '' Pterostichus adulterinus'' Wrase & J.Schmidt, 2006 * '' Pterostichus gaoligongensis'' Wrase & J.Schmidt, 2006 * '' Pterostichus kongshuhensis'' B.Gueorguiev, 2015 * '' Pterostichus phungaraziensis'' Wrase & J.Schmidt, 2006 Subgenus Haplomaseus Reitter, 1896 * '' Pterostichus abagonensis'' Reitter, 1896 * '' Pterostichus andreae'' (Tschitscherine, 1897) * '' Pterostichus arator'' (Faldermann, 1836) * '' Pterostichus armenus'' (Faldermann, 1836) * '' Pterostichus astutus'' (Tschitscherine, 1903) * '' Pterostichus balikliensis'' Hovorka & Skoupy, 2007 * '' Pterostichus caucasicola'' (Tschitscherine, 1893) * '' Pterostichus caucasicus'' (Ménétriés, 1832) * '' Pterostichus chasautianus'' Vysoky, 1984 * '' Pterostichus chydaeus'' (Tschitscherine, 1897) * '' Pterostichus consanguineus'' (Chaudoir, 1878) * '' Pterostichus cristicaudis'' Kurnakov, 1962 * '' Pterostichus denticaudis'' Kurnakov, 1962 * '' Pterostichus giresuni'' Jedlicka, 1965 * '' Pterostichus goriensis'' (Tschitscherine, 1897) * '' Pterostichus heinzi'' Jedlicka, 1965 * '' Pterostichus ikizderensis'' Hovorka & Skoupy, 2007 * '' Pterostichus jugicola'' (Lutshnik, 1916) * ''

Pterostichus krulikovskyi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Lutshnik, 1933) * '' Pterostichus meskheticus'' Belousov, 1991 * ''

Pterostichus novotnyorum ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Vysoky, 1981 * ''

Pterostichus olegi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Belousov, 1991 * '' Pterostichus phaeus'' (Lutshnik, 1922) * ''

Pterostichus porcellus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kurnakov, 1962 * ''

Pterostichus pseudoastutus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hovorka & Skoupy, 2007 * '' Pterostichus rousi'' Kirschenhofer, 1982 * '' Pterostichus skoupyi'' Hovorka & Skoupy, 2007 * '' Pterostichus staveni'' Hovorka & Skoupy, 2007 * ''

Pterostichus tamsii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1831) * '' Pterostichus tanjae'' Hovorka & Skoupy, 2007 * '' Pterostichus woronowi'' (Lutshnik, 1916) Subgenus Haptotapinus Reitter, 1886 * '' Pterostichus aksekianus'' (Straneo, 1988) * '' Pterostichus besucheti'' (Straneo, 1988) * '' Pterostichus bythiniensis'' (Maran, 1944) * '' Pterostichus crassiusculus'' (Chaudoir, 1868) * '' Pterostichus dipojranus'' (Straneo, 1988) * '' Pterostichus matchai'' (Jedlicka, 1930) * ''

Pterostichus oblongoparallelus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Maran, 1932 * ''

Pterostichus odontocnemis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' G.Müller, 1931 * ''

Pterostichus orduensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1988) * '' Pterostichus pavani'' (Straneo, 1988) * ''

Pterostichus reissi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' G.Müller, 1931 * ''

Pterostichus vseteckai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Maran, 1944) Subgenus Huaius Tian; Huang; Chen & Ding, 2019 * '' Pterostichus caoi'' Tian; Huang; Chen & Ding, 2019 * '' Pterostichus hanwang'' Tian & He, 2020 * ''

Pterostichus tiankeng ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tian; Huang; Chen & Ding, 2019 * '' Pterostichus yuae'' Tian; Huang; Chen & Ding, 2019 Subgenus Hypherpes Chaudoir, 1838 * '' Pterostichus adoxus'' (Say, 1823) * '' Pterostichus algidus'' LeConte, 1853 * '' Pterostichus amethystinus'' Mannerheim, 1843 * '' Pterostichus annosus'' Casey, 1913 * '' Pterostichus arcanus'' Casey, 1913 * '' Pterostichus baldwini'' (Casey, 1924) * '' Pterostichus barbarinus'' Casey, 1913 * '' Pterostichus brachylobus'' Kavanaugh & LaBonte, 2006 * ''

Pterostichus californicus ''Pterostichus californicus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' (Dejean, 1828) * '' Pterostichus canallatus'' Casey, 1913 * '' Pterostichus castaneus'' (Dejean, 1828) * '' Pterostichus castanipes'' (Ménétriés, 1843) * ''

Pterostichus congestus ''Pterostichus congestus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Ménétriés, 1843) * '' Pterostichus craterensis'' (Hatch, 1949) * '' Pterostichus crenicollis'' LeConte, 1873 * '' Pterostichus ecarinatus'' Hatch, 1936 * '' Pterostichus esuriens'' Casey, 1913 * '' Pterostichus gliscans'' Casey, 1913 * '' Pterostichus gracilior'' LeConte, 1873 * '' Pterostichus herculaneus'' Mannerheim, 1843 * '' Pterostichus hornii'' LeConte, 1873 * '' Pterostichus illustris'' LeConte, 1851 * '' Pterostichus inermis'' Fall, 1901 * '' Pterostichus isabellae'' LeConte, 1851 * '' Pterostichus jacobinus'' Casey, 1913 * '' Pterostichus laborans'' Casey, 1913 * '' Pterostichus lacertus'' Casey, 1913 * '' Pterostichus lama'' (Ménétriés, 1843) * '' Pterostichus lassulus'' (Casey, 1920) * '' Pterostichus lattini'' LaBonte, 2006 * '' Pterostichus luscus'' (Casey, 1918) * '' Pterostichus menetriesii'' LeConte, 1873 * '' Pterostichus mercedianus'' (Casey, 1918) * ''

Pterostichus miscellus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * '' Pterostichus morionides'' (Chaudoir, 1868) * ''

Pterostichus neobrunneus ''Pterostichus neobrunneus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' Lindroth, 1966 * ''

Pterostichus nigrocaeruleus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Van Dyke, 1926 * '' Pterostichus obsidianus'' Casey, 1913 * '' Pterostichus occultus'' Casey, 1913 * '' Pterostichus ordinarius'' Casey, 1913 * ''

Pterostichus ovalipennis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus panticulatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus pergracilis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Casey, 1920) * ''

Pterostichus planctus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1853 * '' Pterostichus protensiformis'' (Casey, 1924) * ''

Pterostichus protractus ''Pterostichus protractus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found i ...

'' LeConte, 1860 * '' Pterostichus restrictus'' (Casey, 1918) * ''

Pterostichus scutellaris ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1873 * ''

Pterostichus sejungendus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1868) * '' Pterostichus serripes'' (LeConte, 1875) * ''

Pterostichus setosus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hatch, 1951 * ''

Pterostichus sierranus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus sponsor ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * '' Pterostichus spraguei'' LeConte, 1873 * '' Pterostichus suffusus'' Casey, 1913 * ''

Pterostichus tarsalis ''Pterostichus tarsalis'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. References Further reading

* Pterostichus, tarsalis Articles created by Qbugbot Beetles described in 1873 {{Pte ...

'' LeConte, 1873 * ''

Pterostichus tristis ''Pterostichus tristis'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in N ...

'' (Dejean, 1828) * '' Pterostichus tuberculofemoratus'' Hatch, 1936 * '' Pterostichus vandykei'' Schaeffer, 1910 * '' Pterostichus vicinus'' Mannerheim, 1843 * ''

Pterostichus ybousqueti ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O.Berlov, 1999 Subgenus Jedlickaia Sciaky, 1997 * ''

Pterostichus sterbai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1934 Subgenus Koreonialoe Park & Kwon, 1996 * '' Pterostichus adatarasanus'' Sasakawa, 2005 * '' Pterostichus akitai'' Morita, 2004 * '' Pterostichus apiculatiphallus'' Nemoto, 1988 * '' Pterostichus asahinus'' Habu & Baba, 1960 * '' Pterostichus bellatrix'' (Tschitscherine, 1895) * '' Pterostichus bifidiphallus'' Nemoto, 1988 * '' Pterostichus chokaisanus'' Sasakawa, 2009 * '' Pterostichus eboshiyamanus'' Sasakawa, 2009 * '' Pterostichus falcispinus'' Sasakawa, 2005 * '' Pterostichus fukube'' Sugimura, 2005 * '' Pterostichus gassanus'' Sasakawa, 2009 * ''

Pterostichus gujoensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Toda, 2012 * '' Pterostichus ishikawai'' Nemoto, 1988 * '' Pterostichus ishikawaioides'' Sasakawa; J.L.Kim; J.K.Kim & Kubota, 2008 * '' Pterostichus isolatus'' Sasakawa; Kim & Kubota, 2005 * '' Pterostichus iwakiensis'' Sasakawa, 2009 * ''

Pterostichus iwasakii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kasahara, 1990 * '' Pterostichus ixion'' (Tschitscherine, 1902) * '' Pterostichus jiricola'' Sasakawa; J.L.Kim; J.K.Kim & Kubota, 2008 * '' Pterostichus kitakamisanus'' Sasakawa, 2005 * '' Pterostichus koheii'' Nakane, 1963 * '' Pterostichus kuraiyamanus'' Morita & Ohkawa, 2010 * '' Pterostichus kurikomasanus'' Sasakawa, 2005 * '' Pterostichus kuwolsanensis'' Li & Zhang, 2014 * '' Pterostichus macrogenys'' Bates, 1883 * '' Pterostichus microps'' Heyden, 1887 * ''

Pterostichus miyazawai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita & Ohkawa, 2009 * ''

Pterostichus momuranus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita; Ohkawa & Kurihara, 2013 * '' Pterostichus monolineatus'' Sasakawa; Mitsuduka & H.Ito, 2020 * '' Pterostichus nagasawai'' N.Ito & Ogai, 2015 * ''

Pterostichus ohsawacavus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa, 2005 * '' Pterostichus opacipennis'' Jedlicka, 1934 * '' Pterostichus palgongsanus'' Nemoto, 1988 * ''

Pterostichus shikatai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Toda, 2012 * ''

Pterostichus shinbodakensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa & H.Ito, 2017 * '' Pterostichus shirakamisan'' Sasakawa, 2009 * '' Pterostichus shirakamisanus'' Sasakawa, 2005 * ''

Pterostichus shojii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sugimura, 2006 * ''

Pterostichus sumondakensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa, 2005 * '' Pterostichus syleus'' Kirschenhofer, 1997 * ''

Pterostichus taebaegsanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Nemoto, 1988 * ''

Pterostichus takadateyamanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa, 2009 * ''

Pterostichus tanakai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ishida, 1964 * ''

Pterostichus tateishiyamanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa & H.Ito, 2017 * ''

Pterostichus teretis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Park & Kwon, 1996 * '' Pterostichus todai'' Morita & Kanie, 1997 * ''

Pterostichus togyusanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Park & Kwon, 1996 * '' Pterostichus toyodai'' Morita & Kurosa, 1998 * '' Pterostichus uedaorum'' Morita & Hirasawa, 1996 * '' Pterostichus vicinus'' Park & Kwon, 1996 * '' Pterostichus woongbii'' Park & Kwon, 1996 * ''

Pterostichus yahikosanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa, 2009 * ''

Pterostichus yamizosanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa, 2005 * ''

Pterostichus yorikoae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sugimura, 2007 Subgenus Kozyrevius O.Berlov, 1998 * '' Pterostichus militaris'' (Tschitscherine, 1893) Subgenus Lamenius Bousquet, 1999 * '' Pterostichus caudicalis'' (Say, 1823) Subgenus Lenapterus O.Berlov, 1996 * '' Pterostichus agonus'' G.Horn, 1880 * '' Pterostichus cancellatus'' (Motschulsky, 1860) * '' Pterostichus costatus'' (Ménétriés, 1851) * '' Pterostichus galae'' Farkac & Plutenko, 1996 * '' Pterostichus gromykoi'' Sundukov, 2005 * '' Pterostichus marginatus'' Matsumura, 1911 * ''

Pterostichus punctatissimus ''Pterostichus punctatissimus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are fou ...

'' (Randall, 1838) * '' Pterostichus rugosipennis'' Jedlicka, 1932 * ''

Pterostichus saxicola ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1899) * '' Pterostichus vermiculosus'' (Ménétriés, 1851) Subgenus Leptoferonia Casey, 1918 * '' Pterostichus angustus'' (Dejean, 1828) * '' Pterostichus beyeri'' Van Dyke, 1926 * '' Pterostichus blodgettensis'' Will, 2007 * '' Pterostichus caligans'' G.Horn, 1891 * '' Pterostichus cochlearis'' Hacker, 1968 * '' Pterostichus deino'' Will, 2007 * '' Pterostichus enyo'' Will, 2007 * '' Pterostichus falli'' Van Dyke, 1926 * '' Pterostichus fenyesi'' Csiki, 1930 * '' Pterostichus fuchsi'' Schaeffer, 1910 * '' Pterostichus hackerae'' Will, 2016 * '' Pterostichus hatchi'' Hacker, 1968 * '' Pterostichus humilis'' Casey, 1913 * '' Pterostichus idahoae'' Csiki, 1930 * '' Pterostichus inanis'' G.Horn, 1891 * '' Pterostichus infernalis'' Hatch, 1936 * '' Pterostichus inopinus'' (Casey, 1918) * '' Pterostichus lobatus'' Hacker, 1968 * '' Pterostichus marinensis'' Hacker, 1968 * '' Pterostichus mattolensis'' Hacker, 1968 * ''

Pterostichus pemphredo ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Will, 2007 * '' Pterostichus pumilus'' Casey, 1913 * '' Pterostichus rothi'' (Hatch, 1951) * '' Pterostichus sphodrinus'' LeConte, 1863 * '' Pterostichus stapedius'' Hacker, 1968 * ''

Pterostichus trinitensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hacker, 1968 * '' Pterostichus yosemitensis'' Hacker, 1968 Subgenus Lianoe Gozis, 1882 * '' Pterostichus amanoi'' Nakane, 1968 * '' Pterostichus arribasi'' Ortuño, 1991 * '' Pterostichus asturicus'' (Jeanne, 1966) * '' Pterostichus cordatissimus'' (Straneo, 1937) * '' Pterostichus drescoi'' Nègre, 1957 * '' Pterostichus dufourii'' (Dejean, 1828) * '' Pterostichus microphthalmus'' Delarouzée, 1857 * '' Pterostichus muelleri'' (Straneo, 1936) * '' Pterostichus nadari'' (Vuillefroy, 1893) * ''

Pterostichus nakamiyorinus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita; Ohkawa & Kurihara, 2013 * '' Pterostichus rousselli'' Colas, 1963 * '' Pterostichus sakagamii'' Morita, 2001 * '' Pterostichus sudrei'' Sainte-Claire Deville, 1922 Subgenus Lyrothorax Chaudoir, 1838 * '' Pterostichus amagisanus'' Tanaka & Ishida, 1972 * '' Pterostichus caspius'' (Ménétriés, 1832) * '' Pterostichus eoyoritomus'' Sasakawa, 2009 * '' Pterostichus fujitai'' Tanaka & Ishida, 1972 * '' Pterostichus yoritomus'' Bates, 1873 Subgenus Megabea Sciaky, 1997 * '' Pterostichus tienmushanus'' Sciaky, 1997 Subgenus Melanius Bonelli, 1810 * '' Pterostichus aterrimus'' (Herbst, 1784) * '' Pterostichus castor'' Goulet & Bousquet, 1983 * '' Pterostichus corvinus'' (Dejean, 1828) * '' Pterostichus ebeninus'' (Dejean, 1828) * '' Pterostichus elongatus'' (Duftschmid, 1812) * '' Pterostichus licenti'' Jedlicka, 1939 * '' Pterostichus noguchii'' Bates, 1873 Subgenus Metallophilus Chaudoir, 1838 * '' Pterostichus dandongensis'' Kirschenhofer, 1997 * '' Pterostichus interruptus'' (Dejean, 1828) * '' Pterostichus kamtschaticus'' Motschulsky, 1860 * '' Pterostichus mirus'' (Tschitscherine, 1894) * '' Pterostichus orion'' (Tschitscherine, 1901) * ''

Pterostichus pfizenmayeri ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Poppius, 1906 * '' Pterostichus rugosus'' (Gebler, 1823) * '' Pterostichus sublaevis'' (J.Sahlberg, 1880) Subgenus Micronialoe Park; Kwon & Lafer, 1996 * ''

Pterostichus bifoveolatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Park; Kwon & Lafer, 1996 * '' Pterostichus chogyesanus'' Park; Kwon & Lafer, 1996 * '' Pterostichus kaniei'' Morita, 2005 Subgenus Monoferonia Casey, 1918 * '' Pterostichus carolinus'' Darlington, 1932 * '' Pterostichus diligendus'' (Chaudoir, 1868) * '' Pterostichus mancus'' (LeConte, 1853) * '' Pterostichus primus'' Darlington, 1932 Subgenus Morphohaptoderus Tschitscherine, 1898 * '' Pterostichus bowanus'' Sciaky, 1997 * '' Pterostichus cervenkai'' Sciaky, 1994 * '' Pterostichus chungkingi'' Jedlicka, 1932 * '' Pterostichus confucius'' Sciaky & Wrase, 1997 * '' Pterostichus dentellus'' Facchini & Sciaky, 2003 * '' Pterostichus dundai'' Sciaky, 1994 * '' Pterostichus emei'' Sciaky, 1994 * '' Pterostichus expeditus'' (Tschitscherine, 1898) * '' Pterostichus geberti'' Sciaky & Wrase, 1997 * '' Pterostichus giacomazzoi'' Sciaky, 1994 * '' Pterostichus gongga'' Sciaky, 1997 * '' Pterostichus guizhouensis'' Sciaky, 1997 * '' Pterostichus huashanus'' Sciaky, 1994 * '' Pterostichus hubeicus'' Facchini & Sciaky, 2003 * '' Pterostichus irideus'' Sciaky, 1994 * '' Pterostichus janatai'' Sciaky & Wrase, 1997 * '' Pterostichus kalabi'' Sciaky, 1994 * '' Pterostichus kucerai'' Sciaky, 1997 * '' Pterostichus lingshanus'' Sciaky & Wrase, 1997 * '' Pterostichus maximus'' Tschitscherine, 1889 * '' Pterostichus megaloderus'' Sciaky, 1994 * '' Pterostichus ming'' Sciaky & Wrase, 1997 * '' Pterostichus miroslavi'' Sciaky & Wrase, 1997 * '' Pterostichus muellermotzfeldi'' Wrase & J.Schmidt, 2006 * '' Pterostichus parvicollis'' Sciaky & Wrase, 1997 * ''

Pterostichus pseudoplatyderus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sciaky, 1994 * '' Pterostichus saueri'' Sciaky, 1994 * '' Pterostichus schuelkei'' Sciaky & Wrase, 1997 * ''

Pterostichus shennongjianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Facchini & Sciaky, 2003 * ''

Pterostichus straneellus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1938 * ''

Pterostichus toledanoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Facchini & Sciaky, 2003 * ''

Pterostichus wenxianensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Allegro & Sciaky, 2010 * '' Pterostichus yulongshanensis'' Sciaky, 1997 Subgenus Myosodus Fischer von Waldheim, 1823 * '' Pterostichus aapsorum'' Belousov, 1991 * '' Pterostichus aibgensis'' Starck, 1890 * '' Pterostichus avaricus'' Abdurakhmanov & Kryzhanovskij, 1983 * '' Pterostichus batesi'' (Tschitscherine, 1894) * '' Pterostichus buglaniensis'' Kirschenhofer, 1981 * '' Pterostichus filipjevi'' (Lutshnik, 1928) * '' Pterostichus ingusha'' (Lutshnik, 1928) * '' Pterostichus kiritshenkoi'' (Lutshnik, 1928) * '' Pterostichus lacunosus'' (Chaudoir, 1844) * '' Pterostichus lutshnikianus'' Bogachev & Kurnakov, 1958 * ''

Pterostichus nivicola ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Ménétriés, 1832) * '' Pterostichus ordinatus'' (Fischer von Waldheim, 1823) * ''

Pterostichus rudestriatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Reitter, 1883) * '' Pterostichus schoenherri'' Faldermann, 1836 * '' Pterostichus sodalicius'' Heyden, 1885 * '' Pterostichus starcki'' Heyden, 1885 * '' Pterostichus stoeckleini'' (Straneo, 1941) * ''

Pterostichus svanicus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Lutshnik, 1922) * '' Pterostichus swaneticus'' (Reitter, 1883) * ''

Pterostichus variabilis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Ménétriés, 1832) * ''

Pterostichus zamotajlovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Belousov, 1991 Subgenus Neohaptoderus Tschitscherine, 1898 * '' Pterostichus berezowskii'' (Tschitscherine, 1898) * '' Pterostichus catei'' Sciaky & Wrase, 1997 * '' Pterostichus comorus'' Jedlicka, 1932 * '' Pterostichus gravis'' Jedlicka, 1939 * '' Pterostichus haesitatus'' Fairmaire, 1889 * '' Pterostichus ignavus'' (Tschitscherine, 1897) * '' Pterostichus jureceki'' (Jedlicka, 1936) * '' Pterostichus kleinfeldianus'' Sciaky & Wrase, 1997 * '' Pterostichus komalus'' (Jedlicka, 1936) * '' Pterostichus maderi'' Jedlicka, 1938 * ''

Pterostichus molopsoides ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1934 * ''

Pterostichus montigena ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1898) * ''

Pterostichus mundus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1938 * '' Pterostichus oreophilus'' (Tschitscherine, 1898) * ''

Pterostichus orestes ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Jedlicka, 1936) * ''

Pterostichus sinicus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1897) * '' Pterostichus szetschuanensis'' Tschitscherine, 1889 * ''

Pterostichus yunnanensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1934 Subgenus Nialoe Tanaka, 1958 * '' Pterostichus asymmetricus'' Bates, 1883 * '' Pterostichus basilobatus'' Sasakawa, 2005 * '' Pterostichus biexcisus'' Straneo, 1955 * '' Pterostichus bisetosus'' (Straneo, 1938) * '' Pterostichus brunneipennis'' Straneo, 1955 * '' Pterostichus carsticus'' Kasahara & Y.Ito, 1989 * '' Pterostichus chujoi'' Habu, 1959 * '' Pterostichus cristatoides'' Straneo, 1955 * '' Pterostichus daihizanus'' Ishida, 1968 * '' Pterostichus daisenicus'' Ishida, 1958 * '' Pterostichus dandonis'' Kasahara, 1989 * '' Pterostichus enasanus'' Morita, 2007 * '' Pterostichus fujimurai'' Habu, 1958 * '' Pterostichus fujisanus'' Tanaka & Suga, 1972 * ''

Pterostichus fumikoae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kurosa, 2019 * '' Pterostichus hakusanus'' Kasahara, 1989 * ''

Pterostichus hekosanensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa, 2020 * '' Pterostichus himifuriho'' Morita, 2007 * ''

Pterostichus hiraii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2015 * '' Pterostichus hirobane'' Habu & Baba, 1958 * '' Pterostichus hozumii'' Ishida, 1961 * '' Pterostichus ikukoae'' Morita, 2007 * '' Pterostichus ishizukai'' Kasahara, 1995 * '' Pterostichus isumiensis'' Kasahara & Saito, 1997 * '' Pterostichus janoi'' Jedlicka, 1952 * '' Pterostichus jogaesanensis'' Lafer; Paik & Park, 1996 * '' Pterostichus katashinensis'' Habu, 1958 * ''

Pterostichus kaya ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2019 * '' Pterostichus kongosanus'' Nakane, 1963 * '' Pterostichus latistylis'' Tanaka, 1958 * '' Pterostichus masahiroi'' Kasahara, 1988 * '' Pterostichus masatakai'' Morita, 2007 * '' Pterostichus mirificus'' Bates, 1883 * '' Pterostichus mitoyamanus'' Tanaka, 1971 * ''

Pterostichus musashiensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kasahara, 1993 * '' Pterostichus nakanei'' Straneo, 1955 * '' Pterostichus naokii'' Morita, 2003 * ''

Pterostichus napaea ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kasahara, 1988 * ''

Pterostichus nishiyamai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kasahara, 1986 * ''

Pterostichus ogaensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 1995 * '' Pterostichus ohbayashii'' Ishida, 1968 * ''

Pterostichus ohdaisanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Nakane, 1963 * ''

Pterostichus ohkurai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 1996 * ''

Pterostichus okutamae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tanaka, 1963 * ''

Pterostichus omogoensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Nakane, 1972 * ''

Pterostichus ompoensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1932 * '' Pterostichus opaculus'' N.Ito, 2010 * '' Pterostichus ovaliphallus'' Sasakawa, 2005 * ''

Pterostichus parkwon ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Davies, 2004 * ''

Pterostichus praedo ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1901) * ''

Pterostichus rhanis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1902) * ''

Pterostichus ryoheii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2012 * ''

Pterostichus ryomoensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita & Suda, 2007 * '' Pterostichus satoi'' Ishida, 1961 * '' Pterostichus shibatai'' Ishida, 1961 * '' Pterostichus shotaroi'' Morita, 1987 * '' Pterostichus sincerus'' Park & Kwon, 1996 * '' Pterostichus spiculifer'' Bates, 1883 * ''

Pterostichus sugimurai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2007 * '' Pterostichus tahirai'' Kasahara, 1992 * ''

Pterostichus takahashii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ishida, 1958 * '' Pterostichus tanakaorum'' Morita & Ohkura, 1988 * ''

Pterostichus taoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kasahara, 1993 * ''

Pterostichus tokejii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Yoshida & Tanaka, 1960 * ''

Pterostichus tottoriensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2003 * ''

Pterostichus tsurugiyamanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Habu, 1959 * '' Pterostichus uchiyamai'' Morita, 1987 * ''

Pterostichus uenoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Straneo, 1955 * ''

Pterostichus watanabei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Nakane, 1960 * '' Pterostichus yamashitai'' Morita, 2019 * ''

Pterostichus yanoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1953 * ''

Pterostichus yatsuensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Straneo, 1955 * '' Pterostichus yokohamae'' Straneo in Nakane, 1979 Subgenus Oreolyperus Tschitscherine, 1901 * '' Pterostichus heptapotamicus'' (Lutshnik, 1927) * '' Pterostichus korolkowi'' (Tschitscherine, 1901) * ''

Pterostichus necessarius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1894) * '' Pterostichus regeli'' (Tschitscherine, 1894) Subgenus Oreophilus Chaudoir, 1838 * '' Pterostichus alberti'' (Jeannel, 1942) * '' Pterostichus cribratus'' (Dejean, 1828) * ''

Pterostichus dubius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Heer, 1838 * '' Pterostichus duratii'' A. & G.B.Villa, 1835 * '' Pterostichus externepunctatus'' (Dejean, 1828) * '' Pterostichus flavofemoratus'' (Dejean, 1828) * '' Pterostichus franzi'' Nègre, 1955 * '' Pterostichus ienishteai'' Nitzu, 1988 * '' Pterostichus impressus'' (Fairmaire & Laboulbène, 1854) * '' Pterostichus jurinei'' (Panzer, 1802) * '' Pterostichus morio'' (Duftschmid, 1812) * ''

Pterostichus multipunctatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus parnassius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Schaum, 1859 * '' Pterostichus paulini'' (Vuillefroy, 1868) * '' Pterostichus planiusculus'' (Chaudoir, 1859) * '' Pterostichus rufipennis'' Baudi di Selve, 1889 * '' Pterostichus spinolae'' (Dejean, 1828) * ''

Pterostichus variolatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus xatartii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * '' Pterostichus yvanii'' (Dejean, 1828) Subgenus Oreoplatysma Jakobson, 1907 * '' Pterostichus abishirensis'' Belousov, 1991 * '' Pterostichus araraticus'' Kirschenhofer, 1987 * '' Pterostichus asinubas'' Davies, 2004 * '' Pterostichus belizini'' (Lutshnik, 1933) * '' Pterostichus belousovi'' Kryzhanovskij, 1989 * '' Pterostichus borcka'' Jedlicka, 1963 * '' Pterostichus capitolinus'' Kurnakov, 1962 * '' Pterostichus casaleianus'' Kirschenhofer, 1991 * '' Pterostichus cecchiniae'' (Jakobson, 1907) * '' Pterostichus chefsuricus'' Reitter, 1896 * ''Pterostichus colchicus'' (Chaudoir, 1850) * ''Pterostichus cordifer'' Reitter, 1896 * ''Pterostichus cunibakus'' Davies, 2004 * ''Pterostichus daghestanus'' Reitter, 1896 * ''Pterostichus depressidorsis'' Reitter, 1896 * ''Pterostichus faunus'' Kurnakov, 1962 * ''Pterostichus fencli'' Dvorak, 1995 * ''Pterostichus ghilarovi'' Kryzhanovskij, 1988 * ''Pterostichus jakobsonianus'' (Lutshnik, 1928) * ''Pterostichus kadleci'' Dvorak, 1995 * ''Pterostichus kirschenblatti'' Kryzhanovskij, 1988 * ''Pterostichus koenigi'' (Reitter, 1887) * ''Pterostichus krasnopolensis'' Kirschenhofer, 1987 * ''Pterostichus kvirensis'' Belousov, 1991 * ''Pterostichus lodosi'' Heinz, 1977 * ''Pterostichus mediocris'' Dvorak, 1995 * ''Pterostichus nikodymi'' Dvorak, 1995 * ''Pterostichus odvarkai'' Dvorak, 1995 * ''Pterostichus orman'' Jedlicka, 1963 * ''Pterostichus pasanauricus'' Kirschenhofer, 1987 * ''Pterostichus percontator'' Reitter, 1887 * ''Pterostichus planaticollis'' Kirschenhofer, 1987 * ''Pterostichus platyderus'' (Chaudoir, 1850) * ''Pterostichus pulchellus'' (Faldermann, 1836) * ''Pterostichus rousianus'' Kirschenhofer, 1987 * ''Pterostichus rubripalpis'' Csiki, 1930 * ''Pterostichus satunini'' (Tschitscherine, 1903) * ''Pterostichus satyrus'' Kurnakov, 1962 * ''Pterostichus schodaicus'' Kirschenhofer, 1987 * ''Pterostichus setosicus'' Davies, 2004 * ''Pterostichus sojaki'' Kirschenhofer, 1987 * ''Pterostichus stomoides'' (Chaudoir, 1868) * ''Pterostichus strasseri'' Reitter, 1898 * ''Pterostichus tamarae'' Wrase & Kirschenhofer, 1991 * ''Pterostichus validiceps'' Reitter, 1887 * ''Pterostichus zolotarewi'' Reitter, 1911 Subgenus Orientostichus Sciaky & Allegro, 2013 * ''Pterostichus chinensis'' (Jedlicka, 1962) * ''Pterostichus curtatus'' Fairmaire, 1886 * ''Pterostichus davidi'' (Tschitscherine, 1897) * ''Pterostichus deceptrix'' (Tschitscherine, 1898) * ''Pterostichus distinctissimus'' Jedlicka, 1940 * ''Pterostichus ferreroi'' Straneo, 1989 * ''Pterostichus gallopavo'' Sciaky & Wrase, 1997 * ''Pterostichus lesticoides'' (Straneo, 1939) * ''Pterostichus machulkai'' Jedlicka, 1931 * ''Pterostichus perlutus'' Jedlicka, 1938 * ''Pterostichus prattii'' Bates, 1890 * ''Pterostichus pulcher'' Sciaky & Allegro, 2013 * ''Pterostichus semirugosus'' (Andrewes, 1947) * ''Pterostichus simillimus'' Fairmaire, 1886 * ''Pterostichus tachongi'' Jedlicka, 1936 Subgenus Orsonjohnsonus Hatch, 1933 * ''Pterostichus johnsoni'' Ulke, 1889 Subgenus Paraferonia Casey, 1918 * ''Pterostichus lubricus'' LeConte, 1853 Subgenus Parahaptoderus Jeanne, 1969 * ''Pterostichus bielzii'' (Fuss, 1858) * ''Pterostichus brevis'' (Duftschmid, 1812) * ''Pterostichus vecors'' (Tschitscherine, 1897) Subgenus Paralianoe Ishida, 1958 * ''Pterostichus kiiensis'' Morita & Ohkura, 1988 Subgenus Parapterostichus Desbrochers des Loges, 1906 * ''Pterostichus grajus'' (Dejean, 1828) * ''Pterostichus justusii'' W.Redtenbacher, 1842 * ''Pterostichus nodicornis'' (Fairmaire & Laboulbène, 1854) * ''Pterostichus schaschli'' (Marseul, 1880) Subgenus Petrophilus Chaudoir, 1838 * ''Pterostichus abnormis'' (J.Sahlberg, 1880) * ''Pterostichus acutidens'' (Fairmaire, 1888) * ''Pterostichus alexandrovi'' Lafer, 1979 * ''Pterostichus alexeji'' Zamotajlov & Kryzhanovskij, 1992 * ''Pterostichus altaianus'' Jedlicka, 1958 * ''Pterostichus altaicus'' (Germar, 1823) * ''Pterostichus arsenjevi'' Lafer, 1979 * ''Pterostichus bungei'' (Tschitscherine, 1894) * ''Pterostichus calvitarsis'' Breit, 1912 * ''Pterostichus chechcirensis'' Lafer, 1979 * ''Pterostichus chinkiangensis'' Kirschenhofer, 1991 * ''Pterostichus coracinus'' (Newman, 1838) * ''Pterostichus dauricus'' (Gebler, 1832) * ''Pterostichus dilutipes'' (Motschulsky, 1844) * ''Pterostichus elmbergi'' Poppius, 1908 * ''Pterostichus eximius'' A.Morawitz, 1862 * ''Pterostichus fallettii'' Sciaky, 1996 * ''Pterostichus findelii'' (Dejean, 1828) * ''Pterostichus foveolatus'' (Duftschmid, 1812) * ''Pterostichus glaferi'' O. & E.Berlov, 1996 * ''Pterostichus iberlovi'' O.Berlov, 1996 * ''Pterostichus jungens'' (Tschitscherine, 1893) * ''Pterostichus jurecekianus'' Maran, 1940 * ''Pterostichus kokeilii'' L.Miller, 1850 * ''Pterostichus kurentzovi'' Lafer, 1979 * ''Pterostichus kvanmobongus'' Lafer, 2011 * ''Pterostichus lachrymosus'' (Newman, 1838) * ''Pterostichus levadensis'' Lafer, 1979 * ''Pterostichus magoides'' (Straneo, 1937) * ''Pterostichus magus'' (Mannerheim, 1825) * ''Pterostichus maryseae'' Sun & Shi, 2018 * ''Pterostichus melanarius'' (Illiger, 1798) * ''Pterostichus melanodes'' (Chaudoir, 1878) * ''Pterostichus mellyi'' (Gebler, 1843) * ''Pterostichus montanus'' (Motschulsky, 1844) * ''Pterostichus monticoloides'' Shilenkov in Kryzhanovskij et al., 1995 * ''Pterostichus mroczkowskii'' Lafer, 2011 * ''Pterostichus nigellus'' A.Morawitz, 1862 * ''Pterostichus novus'' Straneo, 1944 * ''Pterostichus odaesanensis'' Lafer, 2011 * ''Pterostichus pawlowskii'' Lafer, 2011 * ''Pterostichus pertinax'' (Tschitscherine, 1895) * ''Pterostichus petulans'' Jedlicka, 1938 * ''Pterostichus poppiusi'' (Semenov, 1906) * ''Pterostichus probus'' Park & Kwon, 1996 * ''Pterostichus procax'' A.Morawitz, 1862 * ''Pterostichus rasilis'' Park & Kwon, 1996 * ''Pterostichus relictus'' (Newman, 1838) * ''Pterostichus robustistylis'' Sasakawa, 2005 * ''Pterostichus schoenmanni'' Kirschenhofer, 1991 * ''Pterostichus sejunctus'' Bates, 1883 * ''Pterostichus septentrionis'' (Chaudoir, 1868) * ''Pterostichus seriatus'' (Chaudoir, 1850) * ''Pterostichus seungmoi'' Park & Kwon, 1996 * ''Pterostichus shingarevi'' Lafer, 1979 * ''Pterostichus silvestris'' Sun & Shi, 2018 * ''Pterostichus songoricus'' (Motschulsky, 1845) * ''Pterostichus stygicus'' (Say, 1823) * ''Pterostichus subaeneus'' (Chaudoir, 1850) * ''Pterostichus sungariensis'' Lafer, 1979 * ''Pterostichus sutschanensis'' Jedlicka, 1962 * ''Pterostichus tatianae'' E.Berlov, 1996 * ''Pterostichus thunbergi'' A.Morawitz, 1862 * ''Pterostichus tomensis'' (Gebler, 1847) * ''Pterostichus triseriatus'' (Gebler, 1847) * ''Pterostichus tschitscherinianus'' (Jakobson, 1907) * ''Pterostichus tuberifer'' Sasakawa, 2006 * ''Pterostichus tundrae'' (Tschitscherine, 1894) * ''Pterostichus turanensis'' Jedlicka, 1959 * ''Pterostichus uralensis'' (Motschulsky, 1850) * ''Pterostichus urengaicus'' Jurecek, 1924 * ''Pterostichus vladivostokensis'' Lafer, 1979 Subgenus Phaenoraphis Tschitscherine, 1901 * ''Pterostichus acuspina'' (Tschitscherine, 1901) * ''Pterostichus peninsularis'' Park & Kwon, 1996 * ''Pterostichus sagittus'' Park, 2013 Subgenus Phonias Gozis, 1886 * ''Pterostichus apfelbecki'' Csiki, 1908 * ''Pterostichus burkhan'' Berlov & Anichtchenko, 2005 * ''Pterostichus corrusculus'' LeConte, 1873 * ''Pterostichus datshenkoae'' Sundukov, 2013 * ''Pterostichus defossus'' Bates, 1883 * ''Pterostichus diligens'' (Sturm, 1824) * ''Pterostichus dostali'' Kirschenhofer, 1981 * ''Pterostichus eobius'' (Tschitscherine, 1899) * ''Pterostichus femoralis'' (Kirby, 1837) * ''Pterostichus gyrosus'' (Motschulsky, 1866) * ''Pterostichus innshanensis'' Jedlicka, 1960 * ''Pterostichus jankowskyi'' (Tschitscherine, 1897) * ''Pterostichus kutensis'' Poppius, 1905 * ''Pterostichus liodactylus'' (Tschitscherine, 1898) * ''Pterostichus longinquus'' Bates, 1873 * ''Pterostichus longipennis'' Straneo, 1942 * ''Pterostichus monostigma'' (Tschitscherine, 1898) * ''Pterostichus morawitzianus'' (Lutshnik, 1922) * ''Pterostichus neglectus'' A.Morawitz, 1862 * ''Pterostichus ovoideus'' (Sturm, 1824) * ''Pterostichus patruelis'' (Dejean, 1831) * ''Pterostichus perisi'' Novoa, 1979 * ''Pterostichus ripensis'' (Motschulsky, 1866) * ''Pterostichus sasajii'' Morita, 2007 * ''Pterostichus setipes'' (Tschitscherine, 1898) * ''Pterostichus sotkaensis'' Jedlicka, 1958 * ''Pterostichus strenuus'' (Panzer, 1796) * ''Pterostichus stricticollis'' (Solsky, 1874) * ''Pterostichus subitus'' Csiki, 1930 * ''Pterostichus taksonyis'' Csiki, 1930 * ''Pterostichus ussuriensis'' (Tschitscherine, 1897) Subgenus Platypterus Chaudoir, 1838 * ''Pterostichus dilatatus'' A. & G.B.Villa, 1835 * ''Pterostichus lineatopunctatus'' L.Miller, 1850 * ''Pterostichus lombardus'' K.Daniel, 1900 * ''Pterostichus ottomanus'' Apfelbeck, 1908 * ''Pterostichus panzeri'' (Panzer, 1802) * ''Pterostichus truncatus'' (Dejean, 1828) * ''Pterostichus ziegleri'' (Duftschmid, 1812) Subgenus Platysma Bonelli, 1810 * ''Pterostichus chotanensis'' (Tschitscherine, 1893) * ''Pterostichus cordaticollis'' Heyden, 1884 * ''Pterostichus crenulatopunctatus'' (R.F.Sahlberg, 1844) * ''Pterostichus eschscholtzii'' (Germar, 1823) * ''Pterostichus galinae'' Kabak, 1992 * ''Pterostichus insignicollis'' (Tschitscherine, 1893) * ''Pterostichus leptis'' Bates, 1883 * ''Pterostichus niger'' (Schaller, 1783) * ''Pterostichus planicola'' (Tschitscherine, 1899) * ''Pterostichus sachtlebeni'' Jedlicka, 1962 * ''Pterostichus turcomannicus'' Motschulsky, 1850 Subgenus Plectes Fischer von Waldheim, 1822 * ''Pterostichus consors'' (Tschitscherine, 1893) * ''Pterostichus drescheri'' (Fischer von Waldheim, 1817) Subgenus Pledarus Motschulsky, 1866 * ''Pterostichus gibbicollis'' (Motschulsky, 1844) * ''Pterostichus larisae'' Sundukov, 2013 * ''Pterostichus lutschniki'' Jedlicka, 1962 * ''Pterostichus nigellatus'' Kirschenhofer, 1991 Subgenus Pseudethira Sciaky, 1996 * ''Pterostichus angoarnigi'' Morvan, 1994 * ''Pterostichus atrox'' (Andrewes, 1937) * ''Pterostichus balachowskyi'' Morvan, 1972 * ''Pterostichus balu'' J.Schmidt, 2009 * ''Pterostichus bhutanensis'' Morvan, 1978 * ''Pterostichus brancuccii'' Straneo, 1982 * ''Pterostichus brevilama'' Straneo, 1989 * ''Pterostichus bureli'' Morvan, 1994 * ''Pterostichus chainapaani'' J.Schmidt, 2006 * ''Pterostichus championi'' Andrewes, 1926 * ''Pterostichus conaensis'' J.Schmidt & Tian, 2011 * ''Pterostichus deuvei'' Lassalle, 1985 * ''Pterostichus dhorpatanicus'' Straneo, 1977 * ''Pterostichus dolens'' (Tschitscherine, 1900) * ''Pterostichus dorjulensis'' Morvan, 1978 * ''Pterostichus exochus'' Andrewes, 1930 * ''Pterostichus fritzhiekei'' J.Schmidt, 1994 * ''Pterostichus gagates'' (Hope, 1831) * ''Pterostichus ganesh'' J.Schmidt, 2006 * ''Pterostichus ganja'' J.Schmidt, 1995 * ''Pterostichus gerdi'' J.Schmidt, 2009 * ''Pterostichus gobettii'' Straneo, 1983 * ''Pterostichus gompanus'' Straneo, 1983 * ''Pterostichus harmandi'' (Tschitscherine, 1900) * ''Pterostichus hartmanni'' J.Schmidt, 1995 * ''Pterostichus hedkeinekus'' Morvan, 1995 * ''Pterostichus hoelli'' J.Schmidt, 2009 * ''Pterostichus immarginatus'' Straneo, 1977 * ''Pterostichus jaljaleensis'' J.Schmidt, 2009 * ''Pterostichus janbritoi'' Morvan, 1994 * ''Pterostichus juga'' J.Schmidt, 2006 * ''Pterostichus kadoudali'' Morvan, 1982 * ''Pterostichus kalo'' J.Schmidt, 2012 * ''Pterostichus kangchenjunga'' J.Schmidt, 2012 * ''Pterostichus keltiekus'' Morvan, 1981 * ''Pterostichus kleinfeldi'' Straneo, 1982 * ''Pterostichus kopetzi'' J.Schmidt, 2006 * ''Pterostichus leica'' J.Schmidt, 2012 * ''Pterostichus letensis'' Habu, 1973 * ''Pterostichus lugi'' Morvan, 1982 * ''Pterostichus makalu'' J.Schmidt, 2012 * ''Pterostichus matsumurai'' Habu, 1973 * ''Pterostichus mewakholensis'' J.Schmidt, 2012 * ''Pterostichus miehei'' J.Schmidt, 2006 * ''Pterostichus nepalensis'' Straneo, 1977 * ''Pterostichus nowitzkii'' (Tschitscherine, 1899) * ''Pterostichus olafi'' J.Schmidt, 2006 * ''Pterostichus perlamatus'' Straneo, 1989 * ''Pterostichus pseudoharmandi'' Morvan, 1981 * ''Pterostichus pseudoplatysma'' Straneo, 1982 * ''Pterostichus raraensis'' Morvan, 1980 * ''Pterostichus rugosiceps'' J.Schmidt, 2012 * ''Pterostichus sagarmatha'' J.Schmidt, 2012 * ''Pterostichus santostamangi'' J.Schmidt, 2006 * ''Pterostichus schrettenbrunneri'' J.Schmidt, 2012 * ''Pterostichus thanglaensis'' J.Schmidt, 2012 * ''Pterostichus tulobalu'' J.Schmidt, 2009 * ''Pterostichus weigeli'' J.Schmidt, 2006 * ''Pterostichus weiperti'' J.Schmidt, 2006 * ''Pterostichus wittmeri'' Morvan, 1978 Subgenus Pseudoferonina Ball, 1965 * ''Pterostichus amadeus'' Bousquet, 2012 * ''Pterostichus bousqueti'' Bergdahl in Bergdahl & Kavanaugh, 2011 * ''Pterostichus campbelli'' Bousquet, 1985 * ''Pterostichus humidulus'' (Van Dyke, 1943) * ''Pterostichus lanei'' Van Dyke, 1926 * ''Pterostichus lolo'' Bergdahl in Bergdahl & Kavanaugh, 2011 * ''Pterostichus shulli'' (Hatch, 1949) * ''Pterostichus smetanai'' Bousquet, 1985 * ''Pterostichus spathifer'' Bousquet, 1992 Subgenus Pseudohaptoderus Tschitscherine, 1888 * ''Pterostichus aemiliae'' Facchini & Sciaky, 2000 * ''Pterostichus jugivagus'' (Tschitscherine, 1898) * ''Pterostichus ssemenovi'' Tschitscherine, 1888 Subgenus Pseudomaseus Chaudoir, 1838 * ''Pterostichus aimaki'' Jedlicka, 1968 * ''Pterostichus ambigenus'' Bates, 1883 * ''Pterostichus anthracinus'' (Illiger, 1798) * ''Pterostichus basipunctatus'' Straneo, 1955 * ''Pterostichus carri'' Angus; Galian; Wrase & Chaladze, 2009 * ''Pterostichus chujoiellus'' Jedlicka, 1962 * ''Pterostichus fuscicornis'' (Reiche & Saulcy, 1855) * ''Pterostichus gracilis'' (Dejean, 1828) * ''Pterostichus hejkali'' B.Gueorguiev & Skoupy, 2011 * ''Pterostichus koslovi'' Solodovnikov, 2001 * ''Pterostichus luctuosus'' (Dejean, 1828) * ''Pterostichus michailovi'' Wrase, 1992 * ''Pterostichus minor'' (Gyllenhal, 1827) * ''Pterostichus mukdenensis'' Breit, 1933 * ''Pterostichus nigrita'' (Paykull, 1790) * ''Pterostichus oenotrius'' Ravizza, 1975 * ''Pterostichus piceolus'' (Chaudoir, 1850) * ''Pterostichus rhaeticus'' Heer, 1837 * ''Pterostichus rotundangulus'' A.Morawitz, 1862 * ''Pterostichus tenuis'' (Casey, 1924) * ''Pterostichus tsukubasanus'' Kasahara, 1988 Subgenus Pseudorambousekiella Schweiger, 1967 * ''Pterostichus montisdeorum'' Schweiger, 1967 Subgenus Pseudorites Ganglbauer, 1891 * ''Pterostichus nicaeensis'' (A. & G.B.Villa, 1835) Subgenus Pseudosteropus Chaudoir, 1838 * ''Pterostichus illigeri'' (Panzer, 1802) * ''Pterostichus schmidtii'' (Chaudoir, 1838) Subgenus Pterostichus Bonelli, 1810 * ''Pterostichus andreinii'' Dodero, 1922 * ''Pterostichus auratus'' Heer, 1837 * ''Pterostichus bischoffianus'' Jedlicka, 1936 * ''Pterostichus bruckii'' Schaum, 1859 * ''Pterostichus cantalicus'' (Chaudoir, 1868) * ''Pterostichus cristatus'' (L.Dufour, 1820) * ''Pterostichus devillei'' Puel, 1924 * ''Pterostichus dux'' L.Schaufuss, 1862 * ''Pterostichus epiroticus'' Csiki, 1930 * ''Pterostichus fasciatopunctatus'' (Creutzer, 1799) * ''Pterostichus focarilei'' Casale & Giachino, 1985 * ''Pterostichus funestes'' Csiki, 1930 * ''Pterostichus hagenbachii'' Sturm, 1824 * ''Pterostichus honnoratii'' (Dejean, 1828) * ''Pterostichus impressicollis'' (Fairmaire & Laboulbène, 1854) * ''Pterostichus lanista'' (Tschitscherine, 1898) * ''Pterostichus lasserrei'' (Dejean, 1828) * ''Pterostichus latifianus'' Apfelbeck, 1906 * ''Pterostichus lumensis'' Apfelbeck, 1905 * ''Pterostichus malissorum'' Apfelbeck, 1905 * ''Pterostichus meisteri'' Reitter, 1885 * ''Pterostichus merklii'' (J.Frivaldszky, 1879) * ''Pterostichus micans'' Heer, 1838 * ''Pterostichus pedemontanus'' Ganglbauer, 1891 * ''Pterostichus pentheri'' Apfelbeck, 1918 * ''Pterostichus phaeopus'' (Sainte-Claire Deville, 1903) * ''Pterostichus reiseri'' Ganglbauer, 1889 * ''Pterostichus rhilensis'' Rottenberg, 1874 * ''Pterostichus ruffoi'' Sciaky, 1984 * ''Pterostichus rufipes'' (Dejean, 1828) * ''Pterostichus rutilans'' (Dejean, 1828) * ''Pterostichus sacheri'' I.Frivaldszky von Frivald, 1865 * ''Pterostichus vagepunctatus'' Heer, 1837 * ''Pterostichus walteri'' Reitter, 1883 Subgenus Rambousekiella Knirsch, 1925 * ''Pterostichus ledenikensis'' (Knirsch, 1925) Subgenus Rhagadus Motschulsky, 1866 * ''Pterostichus brittoni'' Habu, 1958 * ''Pterostichus glabripennis'' Jedlicka, 1962 * ''Pterostichus harponifer'' Tanaka, 1987 * ''Pterostichus ishiii'' Morita; Kurosa & Mori, 2009 * ''Pterostichus kalhys'' (Motschulsky, 1866) * ''Pterostichus kimurai'' Morita, 1994 * ''Pterostichus komiyai'' Morita, 2007 * ''Pterostichus laevipunctatus'' Tschitscherine, 1889 * ''Pterostichus latemarginatus'' (Straneo, 1936) * ''Pterostichus microcephalus'' (Motschulsky, 1861) * ''Pterostichus modicellus'' (Tschitscherine, 1897) * ''Pterostichus mundatus'' Jedlicka, 1940 * ''Pterostichus nimbatidius'' (Chaudoir, 1878) * ''Pterostichus polygenus'' Bates, 1883 * ''Pterostichus satsumanus'' Habu, 1958 * ''Pterostichus solskyi'' (Chaudoir, 1878) * ''Pterostichus takaosanus'' Habu, 1958 * ''Pterostichus thorectes'' Bates, 1873 * ''Pterostichus thorectoides'' Jedlicka, 1958 Subgenus Sinoreophilus Sciaky, 1996 * ''Pterostichus potanini'' Tschitscherine, 1889 * ''Pterostichus scalptus'' Sciaky & Wrase, 1997 * ''Pterostichus strigosus'' Sciaky & Wrase, 1997 * ''Pterostichus validior'' Tschitscherine, 1889 Subgenus Sinosteropus Sciaky, 1994 * ''Pterostichus barbarae'' Sciaky & Facchini, 2003 * ''Pterostichus carinatus'' Sciaky & Facchini, 2003 * ''Pterostichus cathaicus'' Sciaky, 1994 * ''Pterostichus curvatus'' Sciaky & Facchini, 2003 * ''Pterostichus horstweiperti'' J.Schmidt & Guorguiev, 2013 * ''Pterostichus jaechianus'' Sciaky & Facchini, 2003 * ''Pterostichus jani'' Sciaky & Facchini, 2003 * ''Pterostichus latitemporis'' Sciaky & Wrase, 1997 * ''Pterostichus microbus'' Sciaky & Facchini, 2003 * ''Pterostichus perhoplites'' J.Schmidt & Tian, 2011 * ''Pterostichus pseudobarbarae'' Sciaky & Facchini, 2003 * ''Pterostichus pseudojugivagus'' J.Schmidt & Tian, 2011 * ''Pterostichus pseudorotundus'' Sciaky & Facchini, 2003 * ''Pterostichus pseudosinensis'' Sciaky & Facchini, 2003 * ''Pterostichus rotundus'' Sciaky, 1994 * ''Pterostichus schneideri'' Sciaky & Facchini, 2003 * ''Pterostichus scuticollis'' (Fairmaire, 1889) * ''Pterostichus septemtrionalis'' Sciaky & Facchini, 2003 * ''Pterostichus sinensis'' Jedlicka, 1962 * ''Pterostichus triangularis'' Sciaky & Facchini, 2003 * ''Pterostichus wrasei'' Sciaky & Facchini, 2003 Subgenus Sphodroferonia Kasahara & Y.Ito, 1987 * ''Pterostichus abaciformis'' Straneo, 1955 * ''Pterostichus arcuaticarinatus'' Kasahara, 1986 * ''Pterostichus audax'' (Tschitscherine, 1895) * ''Pterostichus bilobatus'' N.Ito, 2010 * ''Pterostichus bisulcitarsis'' N.Ito, 2012 * ''Pterostichus confusianus'' N.Ito, 2010 * ''Pterostichus constricticollis'' N.Ito, 2010 * ''Pterostichus esakii'' Ishida, 1959 * ''Pterostichus fenestratus'' Kasahara & Ohtani, 1989 * ''Pterostichus geojensis'' Sasakawa; Kim & Kubota, 2006 * ''Pterostichus gotoensis'' Kasahara & Matsumoto, 1990 * ''Pterostichus gracilitarsis'' N.Ito, 2010 * ''Pterostichus hikosanus'' Kasahara, 1994 * ''Pterostichus hisamatsui'' Ishida & Shibata, 1961 * ''Pterostichus imasakai'' Kasahara & Ohtani, 1988 * ''Pterostichus imitatorius'' N.Ito, 2010 * ''Pterostichus ishizuchiensis'' Kasahara, 1985 * ''Pterostichus itoi'' Kasahara, 1986 * ''Pterostichus kosakai'' Morita, 1998 * ''Pterostichus koujitanakai'' N.Ito, 2010 * ''Pterostichus kyushuensis'' Habu, 1955 * ''Pterostichus laeviceps'' N.Ito, 2010 * ''Pterostichus macrocephalus'' Habu, 1955 * ''Pterostichus marginellus'' Sasakawa; Kim & Kim, 2006 * ''Pterostichus masatakayoshidai'' N.Ito, 2010 * ''Pterostichus masidai'' Ishida, 1959 * ''Pterostichus masumotoi'' Tanaka; Morita & Suga, 1987 * ''Pterostichus miyamai'' Kasahara & Y.Ito, 1987 * ''Pterostichus mosaicus'' Sasakawa, 2005 * ''Pterostichus mucronatus'' Straneo, 1955 * ''Pterostichus multinodosus'' Sasakawa; Kim & Kim, 2006 * ''Pterostichus namedai'' Morita, 2019 * ''Pterostichus nasui'' Kasahara, 1993 * ''Pterostichus nishidai'' Nakane, 1989 * ''Pterostichus obscurimpressus'' N.Ito, 2010 * ''Pterostichus okiensis'' Nakane, 1989 * ''Pterostichus orionis'' Jedlicka, 1962 * ''Pterostichus pachinus'' Bates, 1883 * ''Pterostichus platyocularis'' N.Ito, 2010 * ''Pterostichus platypennis'' N.Ito, 2010 * ''Pterostichus plesiomorphus'' Nemoto, 1989 * ''Pterostichus prominens'' N.Ito, 2010 * ''Pterostichus pseudopachinus'' Nakane, 1963 * ''Pterostichus punctatulus'' N.Ito, 2010 * ''Pterostichus punctibasalis'' N.Ito, 2012 * ''Pterostichus raptor'' (Tschitscherine, 1901) * ''Pterostichus sakuragii'' N.Ito, 2010 * ''Pterostichus sculpturalis'' N.Ito, 2012 * ''Pterostichus scurra'' (Tschitscherine, 1901) * ''Pterostichus scurroides'' Jedlicka, 1958 * ''Pterostichus sediorcrus'' Sasakawa & Kim, 2006 * ''Pterostichus seunglaki'' Park & Kwon, 1996 * ''Pterostichus shiibanus'' Habu, 1958 * ''Pterostichus sphodriformis'' Bates, 1873 * ''Pterostichus strigipennis'' N.Ito, 2012 * ''Pterostichus subabacipennis'' N.Ito, 2012 * ''Pterostichus sulcatus'' N.Ito, 2010 * ''Pterostichus suruganus'' Straneo in Nakane, 1979 * ''Pterostichus symmetricus'' Straneo, 1955 * ''Pterostichus taradakensis'' Kasahara & Ohtani, 1988 * ''Pterostichus tosanus'' Kasahara & Y.Ito, 1997 * ''Pterostichus touzalinioides'' Sasakawa; Kim & Kim, 2007 * ''Pterostichus tsurugaensis'' Ishida, 1959 * ''Pterostichus variicornis'' Sasakawa; Kim & Kim, 2007 * ''Pterostichus yakushimanus'' Nakane & Ishida, 1961 * ''Pterostichus yamajii'' Kasahara, 1993 * ''Pterostichus yoshidai'' Kasahara, 1985 * ''Pterostichus yoshikawai'' Ishida, 1959 * ''Pterostichus yoshiyukiitoi'' N.Ito, 2010 Subgenus Steropanus Fairmaire, 1889 * ''Pterostichus aequus'' (Andrewes, 1937) * ''Pterostichus alveolatus'' Fedorenko, 2018 * ''Pterostichus asulcatus'' Fedorenko, 2018 * ''Pterostichus boriskataevi'' Fedorenko, 2020 * ''Pterostichus cavifrons'' Fedorenko, 2018 * ''Pterostichus deliciatus'' (Andrewes, 1937) * ''Pterostichus felix'' (Andrewes, 1937) * ''Pterostichus forticornis'' (Fairmaire, 1889) * ''Pterostichus fossifrons'' Fedorenko, 2020 * ''Pterostichus glymmiger'' (Andrewes, 1937) * ''Pterostichus infissus'' (Andrewes, 1937) * ''Pterostichus mengtzei'' Jedlicka, 1931 * ''Pterostichus obliteratus'' Fedorenko, 2018 * ''Pterostichus pseudoglymmiger'' Fedorenko, 2020 * ''Pterostichus pseudoviolaceus'' Fedorenko, 2018 * ''Pterostichus securipenis'' Fedorenko, 2020 * ''Pterostichus sulcatipennis'' Fedorenko, 2018 * ''Pterostichus thailandensis'' (Morvan, 1992) * ''Pterostichus thailandicus'' Straneo, 1989 * ''Pterostichus violaceus'' (Straneo, 1949) Subgenus Steropus Dejean, 1821 * ''Pterostichus catalonicus'' J.Daniel in K. & J.Daniel, 1906 * ''Pterostichus ebenus'' (Quensel, 1806) * ''Pterostichus ferreri'' (Español & Mateu, 1942) * ''Pterostichus galaecianus'' Lauffer, 1909 * ''Pterostichus gallega'' (Fairmaire, 1859) * ''Pterostichus ghilianii'' (Putzeys, 1845) * ''Pterostichus insidatrix'' (Piochard de la Brûlerie, 1872) * ''Pterostichus madidus'' (Fabricius, 1775) * ''Pterostichus rifensis'' Antoine, 1933 * ''Pterostichus validus'' (Dejean, 1828) Subgenus Tausternus Fedorenko, 2020 * ''Pterostichus hoii'' Fedorenko, 2020 Subgenus Tinautius Mateu, 1997 * ''Pterostichus exilis'' (Mateu, 2001) * ''Pterostichus troglophilus'' (Mateu, 1997) Subgenus Tschitscherinea Berg, 1898 * ''Pterostichus farkaci'' Sciaky, 1997 * ''Pterostichus filum'' (Tschitscherine, 1897) * ''Pterostichus krali'' Sciaky, 1997 * ''Pterostichus mulensis'' Sciaky, 1997 Subgenus Tubuliphallus Sciaky & Allegro, 2013 * ''Pterostichus megacephalus'' Sciaky & Allegro, 2013 Subgenus Unitrichus Sciaky, 1997 * ''Pterostichus platyops'' Sciaky, 1997 Subgenus Vietosteropus Fedorenko, 2017 * ''Pterostichus abramovi'' Fedorenko, 2017 * ''Pterostichus anichkini'' Fedorenko, 2017 * ''Pterostichus annamita'' (Straneo, 1939) * ''Pterostichus bidoupensis'' Fedorenko, 2017 * ''Pterostichus cavicollis'' Straneo, 1984 * ''Pterostichus chupanphan'' Fedorenko, 2017 * ''Pterostichus dalatensis'' Fedorenko, 2017 * ''Pterostichus gialaiensis'' Fedorenko, 2017 * ''Pterostichus honbaensis'' Fedorenko, 2017 * ''Pterostichus konchurang'' Fedorenko, 2017 * ''Pterostichus konplongensis'' Fedorenko, 2017 * ''Pterostichus kontumensis'' Fedorenko, 2017 * ''Pterostichus kuznetsovi'' Fedorenko, 2020 * ''Pterostichus ngoclinhensis'' Fedorenko, 2017 * ''Pterostichus ngokboci'' Fedorenko, 2017 * ''Pterostichus semiopacus'' Fedorenko, 2017 * ''Pterostichus sulcicollis'' Fedorenko, 2017 Subgenus Wraseiellus Shi & Sciaky, 2013 * ''Pterostichus andrewesi'' Jedlicka, 1931 * ''Pterostichus comatus'' Shi & Sciaky, 2013 * ''Pterostichus crassiapex'' Shi & Sciaky, 2013 * ''Pterostichus diversus'' (Fairmaire, 1886) * ''Pterostichus kambaiti'' (Andrewes, 1947) * ''Pterostichus meyeri'' Jedlicka, 1934 * ''Pterostichus pseudodiversus'' Shi & Sciaky, 2013 * ''Pterostichus stictopleurus'' (Fairmaire, 1889) Unassigned Subgenus * ''Pterostichus analis'' Jedlicka, 1963 * ''Pterostichus antepunctatus'' Straneo, 1949 * ''Pterostichus aristochroides'' Deuve, 2006 * ''Pterostichus chenpengi'' Li, 1992 * ''Pterostichus clepsydra'' Sciaky & Wrase, 1997 * ''Pterostichus deuvesianus'' Morvan, 1995 * ''Pterostichus georgi'' Straneo, 1963 * ''Pterostichus glabricollis'' Jedlicka, 1962 * ''Pterostichus gwervaenus'' Morvan, 1995 * ''Pterostichus heilongjiangensis'' Li, 1992 * ''Pterostichus imitatus'' Morvan, 1978 * ''Pterostichus jelepus'' Andrewes, 1932 * ''Pterostichus kiangsu'' Jedlicka, 1965 * ''Pterostichus kolosovi'' (Lutshnik, 1928) * ''Pterostichus leviculus'' Andrewes, 1933 * ''Pterostichus lidarus'' (Andrewes, 1937) * ''Pterostichus lukjanovitshi'' (Lutshnik, 1927) * ''Pterostichus malaisei'' Jedlicka, 1965 * ''Pterostichus migliaccioi'' Straneo, 1982 * ''Pterostichus orobius'' Fairmaire, 1861 * ''Pterostichus peilingi'' Jedlicka, 1937 * ''Pterostichus punctisternus'' Straneo, 1989 * ''Pterostichus quadriimpressus'' Straneo, 1983 * ''Pterostichus reuteri'' Wrase & J.Schmidt, 2006 * ''Pterostichus rolex'' Morvan, 1995 * ''Pterostichus sintanus'' Andrewes, 1927 * ''Pterostichus styx'' (Andrewes, 1937) * ''Pterostichus szekessyianus'' Sciaky, 1996 * ''Pterostichus tantillus'' (Fairmaire, 1889) * ''Pterostichus tonkinensis'' Straneo, 1980 * ''Pterostichus zhejiangensis'' Kirschenhofer, 1997 * †''Pterostichus abrogatus'' Scudder, 1890 * †''Pterostichus depletus'' Scudder, 1900 * †''Pterostichus destitutus'' Scudder, 1890 * †''Pterostichus destructus'' Scudder, 1890 * †''Pterostichus dormitans'' Scudder, 1890 * †''Pterostichus fernquisti'' Wickham, 1931 * †''Pterostichus fractus'' Scudder, 1890 * †''Pterostichus laevigatus'' Scudder, 1890 * †''Pterostichus minax'' (Oustalet, 1874) * †''Pterostichus minutulus'' Heer, 1862 * †''Pterostichus provincialis'' (Oustalet, 1874) * †''Pterostichus pumpellyi'' Scudder, 1900 * †''Pterostichus walcotti'' Scudder, 1900

References

{{Taxonbar, from=Q1540893 Pterostichus, Pterostichinae Taxa named by Franco Andrea Bonelli