''Pterostichus'' is a very large genus of

ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it ...

s with a

Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...

distribution in the subfamily

Harpalinae Harpalinae is the largest subfamily of ground beetles, containing more than 19,000 species worldwide. Biology This subfamily contains the most apomorphic ground beetles, displaying a wide range of forms and behaviors. The morphology of spe ...

. It has over 1,200 species. The beetles are

predatory Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...

, but sometimes feed on

strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...

. They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species.

Species

These species and subgenera are included in the genus ''Pterostichus''. Subgenus Abea Morita, 1992 * ''

Pterostichus yamauchii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 1992 Subgenus Adelopterus Reitter, 1886 * '' Pterostichus ambiguus'' (Fairmaire, 1858) Subgenus Adelosia Stephens, 1835 * '' Pterostichus macer'' (Marsham, 1802) Subgenus Agastillus Reitter, 1892 * '' Pterostichus capitatus'' (Chaudoir, 1850) * '' Pterostichus cratocephalus'' (Tschitscherine, 1897) * '' Pterostichus cucujinus'' Reitter, 1892 * '' Pterostichus eriwanicus'' (Tschitscherine, 1897) Subgenus Anilloferonia Van Dyke, 1926 * '' Pterostichus diana'' LaBonte, 2013 * '' Pterostichus malkini'' (Hatch, 1953) * ''

Pterostichus testaceus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Van Dyke, 1926) Subgenus Anomostichus Sciaky, 1995 * '' Pterostichus anomostriatus'' Sciaky, 1995 Subgenus Aphaonus Reitter, 1887 * ''

Pterostichus arcanoides ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries The garden strawberry (or simply ...

'' (Lorenz, 1998) * '' Pterostichus compressus'' (Rost, 1892) * '' Pterostichus cylindriformis'' (Reitter, 1887) * '' Pterostichus koenigianus'' (Tschitscherine, 1891) * ''

Pterostichus mariamae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Zamotajlov; Solodovnikov & Fominykh, 2015 * ''

Pterostichus miroshnikovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Zamotajlov, 1991) * ''

Pterostichus neilgaimani ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries The garden strawberry (or simply ...

'' Chaladze & Kalatozishvili, 2017 * ''

Pterostichus pseudopercus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Reitter, 1889) * ''

Pterostichus starckianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Reitter, 1887) * ''

Pterostichus svetlanae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Zamotajlov; Solodovnikov & Fominykh, 2015 * ''

Pterostichus trubilini ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Zamotajlov, 1997 Subgenus Argutor Dejean, 1821 * ''

Pterostichus aanistschenkoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O. & E.Berlov, 1999 * ''

Pterostichus akozyrevi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O. & E.Berlov, 1999 * ''

Pterostichus bhadravati ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1965 * '' Pterostichus chameleon'' (Motschulsky, 1866) * '' Pterostichus commutabilis'' (Motschulsky, 1866) * '' Pterostichus cryobioides'' (Chaudoir, 1868) * '' Pterostichus cursor'' (Dejean, 1828) * '' Pterostichus dulcis'' (Bates, 1883) * '' Pterostichus kerzhneri'' Lafer, 1983 * '' Pterostichus leonisi'' Apfelbeck, 1904 * ''

Pterostichus praetermissus ''Pterostichus praetermissus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' (Chaudoir, 1868) * ''

Pterostichus sulcitarsis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' A.Morawitz, 1862 * '' Pterostichus vernalis'' (Panzer, 1796) Subgenus Asioplatysma Kryzhanovskij, 1968 * '' Pterostichus capito'' (Tschitscherine, 1900) * '' Pterostichus darvazicus'' Kryzhanovskij, 1968 * '' Pterostichus karateghinicus'' Mikhailov, 1972 * '' Pterostichus klapperichi'' Jedlicka, 1956 * '' Pterostichus medvedevi'' Kryzhanovskij & Mikhailov, 1972 * ''

Pterostichus mujahedeeni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Savich, 1999 * ''

Pterostichus rufopiceus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Heyden, 1890 * ''

Pterostichus talibani ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Savich, 1999 Subgenus Badistrinus Motschulsky, 1866 * '' Pterostichus arrowi'' Jedlicka, 1936 * '' Pterostichus arrowianus'' Jedlicka, 1938 * '' Pterostichus bandotaro'' Tanaka, 1958 * '' Pterostichus doris'' Jedlicka, 1962 * '' Pterostichus haptoderoides'' Tschitscherine, 1889 * '' Pterostichus kajimurai'' Habu & Tanaka, 1957 * '' Pterostichus laticollis'' (Motschulsky, 1844) * ''

Pterostichus procephalus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Bates, 1873 Subgenus Bothriopterus Chaudoir, 1835 * '' Pterostichus ademidovae'' O.Berlov, 1996 * '' Pterostichus adstrictus'' Eschscholtz, 1823 * '' Pterostichus aeneocupreus'' (Fairmaire, 1887) * '' Pterostichus bhunetansis'' Davies, 2004 * '' Pterostichus commixtiformis'' Roubal, 1920 * '' Pterostichus ferghanicus'' Kabak, 2001 * '' Pterostichus lustrans'' LeConte, 1851 * '' Pterostichus mariae'' (Lutshnik, 1921) * ''

Pterostichus mutus ''Pterostichus mutus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in Nor ...

'' (Say, 1823) * '' Pterostichus oblongopunctatus'' (Fabricius, 1787) * ''

Pterostichus oregonus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1861 * ''

Pterostichus pensylvanicus ''Pterostichus pensylvanicus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' LeConte, 1873 * '' Pterostichus quadrifoveolatus'' Letzner, 1852 * ''

Pterostichus subovatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1861) * ''

Pterostichus trinarius ''Pterostichus trinarius'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Casey, 1918) * ''

Pterostichus tropicalis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Bates, 1882) * ''

Pterostichus zyzzovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O. & E.Berlov, 1996 Subgenus Calopterus Chaudoir, 1838 * ''

Pterostichus pilosus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Host, 1790) * ''

Pterostichus selmanni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Duftschmid, 1812) Subgenus Carllindrothius Habu, 1984 * '' Pterostichus colonus'' (Bates, 1883) Subgenus Cheporus Latreille, 1829 * '' Pterostichus burmeisteri'' Heer, 1837 * '' Pterostichus dissimilis'' (A. & G.B.Villa, 1833) * ''

Pterostichus muehlfeldii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Duftschmid, 1812) * ''

Pterostichus transversalis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Duftschmid, 1812) Subgenus Chinapterus O.Berlov, 1998 * ''

Pterostichus lianhuaensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Dorjerem; Shi & Liang, 2020 * ''

Pterostichus liupanensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Dorjerem; Shi & Liang, 2020 * ''

Pterostichus przewalskyi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tschitscherine, 1888 * ''

Pterostichus singularis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tschitscherine, 1889 Subgenus Circinatus Sciaky, 1996 * ''

Pterostichus adelphus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * '' Pterostichus agilis'' Allegro & Sciaky, 2010 * ''

Pterostichus ailaoicus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * '' Pterostichus baenningeri'' Jedlicka, 1931 * ''

Pterostichus batxatensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * '' Pterostichus beneshi'' Sciaky, 1996 * '' Pterostichus bullatus'' Allegro & Sciaky, 2010 * ''

Pterostichus camelus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus caobang ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * ''

Pterostichus cavazzutianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * '' Pterostichus dentifer'' Allegro & Sciaky, 2010 * ''

Pterostichus dimorphus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus expolitus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * ''

Pterostichus hoanglien ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * ''

Pterostichus laocaiensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * '' Pterostichus liciniformis'' Csiki, 1930 * ''

Pterostichus maitreya ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus miao ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus phangxipang ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * ''

Pterostichus phiaoc ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fedorenko, 2019 * ''

Pterostichus pohnerti ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1934 * ''

Pterostichus subtilissimus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sciaky, 1996 * ''

Pterostichus tumulus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus wangjiani ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus xilingensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Allegro & Sciaky, 2010 * ''

Pterostichus yan ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus yuxiaodongi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus zhygealu ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shi & Liang, 2015 * ''

Pterostichus zoiai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sciaky, 1996 Subgenus Cophosus Dejean, 1821 * '' Pterostichus cylindricus'' (Herbst, 1784) Subgenus Cryobius Chaudoir, 1838 * '' Pterostichus abaxoides'' (Dejean, 1828) * '' Pterostichus acrogonus'' (Chaudoir, 1843) * '' Pterostichus akkusianus'' Kirschenhofer, 1981 * '' Pterostichus altaiensis'' (Poppius, 1906) * '' Pterostichus amaroides'' (Dejean, 1828) * '' Pterostichus amblypterus'' (Chaudoir, 1868) * '' Pterostichus amoenus'' (Dejean, 1828) * '' Pterostichus amurensis'' (Poppius, 1906) * '' Pterostichus anatolicus'' Jedlicka, 1963 * '' Pterostichus apenninus'' (Dejean, 1831) * '' Pterostichus aralarensis'' (Mateu, 1945) * '' Pterostichus archangaicus'' Shilenkov, 2000 * '' Pterostichus arcticola'' (Chaudoir, 1868) * '' Pterostichus argutoriformis'' (Poppius, 1906) * '' Pterostichus auriga'' Ball, 1962 * '' Pterostichus bargusinicus'' Shilenkov, 2000 * '' Pterostichus barlensis'' Straneo, 1935 * '' Pterostichus barryorum'' Ball, 1962 * '' Pterostichus biocryus'' Ball, 1962 * '' Pterostichus blandulus'' L.Miller, 1859 * '' Pterostichus blumenthali'' Heinz, 1965 * '' Pterostichus brevicornis'' (Kirby, 1837) * '' Pterostichus breviusculus'' (R.F.Sahlberg, 1844) * '' Pterostichus bryanti'' (Van Dyke, 1951) * '' Pterostichus bryantoides'' Ball, 1962 * '' Pterostichus burjaticus'' (Poppius, 1906) * '' Pterostichus cacumensis'' Ball, 1966 * '' Pterostichus cantabricus'' (L.Schaufuss, 1862) * '' Pterostichus caribou'' Ball, 1962 * '' Pterostichus carradei'' (Gautier des Cottes, 1866) * '' Pterostichus champenoisi'' (Croissandeau, 1893) * '' Pterostichus chipewyan'' Ball, 1962 * '' Pterostichus colasi'' (Jeannel, 1937) * '' Pterostichus convexus'' (Gebler, 1847) * '' Pterostichus coreicus'' Jedlicka, 1962 * '' Pterostichus davshensis'' Shilenkov, 2000 * ''

Pterostichus dubiosus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1894) * '' Pterostichus ehlersi'' (Heyden, 1881) * '' Pterostichus empetricola'' (Dejean, 1828) * '' Pterostichus espanoli'' (J. & E.Vives, 1977) * '' Pterostichus euxinus'' Straneo, 1935 * '' Pterostichus exceptus'' (J.Sahlberg, 1885) * '' Pterostichus firmus'' (Lutshnik, 1933) * '' Pterostichus fulvescens'' (Motschulsky, 1844) * '' Pterostichus gerstlensis'' Ball, 1962 * '' Pterostichus glacialis'' (Brisout de Barneville, 1863) * '' Pterostichus glukhomanka'' Sundukov, 2013 * '' Pterostichus haftorni'' Lindroth, 1969 * '' Pterostichus herzi'' (Poppius, 1906) * '' Pterostichus homalonotus'' (Tschitscherine, 1894) * '' Pterostichus horvatovichi'' Kirschenhofer, 1991 * '' Pterostichus hudsonicus'' LeConte, 1863 * '' Pterostichus infimus'' (Chaudoir, 1868) * '' Pterostichus inopinatus'' (Lutshnik, 1933) * '' Pterostichus iripennis'' (Chaudoir, 1868) * '' Pterostichus ishiharai'' Ishida & Shibata, 1961 * '' Pterostichus jacobsoni'' (Poppius, 1906) * '' Pterostichus jaechi'' Kirschenhofer, 1997 * '' Pterostichus josephi'' Csiki, 1930 * '' Pterostichus kadjaranze'' (Jedlicka, 1947) * '' Pterostichus kaninensis'' (Poppius, 1906) * '' Pterostichus kavanaughi'' Shilenkov, 2000 * '' Pterostichus kaynashensis'' Schweiger, 1967 * '' Pterostichus kolymensis'' Erjiomin, 1998 * '' Pterostichus korgei'' Jedlicka, 1964 * '' Pterostichus kotzebuei'' Ball, 1962 * '' Pterostichus kultianus'' Jedlicka, 1946 * '' Pterostichus kuraicus'' Shilenkov, 2000 * '' Pterostichus kurosai'' Tanaka, 1958 * '' Pterostichus kurosawai'' Tanaka, 1958 * '' Pterostichus lamuticus'' (Poppius, 1906) * '' Pterostichus latiusculus'' (Chaudoir, 1868) * '' Pterostichus longipes'' (Poppius, 1906) * '' Pterostichus lucidus'' (Motschulsky, 1844) * '' Pterostichus macedonicus'' Apfelbeck, 1918 * '' Pterostichus macrothorax'' (Poppius, 1906) * '' Pterostichus maeklini'' (Poppius, 1906) * '' Pterostichus mandibularoides'' Ball, 1966 * '' Pterostichus marani'' Jedlicka, 1967 * '' Pterostichus middendorffi'' (J.Sahlberg, 1875) * ''

Pterostichus negligens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Sturm, 1824) * ''

Pterostichus nemoralis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Graells, 1851) * ''

Pterostichus nigripalpis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Poppius, 1906) * ''

Pterostichus nivalis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (R.F.Sahlberg, 1844) * ''

Pterostichus nordqvisti ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (J.Sahlberg, 1885) * ''

Pterostichus oblongiusculus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1850) * ''

Pterostichus ochoticus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (R.F.Sahlberg, 1844) * ''

Pterostichus palekhai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' E. & O.Berlov, 1996 * ''

Pterostichus parasimilis ''Pterostichus parasimilis'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' Ball, 1962 * ''

Pterostichus parviceps ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Poppius, 1906) * ''

Pterostichus pinguedineus ''Pterostichus pinguedineus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' (Eschscholtz, 1823) * ''

Pterostichus pithyusicus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Lutshnik, 1933) * ''

Pterostichus planus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (J.Sahlberg, 1885) * ''

Pterostichus poppiusianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Jakobson, 1907) * ''

Pterostichus procerulus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Heyden, 1880) * ''

Pterostichus properans ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1868) * ''

Pterostichus pseudostuxbergi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Poppius, 1906) * ''

Pterostichus pumilio ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus punctiger ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (J.Sahlberg, 1880) * ''

Pterostichus pusillus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus quadraticollis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1846) * ''

Pterostichus riparius ''Pterostichus riparius'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Dejean, 1828) * ''

Pterostichus rodiovoni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus rufonitens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Fairmaire, 1866) * ''

Pterostichus safonovae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Budarin, 1995 * ''

Pterostichus sahlbergi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1894) * ''

Pterostichus scitus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Mäklin, 1878) * ''

Pterostichus shilenkovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Erjiomin & Kabak, 1991 * ''

Pterostichus similis ''Pterostichus similis'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in N ...

'' Mannerheim, 1852 * ''

Pterostichus sojot ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus soperi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1966 * ''

Pterostichus splendidus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Park, 2013 * ''

Pterostichus spornyi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sundukov, 2013 * ''

Pterostichus stantonensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1966 * ''

Pterostichus subgibbus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1860) * ''

Pterostichus subiasi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Ortuño & Zaballos, 1992) * ''

Pterostichus subsinuatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus surgens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1878 * ''

Pterostichus tareumiut ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1962 * ''

Pterostichus taskylensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus tatricus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kult, 1947 * ''

Pterostichus theeli ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Mäklin, 1878) * ''

Pterostichus tichomirovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Erjiomin, 1990 * ''

Pterostichus tiliaceoradix ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball, 1962 * ''

Pterostichus tokmakovae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sundukov, 2013 * ''

Pterostichus tunkinensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus udokanensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Shilenkov, 2000 * ''

Pterostichus ulubeyensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kirschenhofer, 1981 * ''

Pterostichus unctulatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Duftschmid, 1812) * ''

Pterostichus ventricosus ''Pterostichus ventricosus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' (Eschscholtz, 1823) * ''

Pterostichus woodi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Ball & Currie, 1997 Subgenus Cylindrocharis Casey, 1918 * '' Pterostichus acutipes'' Barr, 1971 * '' Pterostichus hypogeus'' Barr, 1971 * ''

Pterostichus rostratus ''Pterostichus rostratus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Newman, 1838) Subgenus Eosteropus Tschitscherine, 1902 * '' Pterostichus abensis'' Morita, 2010 * '' Pterostichus abudarini'' O. & E.Berlov, 1997 * '' Pterostichus aenescens'' (Chaudoir, 1850) * '' Pterostichus aereipennis'' (Solsky, 1872) * '' Pterostichus aethiops'' (Panzer, 1796) * '' Pterostichus ainus'' Plutenko, 2005 * '' Pterostichus alacer'' A.Morawitz, 1862 * '' Pterostichus bituberculatus'' (Tschitscherine, 1899) * '' Pterostichus circulosus'' Lindroth, 1966 * '' Pterostichus coruscus'' (Tschitscherine, 1895) * '' Pterostichus creper'' (Tschitscherine, 1902) * '' Pterostichus discrepans'' A.Morawitz, 1862 * '' Pterostichus dudkoi'' Sundukov, 2013 * '' Pterostichus faldermanni'' Schatzmayr, 1929 * '' Pterostichus funakoshii'' Morita, 2007 * '' Pterostichus hidanus'' Morita, 2010 * '' Pterostichus hiramatsui'' Morita, 2010 * '' Pterostichus hirasawai'' Morita, 2010 * '' Pterostichus imurai'' Morita, 2007 * '' Pterostichus japonicus'' (Motschulsky, 1861) * '' Pterostichus karasawai'' Tanaka, 1958 * ''

Pterostichus kimioi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2011 * '' Pterostichus makolskii'' G.Müller, 1933 * '' Pterostichus mandzhuricus'' (Lutshnik, 1916) * '' Pterostichus mannerheimii'' (Dejean, 1831) * ''

Pterostichus matsunagai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2011 * '' Pterostichus maurusiacus'' (Mannerheim, 1825) * '' Pterostichus mizunoyai'' Morita, 2007 * '' Pterostichus moestus'' (Say, 1823) * ''

Pterostichus nasuensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2007 * ''

Pterostichus noburai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2008 * ''

Pterostichus ohkawai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2010 * ''

Pterostichus orientalis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Motschulsky, 1844) * ''

Pterostichus rengensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2010 * ''

Pterostichus rufitarsis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1828) * ''

Pterostichus sakuraii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2008 * ''

Pterostichus shimizui ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2010 * ''

Pterostichus sudai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 1991 * ''

Pterostichus superciliosus ''Pterostichus superciliosus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' (Say, 1823) * ''

Pterostichus tenuimarginatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1868) * ''

Pterostichus tokui ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2007 * ''

Pterostichus tuberculiger ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Tschitscherine, 1897) * ''

Pterostichus virescens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Gebler, 1833) * ''

Pterostichus yoshizawai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita, 2010 Subgenus Ethira Andrewes, 1936 * '' Pterostichus cometes'' (Andrewes, 1936) * '' Pterostichus depilatus'' (Bates, 1889) * '' Pterostichus heinzianus'' Sciaky, 1996 * ''

Pterostichus multiseta ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * ''

Pterostichus piliferus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Bates, 1878) * ''

Pterostichus pseudopilifer ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1957) * ''

Pterostichus rugiceps ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * ''

Pterostichus seticeps ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * ''

Pterostichus sharanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * ''

Pterostichus variseta ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1984) * ''

Pterostichus viridellus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Straneo, 1984 Subgenus Falsargutor Kryzhanovskij, 1983 * '' Pterostichus kataevi'' Kryzhanovskij, 1989 * ''

Pterostichus ponticus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kirschenhofer, 1987 * ''

Pterostichus pseudopedius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Reitter, 1887 Subgenus Feronidius Jeannel, 1942 * '' Pterostichus brevipennis'' (Chevrolat, 1840) * '' Pterostichus fornicatus'' (Kolenati, 1845) * '' Pterostichus hungaricus'' (Dejean, 1828) * '' Pterostichus incommodus'' Schaum, 1858 * '' Pterostichus melas'' (Creutzer, 1799) Subgenus Feronina Casey, 1918 * '' Pterostichus barri'' Bousquet, 2006 * ''

Pterostichus palmi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Schaeffer, 1910 Subgenus Fukienostichus Lafer, 1983 * ''

Pterostichus shaovuensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1956 Subgenus Gastrosticta Casey, 1918 * '' Pterostichus enodis'' Bousquet, 1992 * ''

Pterostichus mutoides ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Bousquet, 1992 * ''

Pterostichus obesulus ''Pterostichus obesulus'' is a species of beetle first discovered in 1873. No sub-species are listed in the Catalogue of Life The Catalogue of Life (CoL) is an online database that provides an index of known species of animals, plants, fungi, ...

'' LeConte, 1873 * ''

Pterostichus ophryoderus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1878) * ''

Pterostichus punctiventris ''Pterostichus punctiventris'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are foun ...

'' (Chaudoir, 1878) * ''

Pterostichus putus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus sayanus ''Pterostichus sayanus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in N ...

'' Csiki, 1930 * ''

Pterostichus subacutus ''Pterostichus subacutus'' is a species of woodland ground beetle in the family Carabidae which is found in North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. N ...

'' (Casey, 1918) * ''

Pterostichus tumescens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1863 * ''

Pterostichus ventralis ''Pterostichus ventralis'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Say, 1823) Subgenus Georgeballius Habu, 1984 * '' Pterostichus hoplites'' (Bates, 1883) Subgenus Gutta Wrase & J.Schmidt, 2006 * '' Pterostichus adulterinus'' Wrase & J.Schmidt, 2006 * '' Pterostichus gaoligongensis'' Wrase & J.Schmidt, 2006 * ''

Pterostichus kongshuhensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' B.Gueorguiev, 2015 * ''

Pterostichus phungaraziensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Wrase & J.Schmidt, 2006 Subgenus Haplomaseus Reitter, 1896 * '' Pterostichus abagonensis'' Reitter, 1896 * '' Pterostichus andreae'' (Tschitscherine, 1897) * '' Pterostichus arator'' (Faldermann, 1836) * '' Pterostichus armenus'' (Faldermann, 1836) * '' Pterostichus astutus'' (Tschitscherine, 1903) * '' Pterostichus balikliensis'' Hovorka & Skoupy, 2007 * '' Pterostichus caucasicola'' (Tschitscherine, 1893) * '' Pterostichus caucasicus'' (Ménétriés, 1832) * '' Pterostichus chasautianus'' Vysoky, 1984 * '' Pterostichus chydaeus'' (Tschitscherine, 1897) * '' Pterostichus consanguineus'' (Chaudoir, 1878) * '' Pterostichus cristicaudis'' Kurnakov, 1962 * '' Pterostichus denticaudis'' Kurnakov, 1962 * '' Pterostichus giresuni'' Jedlicka, 1965 * '' Pterostichus goriensis'' (Tschitscherine, 1897) * '' Pterostichus heinzi'' Jedlicka, 1965 * '' Pterostichus ikizderensis'' Hovorka & Skoupy, 2007 * '' Pterostichus jugicola'' (Lutshnik, 1916) * ''

Pterostichus krulikovskyi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Lutshnik, 1933) * '' Pterostichus meskheticus'' Belousov, 1991 * ''

Pterostichus novotnyorum ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Vysoky, 1981 * ''

Pterostichus olegi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Belousov, 1991 * ''

Pterostichus phaeus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Lutshnik, 1922) * ''

Pterostichus porcellus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kurnakov, 1962 * ''

Pterostichus pseudoastutus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hovorka & Skoupy, 2007 * ''

Pterostichus rousi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kirschenhofer, 1982 * ''

Pterostichus skoupyi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hovorka & Skoupy, 2007 * ''

Pterostichus staveni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hovorka & Skoupy, 2007 * ''

Pterostichus tamsii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Dejean, 1831) * ''

Pterostichus tanjae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hovorka & Skoupy, 2007 * ''

Pterostichus woronowi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Lutshnik, 1916) Subgenus Haptotapinus Reitter, 1886 * '' Pterostichus aksekianus'' (Straneo, 1988) * '' Pterostichus besucheti'' (Straneo, 1988) * '' Pterostichus bythiniensis'' (Maran, 1944) * '' Pterostichus crassiusculus'' (Chaudoir, 1868) * '' Pterostichus dipojranus'' (Straneo, 1988) * '' Pterostichus matchai'' (Jedlicka, 1930) * ''

Pterostichus oblongoparallelus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Maran, 1932 * ''

Pterostichus odontocnemis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' G.Müller, 1931 * ''

Pterostichus orduensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1988) * ''

Pterostichus pavani ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Straneo, 1988) * ''

Pterostichus reissi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' G.Müller, 1931 * ''

Pterostichus vseteckai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Maran, 1944) Subgenus Huaius Tian; Huang; Chen & Ding, 2019 * ''

Pterostichus caoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tian; Huang; Chen & Ding, 2019 * ''

Pterostichus hanwang ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tian & He, 2020 * ''

Pterostichus tiankeng ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tian; Huang; Chen & Ding, 2019 * ''

Pterostichus yuae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Tian; Huang; Chen & Ding, 2019 Subgenus Hypherpes Chaudoir, 1838 * '' Pterostichus adoxus'' (Say, 1823) * '' Pterostichus algidus'' LeConte, 1853 * '' Pterostichus amethystinus'' Mannerheim, 1843 * '' Pterostichus annosus'' Casey, 1913 * '' Pterostichus arcanus'' Casey, 1913 * '' Pterostichus baldwini'' (Casey, 1924) * '' Pterostichus barbarinus'' Casey, 1913 * '' Pterostichus brachylobus'' Kavanaugh & LaBonte, 2006 * ''

Pterostichus californicus ''Pterostichus californicus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' (Dejean, 1828) * '' Pterostichus canallatus'' Casey, 1913 * '' Pterostichus castaneus'' (Dejean, 1828) * '' Pterostichus castanipes'' (Ménétriés, 1843) * ''

Pterostichus congestus ''Pterostichus congestus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in ...

'' (Ménétriés, 1843) * '' Pterostichus craterensis'' (Hatch, 1949) * '' Pterostichus crenicollis'' LeConte, 1873 * '' Pterostichus ecarinatus'' Hatch, 1936 * '' Pterostichus esuriens'' Casey, 1913 * '' Pterostichus gliscans'' Casey, 1913 * '' Pterostichus gracilior'' LeConte, 1873 * '' Pterostichus herculaneus'' Mannerheim, 1843 * '' Pterostichus hornii'' LeConte, 1873 * '' Pterostichus illustris'' LeConte, 1851 * '' Pterostichus inermis'' Fall, 1901 * '' Pterostichus isabellae'' LeConte, 1851 * '' Pterostichus jacobinus'' Casey, 1913 * '' Pterostichus laborans'' Casey, 1913 * '' Pterostichus lacertus'' Casey, 1913 * '' Pterostichus lama'' (Ménétriés, 1843) * '' Pterostichus lassulus'' (Casey, 1920) * '' Pterostichus lattini'' LaBonte, 2006 * '' Pterostichus luscus'' (Casey, 1918) * '' Pterostichus menetriesii'' LeConte, 1873 * '' Pterostichus mercedianus'' (Casey, 1918) * ''

Pterostichus miscellus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * '' Pterostichus morionides'' (Chaudoir, 1868) * ''

Pterostichus neobrunneus ''Pterostichus neobrunneus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found ...

'' Lindroth, 1966 * ''

Pterostichus nigrocaeruleus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Van Dyke, 1926 * ''

Pterostichus obsidianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus occultus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus ordinarius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus ovalipennis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus panticulatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus pergracilis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Casey, 1920) * ''

Pterostichus planctus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1853 * ''

Pterostichus protensiformis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Casey, 1924) * ''

Pterostichus protractus ''Pterostichus protractus'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found i ...

'' LeConte, 1860 * ''

Pterostichus restrictus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Casey, 1918) * ''

Pterostichus scutellaris ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1873 * ''

Pterostichus sejungendus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (Chaudoir, 1868) * ''

Pterostichus serripes ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' (LeConte, 1875) * ''

Pterostichus setosus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hatch, 1951 * ''

Pterostichus sierranus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus sponsor ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus spraguei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' LeConte, 1873 * ''

Pterostichus suffusus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Casey, 1913 * ''

Pterostichus tarsalis ''Pterostichus tarsalis'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. References Further reading

* Pterostichus, tarsalis Articles created by Qbugbot Beetles described in 1873 {{Pte ...

'' LeConte, 1873 * ''

Pterostichus tristis ''Pterostichus tristis'' is a species of woodland ground beetle in the family Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in N ...

'' (Dejean, 1828) * ''

Pterostichus tuberculofemoratus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Hatch, 1936 * ''

Pterostichus vandykei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Schaeffer, 1910 * ''

Pterostichus vicinus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Mannerheim, 1843 * ''

Pterostichus ybousqueti ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' O.Berlov, 1999 Subgenus Jedlickaia Sciaky, 1997 * ''

Pterostichus sterbai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Jedlicka, 1934 Subgenus Koreonialoe Park & Kwon, 1996 * '' Pterostichus adatarasanus'' Sasakawa, 2005 * '' Pterostichus akitai'' Morita, 2004 * '' Pterostichus apiculatiphallus'' Nemoto, 1988 * '' Pterostichus asahinus'' Habu & Baba, 1960 * '' Pterostichus bellatrix'' (Tschitscherine, 1895) * '' Pterostichus bifidiphallus'' Nemoto, 1988 * '' Pterostichus chokaisanus'' Sasakawa, 2009 * '' Pterostichus eboshiyamanus'' Sasakawa, 2009 * '' Pterostichus falcispinus'' Sasakawa, 2005 * '' Pterostichus fukube'' Sugimura, 2005 * '' Pterostichus gassanus'' Sasakawa, 2009 * ''

Pterostichus gujoensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Toda, 2012 * '' Pterostichus ishikawai'' Nemoto, 1988 * '' Pterostichus ishikawaioides'' Sasakawa; J.L.Kim; J.K.Kim & Kubota, 2008 * '' Pterostichus isolatus'' Sasakawa; Kim & Kubota, 2005 * '' Pterostichus iwakiensis'' Sasakawa, 2009 * ''

Pterostichus iwasakii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Kasahara, 1990 * '' Pterostichus ixion'' (Tschitscherine, 1902) * '' Pterostichus jiricola'' Sasakawa; J.L.Kim; J.K.Kim & Kubota, 2008 * '' Pterostichus kitakamisanus'' Sasakawa, 2005 * '' Pterostichus koheii'' Nakane, 1963 * ''

Pterostichus kuraiyamanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita & Ohkawa, 2010 * '' Pterostichus kurikomasanus'' Sasakawa, 2005 * '' Pterostichus kuwolsanensis'' Li & Zhang, 2014 * '' Pterostichus macrogenys'' Bates, 1883 * '' Pterostichus microps'' Heyden, 1887 * ''

Pterostichus miyazawai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita & Ohkawa, 2009 * ''

Pterostichus momuranus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Morita; Ohkawa & Kurihara, 2013 * ''

Pterostichus monolineatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,200 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Sasakawa; Mitsuduka & H.Ito, 2020 * ''Pterostichus nagasawai'' N.Ito & Ogai, 2015 * ''Pterostichus ohsawacavus'' Sasakawa, 2005 * ''Pterostichus opacipennis'' Jedlicka, 1934 * ''Pterostichus palgongsanus'' Nemoto, 1988 * ''Pterostichus shikatai'' Toda, 2012 * ''Pterostichus shinbodakensis'' Sasakawa & H.Ito, 2017 * ''Pterostichus shirakamisan'' Sasakawa, 2009 * ''Pterostichus shirakamisanus'' Sasakawa, 2005 * ''Pterostichus shojii'' Sugimura, 2006 * ''Pterostichus sumondakensis'' Sasakawa, 2005 * ''Pterostichus syleus'' Kirschenhofer, 1997 * ''Pterostichus taebaegsanus'' Nemoto, 1988 * ''Pterostichus takadateyamanus'' Sasakawa, 2009 * ''Pterostichus tanakai'' Ishida, 1964 * ''Pterostichus tateishiyamanus'' Sasakawa & H.Ito, 2017 * ''Pterostichus teretis'' Park & Kwon, 1996 * ''Pterostichus todai'' Morita & Kanie, 1997 * ''Pterostichus togyusanus'' Park & Kwon, 1996 * ''Pterostichus toyodai'' Morita & Kurosa, 1998 * ''Pterostichus uedaorum'' Morita & Hirasawa, 1996 * ''

'' Park & Kwon, 1996 * ''Pterostichus woongbii'' Park & Kwon, 1996 * ''Pterostichus yahikosanus'' Sasakawa, 2009 * ''Pterostichus yamizosanus'' Sasakawa, 2005 * ''Pterostichus yorikoae'' Sugimura, 2007 Subgenus Kozyrevius O.Berlov, 1998 * ''Pterostichus militaris'' (Tschitscherine, 1893) Subgenus Lamenius Bousquet, 1999 * ''Pterostichus caudicalis'' (Say, 1823) Subgenus Lenapterus O.Berlov, 1996 * ''Pterostichus agonus'' G.Horn, 1880 * ''Pterostichus cancellatus'' (Motschulsky, 1860) * ''Pterostichus costatus'' (Ménétriés, 1851) * ''Pterostichus galae'' Farkac & Plutenko, 1996 * ''Pterostichus gromykoi'' Sundukov, 2005 * ''Pterostichus marginatus'' Matsumura, 1911 * ''Pterostichus punctatissimus'' (Randall, 1838) * ''Pterostichus rugosipennis'' Jedlicka, 1932 * ''Pterostichus saxicola'' (Tschitscherine, 1899) * ''Pterostichus vermiculosus'' (Ménétriés, 1851) Subgenus Leptoferonia Casey, 1918 * ''Pterostichus angustus'' (Dejean, 1828) * ''Pterostichus beyeri'' Van Dyke, 1926 * ''Pterostichus blodgettensis'' Will, 2007 * ''Pterostichus caligans'' G.Horn, 1891 * ''Pterostichus cochlearis'' Hacker, 1968 * ''Pterostichus deino'' Will, 2007 * ''Pterostichus enyo'' Will, 2007 * ''Pterostichus falli'' Van Dyke, 1926 * ''Pterostichus fenyesi'' Csiki, 1930 * ''Pterostichus fuchsi'' Schaeffer, 1910 * ''Pterostichus hackerae'' Will, 2016 * ''Pterostichus hatchi'' Hacker, 1968 * ''Pterostichus humilis'' Casey, 1913 * ''Pterostichus idahoae'' Csiki, 1930 * ''Pterostichus inanis'' G.Horn, 1891 * ''Pterostichus infernalis'' Hatch, 1936 * ''Pterostichus inopinus'' (Casey, 1918) * ''Pterostichus lobatus'' Hacker, 1968 * ''Pterostichus marinensis'' Hacker, 1968 * ''Pterostichus mattolensis'' Hacker, 1968 * ''Pterostichus pemphredo'' Will, 2007 * ''Pterostichus pumilus'' Casey, 1913 * ''Pterostichus rothi'' (Hatch, 1951) * ''Pterostichus sphodrinus'' LeConte, 1863 * ''Pterostichus stapedius'' Hacker, 1968 * ''Pterostichus trinitensis'' Hacker, 1968 * ''Pterostichus yosemitensis'' Hacker, 1968 Subgenus Lianoe Gozis, 1882 * ''Pterostichus amanoi'' Nakane, 1968 * ''Pterostichus arribasi'' Ortuño, 1991 * ''Pterostichus asturicus'' (Jeanne, 1966) * ''Pterostichus cordatissimus'' (Straneo, 1937) * ''Pterostichus drescoi'' Nègre, 1957 * ''Pterostichus dufourii'' (Dejean, 1828) * ''Pterostichus microphthalmus'' Delarouzée, 1857 * ''Pterostichus muelleri'' (Straneo, 1936) * ''Pterostichus nadari'' (Vuillefroy, 1893) * ''Pterostichus nakamiyorinus'' Morita; Ohkawa & Kurihara, 2013 * ''Pterostichus rousselli'' Colas, 1963 * ''Pterostichus sakagamii'' Morita, 2001 * ''Pterostichus sudrei'' Sainte-Claire Deville, 1922 Subgenus Lyrothorax Chaudoir, 1838 * ''Pterostichus amagisanus'' Tanaka & Ishida, 1972 * ''Pterostichus caspius'' (Ménétriés, 1832) * ''Pterostichus eoyoritomus'' Sasakawa, 2009 * ''Pterostichus fujitai'' Tanaka & Ishida, 1972 * ''Pterostichus yoritomus'' Bates, 1873 Subgenus Megabea Sciaky, 1997 * ''Pterostichus tienmushanus'' Sciaky, 1997 Subgenus Melanius Bonelli, 1810 * ''Pterostichus aterrimus'' (Herbst, 1784) * ''Pterostichus castor'' Goulet & Bousquet, 1983 * ''Pterostichus corvinus'' (Dejean, 1828) * ''Pterostichus ebeninus'' (Dejean, 1828) * ''Pterostichus elongatus'' (Duftschmid, 1812) * ''Pterostichus licenti'' Jedlicka, 1939 * ''Pterostichus noguchii'' Bates, 1873 Subgenus Metallophilus Chaudoir, 1838 * ''Pterostichus dandongensis'' Kirschenhofer, 1997 * ''Pterostichus interruptus'' (Dejean, 1828) * ''Pterostichus kamtschaticus'' Motschulsky, 1860 * ''Pterostichus mirus'' (Tschitscherine, 1894) * ''Pterostichus orion'' (Tschitscherine, 1901) * ''Pterostichus pfizenmayeri'' Poppius, 1906 * ''Pterostichus rugosus'' (Gebler, 1823) * ''Pterostichus sublaevis'' (J.Sahlberg, 1880) Subgenus Micronialoe Park; Kwon & Lafer, 1996 * ''Pterostichus bifoveolatus'' Park; Kwon & Lafer, 1996 * ''Pterostichus chogyesanus'' Park; Kwon & Lafer, 1996 * ''Pterostichus kaniei'' Morita, 2005 Subgenus Monoferonia Casey, 1918 * ''Pterostichus carolinus'' Darlington, 1932 * ''Pterostichus diligendus'' (Chaudoir, 1868) * ''Pterostichus mancus'' (LeConte, 1853) * ''Pterostichus primus'' Darlington, 1932 Subgenus Morphohaptoderus Tschitscherine, 1898 * ''Pterostichus bowanus'' Sciaky, 1997 * ''Pterostichus cervenkai'' Sciaky, 1994 * ''Pterostichus chungkingi'' Jedlicka, 1932 * ''Pterostichus confucius'' Sciaky & Wrase, 1997 * ''Pterostichus dentellus'' Facchini & Sciaky, 2003 * ''Pterostichus dundai'' Sciaky, 1994 * ''Pterostichus emei'' Sciaky, 1994 * ''Pterostichus expeditus'' (Tschitscherine, 1898) * ''Pterostichus geberti'' Sciaky & Wrase, 1997 * ''Pterostichus giacomazzoi'' Sciaky, 1994 * ''Pterostichus gongga'' Sciaky, 1997 * ''Pterostichus guizhouensis'' Sciaky, 1997 * ''Pterostichus huashanus'' Sciaky, 1994 * ''Pterostichus hubeicus'' Facchini & Sciaky, 2003 * ''Pterostichus irideus'' Sciaky, 1994 * ''Pterostichus janatai'' Sciaky & Wrase, 1997 * ''Pterostichus kalabi'' Sciaky, 1994 * ''Pterostichus kucerai'' Sciaky, 1997 * ''Pterostichus lingshanus'' Sciaky & Wrase, 1997 * ''Pterostichus maximus'' Tschitscherine, 1889 * ''Pterostichus megaloderus'' Sciaky, 1994 * ''Pterostichus ming'' Sciaky & Wrase, 1997 * ''Pterostichus miroslavi'' Sciaky & Wrase, 1997 * ''Pterostichus muellermotzfeldi'' Wrase & J.Schmidt, 2006 * ''Pterostichus parvicollis'' Sciaky & Wrase, 1997 * ''Pterostichus pseudoplatyderus'' Sciaky, 1994 * ''Pterostichus saueri'' Sciaky, 1994 * ''Pterostichus schuelkei'' Sciaky & Wrase, 1997 * ''Pterostichus shennongjianus'' Facchini & Sciaky, 2003 * ''Pterostichus straneellus'' Jedlicka, 1938 * ''Pterostichus toledanoi'' Facchini & Sciaky, 2003 * ''Pterostichus wenxianensis'' Allegro & Sciaky, 2010 * ''Pterostichus yulongshanensis'' Sciaky, 1997 Subgenus Myosodus Fischer von Waldheim, 1823 * ''Pterostichus aapsorum'' Belousov, 1991 * ''Pterostichus aibgensis'' Starck, 1890 * ''Pterostichus avaricus'' Abdurakhmanov & Kryzhanovskij, 1983 * ''Pterostichus batesi'' (Tschitscherine, 1894) * ''Pterostichus buglaniensis'' Kirschenhofer, 1981 * ''Pterostichus filipjevi'' (Lutshnik, 1928) * ''Pterostichus ingusha'' (Lutshnik, 1928) * ''Pterostichus kiritshenkoi'' (Lutshnik, 1928) * ''Pterostichus lacunosus'' (Chaudoir, 1844) * ''Pterostichus lutshnikianus'' Bogachev & Kurnakov, 1958 * ''Pterostichus nivicola'' (Ménétriés, 1832) * ''Pterostichus ordinatus'' (Fischer von Waldheim, 1823) * ''Pterostichus rudestriatus'' (Reitter, 1883) * ''Pterostichus schoenherri'' Faldermann, 1836 * ''Pterostichus sodalicius'' Heyden, 1885 * ''Pterostichus starcki'' Heyden, 1885 * ''Pterostichus stoeckleini'' (Straneo, 1941) * ''Pterostichus svanicus'' (Lutshnik, 1922) * ''Pterostichus swaneticus'' (Reitter, 1883) * ''Pterostichus variabilis'' (Ménétriés, 1832) * ''Pterostichus zamotajlovi'' Belousov, 1991 Subgenus Neohaptoderus Tschitscherine, 1898 * ''Pterostichus berezowskii'' (Tschitscherine, 1898) * ''Pterostichus catei'' Sciaky & Wrase, 1997 * ''Pterostichus comorus'' Jedlicka, 1932 * ''Pterostichus gravis'' Jedlicka, 1939 * ''Pterostichus haesitatus'' Fairmaire, 1889 * ''Pterostichus ignavus'' (Tschitscherine, 1897) * ''Pterostichus jureceki'' (Jedlicka, 1936) * ''Pterostichus kleinfeldianus'' Sciaky & Wrase, 1997 * ''Pterostichus komalus'' (Jedlicka, 1936) * ''Pterostichus maderi'' Jedlicka, 1938 * ''Pterostichus molopsoides'' Jedlicka, 1934 * ''Pterostichus montigena'' (Tschitscherine, 1898) * ''Pterostichus mundus'' Jedlicka, 1938 * ''Pterostichus oreophilus'' (Tschitscherine, 1898) * ''Pterostichus orestes'' (Jedlicka, 1936) * ''Pterostichus sinicus'' (Tschitscherine, 1897) * ''Pterostichus szetschuanensis'' Tschitscherine, 1889 * ''Pterostichus yunnanensis'' Jedlicka, 1934 Subgenus Nialoe Tanaka, 1958 * ''Pterostichus asymmetricus'' Bates, 1883 * ''Pterostichus basilobatus'' Sasakawa, 2005 * ''Pterostichus biexcisus'' Straneo, 1955 * ''Pterostichus bisetosus'' (Straneo, 1938) * ''Pterostichus brunneipennis'' Straneo, 1955 * ''Pterostichus carsticus'' Kasahara & Y.Ito, 1989 * ''Pterostichus chujoi'' Habu, 1959 * ''Pterostichus cristatoides'' Straneo, 1955 * ''Pterostichus daihizanus'' Ishida, 1968 * ''Pterostichus daisenicus'' Ishida, 1958 * ''Pterostichus dandonis'' Kasahara, 1989 * ''Pterostichus enasanus'' Morita, 2007 * ''Pterostichus fujimurai'' Habu, 1958 * ''Pterostichus fujisanus'' Tanaka & Suga, 1972 * ''Pterostichus fumikoae'' Kurosa, 2019 * ''Pterostichus hakusanus'' Kasahara, 1989 * ''Pterostichus hekosanensis'' Sasakawa, 2020 * ''Pterostichus himifuriho'' Morita, 2007 * ''Pterostichus hiraii'' Morita, 2015 * ''Pterostichus hirobane'' Habu & Baba, 1958 * ''Pterostichus hozumii'' Ishida, 1961 * ''Pterostichus ikukoae'' Morita, 2007 * ''Pterostichus ishizukai'' Kasahara, 1995 * ''Pterostichus isumiensis'' Kasahara & Saito, 1997 * ''Pterostichus janoi'' Jedlicka, 1952 * ''Pterostichus jogaesanensis'' Lafer; Paik & Park, 1996 * ''Pterostichus katashinensis'' Habu, 1958 * ''Pterostichus kaya'' Morita, 2019 * ''Pterostichus kongosanus'' Nakane, 1963 * ''Pterostichus latistylis'' Tanaka, 1958 * ''Pterostichus masahiroi'' Kasahara, 1988 * ''Pterostichus masatakai'' Morita, 2007 * ''Pterostichus mirificus'' Bates, 1883 * ''Pterostichus mitoyamanus'' Tanaka, 1971 * ''Pterostichus musashiensis'' Kasahara, 1993 * ''Pterostichus nakanei'' Straneo, 1955 * ''Pterostichus naokii'' Morita, 2003 * ''Pterostichus napaea'' Kasahara, 1988 * ''Pterostichus nishiyamai'' Kasahara, 1986 * ''Pterostichus ogaensis'' Morita, 1995 * ''Pterostichus ohbayashii'' Ishida, 1968 * ''Pterostichus ohdaisanus'' Nakane, 1963 * ''Pterostichus ohkurai'' Morita, 1996 * ''Pterostichus okutamae'' Tanaka, 1963 * ''Pterostichus omogoensis'' Nakane, 1972 * ''Pterostichus ompoensis'' Jedlicka, 1932 * ''Pterostichus opaculus'' N.Ito, 2010 * ''Pterostichus ovaliphallus'' Sasakawa, 2005 * ''Pterostichus parkwon'' Davies, 2004 * ''Pterostichus praedo'' (Tschitscherine, 1901) * ''Pterostichus rhanis'' (Tschitscherine, 1902) * ''Pterostichus ryoheii'' Morita, 2012 * ''Pterostichus ryomoensis'' Morita & Suda, 2007 * ''Pterostichus satoi'' Ishida, 1961 * ''Pterostichus shibatai'' Ishida, 1961 * ''Pterostichus shotaroi'' Morita, 1987 * ''Pterostichus sincerus'' Park & Kwon, 1996 * ''Pterostichus spiculifer'' Bates, 1883 * ''Pterostichus sugimurai'' Morita, 2007 * ''Pterostichus tahirai'' Kasahara, 1992 * ''Pterostichus takahashii'' Ishida, 1958 * ''Pterostichus tanakaorum'' Morita & Ohkura, 1988 * ''Pterostichus taoi'' Kasahara, 1993 * ''Pterostichus tokejii'' Yoshida & Tanaka, 1960 * ''Pterostichus tottoriensis'' Morita, 2003 * ''Pterostichus tsurugiyamanus'' Habu, 1959 * ''Pterostichus uchiyamai'' Morita, 1987 * ''Pterostichus uenoi'' Straneo, 1955 * ''Pterostichus watanabei'' Nakane, 1960 * ''Pterostichus yamashitai'' Morita, 2019 * ''Pterostichus yanoi'' Jedlicka, 1953 * ''Pterostichus yatsuensis'' Straneo, 1955 * ''Pterostichus yokohamae'' Straneo in Nakane, 1979 Subgenus Oreolyperus Tschitscherine, 1901 * ''Pterostichus heptapotamicus'' (Lutshnik, 1927) * ''Pterostichus korolkowi'' (Tschitscherine, 1901) * ''Pterostichus necessarius'' (Tschitscherine, 1894) * ''Pterostichus regeli'' (Tschitscherine, 1894) Subgenus Oreophilus Chaudoir, 1838 * ''Pterostichus alberti'' (Jeannel, 1942) * ''Pterostichus cribratus'' (Dejean, 1828) * ''Pterostichus dubius'' Heer, 1838 * ''Pterostichus duratii'' A. & G.B.Villa, 1835 * ''Pterostichus externepunctatus'' (Dejean, 1828) * ''Pterostichus flavofemoratus'' (Dejean, 1828) * ''Pterostichus franzi'' Nègre, 1955 * ''Pterostichus ienishteai'' Nitzu, 1988 * ''Pterostichus impressus'' (Fairmaire & Laboulbène, 1854) * ''Pterostichus jurinei'' (Panzer, 1802) * ''Pterostichus morio'' (Duftschmid, 1812) * ''Pterostichus multipunctatus'' (Dejean, 1828) * ''Pterostichus parnassius'' Schaum, 1859 * ''Pterostichus paulini'' (Vuillefroy, 1868) * ''Pterostichus planiusculus'' (Chaudoir, 1859) * ''Pterostichus rufipennis'' Baudi di Selve, 1889 * ''Pterostichus spinolae'' (Dejean, 1828) * ''Pterostichus variolatus'' (Dejean, 1828) * ''Pterostichus xatartii'' (Dejean, 1828) * ''Pterostichus yvanii'' (Dejean, 1828) Subgenus Oreoplatysma Jakobson, 1907 * ''Pterostichus abishirensis'' Belousov, 1991 * ''Pterostichus araraticus'' Kirschenhofer, 1987 * ''Pterostichus asinubas'' Davies, 2004 * ''Pterostichus belizini'' (Lutshnik, 1933) * ''Pterostichus belousovi'' Kryzhanovskij, 1989 * ''Pterostichus borcka'' Jedlicka, 1963 * ''Pterostichus capitolinus'' Kurnakov, 1962 * ''Pterostichus casaleianus'' Kirschenhofer, 1991 * ''Pterostichus cecchiniae'' (Jakobson, 1907) * ''Pterostichus chefsuricus'' Reitter, 1896 * ''Pterostichus colchicus'' (Chaudoir, 1850) * ''Pterostichus cordifer'' Reitter, 1896 * ''Pterostichus cunibakus'' Davies, 2004 * ''Pterostichus daghestanus'' Reitter, 1896 * ''Pterostichus depressidorsis'' Reitter, 1896 * ''Pterostichus faunus'' Kurnakov, 1962 * ''Pterostichus fencli'' Dvorak, 1995 * ''Pterostichus ghilarovi'' Kryzhanovskij, 1988 * ''Pterostichus jakobsonianus'' (Lutshnik, 1928) * ''Pterostichus kadleci'' Dvorak, 1995 * ''Pterostichus kirschenblatti'' Kryzhanovskij, 1988 * ''Pterostichus koenigi'' (Reitter, 1887) * ''Pterostichus krasnopolensis'' Kirschenhofer, 1987 * ''Pterostichus kvirensis'' Belousov, 1991 * ''Pterostichus lodosi'' Heinz, 1977 * ''Pterostichus mediocris'' Dvorak, 1995 * ''Pterostichus nikodymi'' Dvorak, 1995 * ''Pterostichus odvarkai'' Dvorak, 1995 * ''Pterostichus orman'' Jedlicka, 1963 * ''Pterostichus pasanauricus'' Kirschenhofer, 1987 * ''Pterostichus percontator'' Reitter, 1887 * ''Pterostichus planaticollis'' Kirschenhofer, 1987 * ''Pterostichus platyderus'' (Chaudoir, 1850) * ''Pterostichus pulchellus'' (Faldermann, 1836) * ''Pterostichus rousianus'' Kirschenhofer, 1987 * ''Pterostichus rubripalpis'' Csiki, 1930 * ''Pterostichus satunini'' (Tschitscherine, 1903) * ''Pterostichus satyrus'' Kurnakov, 1962 * ''Pterostichus schodaicus'' Kirschenhofer, 1987 * ''Pterostichus setosicus'' Davies, 2004 * ''Pterostichus sojaki'' Kirschenhofer, 1987 * ''Pterostichus stomoides'' (Chaudoir, 1868) * ''Pterostichus strasseri'' Reitter, 1898 * ''Pterostichus tamarae'' Wrase & Kirschenhofer, 1991 * ''Pterostichus validiceps'' Reitter, 1887 * ''Pterostichus zolotarewi'' Reitter, 1911 Subgenus Orientostichus Sciaky & Allegro, 2013 * ''Pterostichus chinensis'' (Jedlicka, 1962) * ''Pterostichus curtatus'' Fairmaire, 1886 * ''Pterostichus davidi'' (Tschitscherine, 1897) * ''Pterostichus deceptrix'' (Tschitscherine, 1898) * ''Pterostichus distinctissimus'' Jedlicka, 1940 * ''Pterostichus ferreroi'' Straneo, 1989 * ''Pterostichus gallopavo'' Sciaky & Wrase, 1997 * ''Pterostichus lesticoides'' (Straneo, 1939) * ''Pterostichus machulkai'' Jedlicka, 1931 * ''Pterostichus perlutus'' Jedlicka, 1938 * ''Pterostichus prattii'' Bates, 1890 * ''Pterostichus pulcher'' Sciaky & Allegro, 2013 * ''Pterostichus semirugosus'' (Andrewes, 1947) * ''Pterostichus simillimus'' Fairmaire, 1886 * ''Pterostichus tachongi'' Jedlicka, 1936 Subgenus Orsonjohnsonus Hatch, 1933 * ''Pterostichus johnsoni'' Ulke, 1889 Subgenus Paraferonia Casey, 1918 * ''Pterostichus lubricus'' LeConte, 1853 Subgenus Parahaptoderus Jeanne, 1969 * ''Pterostichus bielzii'' (Fuss, 1858) * ''Pterostichus brevis'' (Duftschmid, 1812) * ''Pterostichus vecors'' (Tschitscherine, 1897) Subgenus Paralianoe Ishida, 1958 * ''Pterostichus kiiensis'' Morita & Ohkura, 1988 Subgenus Parapterostichus Desbrochers des Loges, 1906 * ''Pterostichus grajus'' (Dejean, 1828) * ''Pterostichus justusii'' W.Redtenbacher, 1842 * ''Pterostichus nodicornis'' (Fairmaire & Laboulbène, 1854) * ''Pterostichus schaschli'' (Marseul, 1880) Subgenus Petrophilus Chaudoir, 1838 * ''Pterostichus abnormis'' (J.Sahlberg, 1880) * ''Pterostichus acutidens'' (Fairmaire, 1888) * ''Pterostichus alexandrovi'' Lafer, 1979 * ''Pterostichus alexeji'' Zamotajlov & Kryzhanovskij, 1992 * ''Pterostichus altaianus'' Jedlicka, 1958 * ''Pterostichus altaicus'' (Germar, 1823) * ''Pterostichus arsenjevi'' Lafer, 1979 * ''Pterostichus bungei'' (Tschitscherine, 1894) * ''Pterostichus calvitarsis'' Breit, 1912 * ''Pterostichus chechcirensis'' Lafer, 1979 * ''Pterostichus chinkiangensis'' Kirschenhofer, 1991 * ''Pterostichus coracinus'' (Newman, 1838) * ''Pterostichus dauricus'' (Gebler, 1832) * ''Pterostichus dilutipes'' (Motschulsky, 1844) * ''Pterostichus elmbergi'' Poppius, 1908 * ''Pterostichus eximius'' A.Morawitz, 1862 * ''Pterostichus fallettii'' Sciaky, 1996 * ''Pterostichus findelii'' (Dejean, 1828) * ''Pterostichus foveolatus'' (Duftschmid, 1812) * ''Pterostichus glaferi'' O. & E.Berlov, 1996 * ''Pterostichus iberlovi'' O.Berlov, 1996 * ''Pterostichus jungens'' (Tschitscherine, 1893) * ''Pterostichus jurecekianus'' Maran, 1940 * ''Pterostichus kokeilii'' L.Miller, 1850 * ''Pterostichus kurentzovi'' Lafer, 1979 * ''Pterostichus kvanmobongus'' Lafer, 2011 * ''Pterostichus lachrymosus'' (Newman, 1838) * ''Pterostichus levadensis'' Lafer, 1979 * ''Pterostichus magoides'' (Straneo, 1937) * ''Pterostichus magus'' (Mannerheim, 1825) * ''Pterostichus maryseae'' Sun & Shi, 2018 * ''Pterostichus melanarius'' (Illiger, 1798) * ''Pterostichus melanodes'' (Chaudoir, 1878) * ''Pterostichus mellyi'' (Gebler, 1843) * ''Pterostichus montanus'' (Motschulsky, 1844) * ''Pterostichus monticoloides'' Shilenkov in Kryzhanovskij et al., 1995 * ''Pterostichus mroczkowskii'' Lafer, 2011 * ''Pterostichus nigellus'' A.Morawitz, 1862 * ''Pterostichus novus'' Straneo, 1944 * ''Pterostichus odaesanensis'' Lafer, 2011 * ''Pterostichus pawlowskii'' Lafer, 2011 * ''Pterostichus pertinax'' (Tschitscherine, 1895) * ''Pterostichus petulans'' Jedlicka, 1938 * ''Pterostichus poppiusi'' (Semenov, 1906) * ''Pterostichus probus'' Park & Kwon, 1996 * ''Pterostichus procax'' A.Morawitz, 1862 * ''Pterostichus rasilis'' Park & Kwon, 1996 * ''Pterostichus relictus'' (Newman, 1838) * ''Pterostichus robustistylis'' Sasakawa, 2005 * ''Pterostichus schoenmanni'' Kirschenhofer, 1991 * ''Pterostichus sejunctus'' Bates, 1883 * ''Pterostichus septentrionis'' (Chaudoir, 1868) * ''Pterostichus seriatus'' (Chaudoir, 1850) * ''Pterostichus seungmoi'' Park & Kwon, 1996 * ''Pterostichus shingarevi'' Lafer, 1979 * ''Pterostichus silvestris'' Sun & Shi, 2018 * ''Pterostichus songoricus'' (Motschulsky, 1845) * ''Pterostichus stygicus'' (Say, 1823) * ''Pterostichus subaeneus'' (Chaudoir, 1850) * ''Pterostichus sungariensis'' Lafer, 1979 * ''Pterostichus sutschanensis'' Jedlicka, 1962 * ''Pterostichus tatianae'' E.Berlov, 1996 * ''Pterostichus thunbergi'' A.Morawitz, 1862 * ''Pterostichus tomensis'' (Gebler, 1847) * ''Pterostichus triseriatus'' (Gebler, 1847) * ''Pterostichus tschitscherinianus'' (Jakobson, 1907) * ''Pterostichus tuberifer'' Sasakawa, 2006 * ''Pterostichus tundrae'' (Tschitscherine, 1894) * ''Pterostichus turanensis'' Jedlicka, 1959 * ''Pterostichus uralensis'' (Motschulsky, 1850) * ''Pterostichus urengaicus'' Jurecek, 1924 * ''Pterostichus vladivostokensis'' Lafer, 1979 Subgenus Phaenoraphis Tschitscherine, 1901 * ''Pterostichus acuspina'' (Tschitscherine, 1901) * ''Pterostichus peninsularis'' Park & Kwon, 1996 * ''Pterostichus sagittus'' Park, 2013 Subgenus Phonias Gozis, 1886 * ''Pterostichus apfelbecki'' Csiki, 1908 * ''Pterostichus burkhan'' Berlov & Anichtchenko, 2005 * ''Pterostichus corrusculus'' LeConte, 1873 * ''Pterostichus datshenkoae'' Sundukov, 2013 * ''Pterostichus defossus'' Bates, 1883 * ''Pterostichus diligens'' (Sturm, 1824) * ''Pterostichus dostali'' Kirschenhofer, 1981 * ''Pterostichus eobius'' (Tschitscherine, 1899) * ''Pterostichus femoralis'' (Kirby, 1837) * ''Pterostichus gyrosus'' (Motschulsky, 1866) * ''Pterostichus innshanensis'' Jedlicka, 1960 * ''Pterostichus jankowskyi'' (Tschitscherine, 1897) * ''Pterostichus kutensis'' Poppius, 1905 * ''Pterostichus liodactylus'' (Tschitscherine, 1898) * ''Pterostichus longinquus'' Bates, 1873 * ''Pterostichus longipennis'' Straneo, 1942 * ''Pterostichus monostigma'' (Tschitscherine, 1898) * ''Pterostichus morawitzianus'' (Lutshnik, 1922) * ''Pterostichus neglectus'' A.Morawitz, 1862 * ''Pterostichus ovoideus'' (Sturm, 1824) * ''Pterostichus patruelis'' (Dejean, 1831) * ''Pterostichus perisi'' Novoa, 1979 * ''Pterostichus ripensis'' (Motschulsky, 1866) * ''Pterostichus sasajii'' Morita, 2007 * ''Pterostichus setipes'' (Tschitscherine, 1898) * ''Pterostichus sotkaensis'' Jedlicka, 1958 * ''Pterostichus strenuus'' (Panzer, 1796) * ''Pterostichus stricticollis'' (Solsky, 1874) * ''Pterostichus subitus'' Csiki, 1930 * ''Pterostichus taksonyis'' Csiki, 1930 * ''Pterostichus ussuriensis'' (Tschitscherine, 1897) Subgenus Platypterus Chaudoir, 1838 * ''Pterostichus dilatatus'' A. & G.B.Villa, 1835 * ''Pterostichus lineatopunctatus'' L.Miller, 1850 * ''Pterostichus lombardus'' K.Daniel, 1900 * ''Pterostichus ottomanus'' Apfelbeck, 1908 * ''Pterostichus panzeri'' (Panzer, 1802) * ''Pterostichus truncatus'' (Dejean, 1828) * ''Pterostichus ziegleri'' (Duftschmid, 1812) Subgenus Platysma Bonelli, 1810 * ''Pterostichus chotanensis'' (Tschitscherine, 1893) * ''Pterostichus cordaticollis'' Heyden, 1884 * ''Pterostichus crenulatopunctatus'' (R.F.Sahlberg, 1844) * ''Pterostichus eschscholtzii'' (Germar, 1823) * ''Pterostichus galinae'' Kabak, 1992 * ''Pterostichus insignicollis'' (Tschitscherine, 1893) * ''Pterostichus leptis'' Bates, 1883 * ''Pterostichus niger'' (Schaller, 1783) * ''Pterostichus planicola'' (Tschitscherine, 1899) * ''Pterostichus sachtlebeni'' Jedlicka, 1962 * ''Pterostichus turcomannicus'' Motschulsky, 1850 Subgenus Plectes Fischer von Waldheim, 1822 * ''Pterostichus consors'' (Tschitscherine, 1893) * ''Pterostichus drescheri'' (Fischer von Waldheim, 1817) Subgenus Pledarus Motschulsky, 1866 * ''Pterostichus gibbicollis'' (Motschulsky, 1844) * ''Pterostichus larisae'' Sundukov, 2013 * ''Pterostichus lutschniki'' Jedlicka, 1962 * ''Pterostichus nigellatus'' Kirschenhofer, 1991 Subgenus Pseudethira Sciaky, 1996 * ''Pterostichus angoarnigi'' Morvan, 1994 * ''Pterostichus atrox'' (Andrewes, 1937) * ''Pterostichus balachowskyi'' Morvan, 1972 * ''Pterostichus balu'' J.Schmidt, 2009 * ''Pterostichus bhutanensis'' Morvan, 1978 * ''Pterostichus brancuccii'' Straneo, 1982 * ''Pterostichus brevilama'' Straneo, 1989 * ''Pterostichus bureli'' Morvan, 1994 * ''Pterostichus chainapaani'' J.Schmidt, 2006 * ''Pterostichus championi'' Andrewes, 1926 * ''Pterostichus conaensis'' J.Schmidt & Tian, 2011 * ''Pterostichus deuvei'' Lassalle, 1985 * ''Pterostichus dhorpatanicus'' Straneo, 1977 * ''Pterostichus dolens'' (Tschitscherine, 1900) * ''Pterostichus dorjulensis'' Morvan, 1978 * ''Pterostichus exochus'' Andrewes, 1930 * ''Pterostichus fritzhiekei'' J.Schmidt, 1994 * ''Pterostichus gagates'' (Hope, 1831) * ''Pterostichus ganesh'' J.Schmidt, 2006 * ''Pterostichus ganja'' J.Schmidt, 1995 * ''Pterostichus gerdi'' J.Schmidt, 2009 * ''Pterostichus gobettii'' Straneo, 1983 * ''Pterostichus gompanus'' Straneo, 1983 * ''Pterostichus harmandi'' (Tschitscherine, 1900) * ''Pterostichus hartmanni'' J.Schmidt, 1995 * ''Pterostichus hedkeinekus'' Morvan, 1995 * ''Pterostichus hoelli'' J.Schmidt, 2009 * ''Pterostichus immarginatus'' Straneo, 1977 * ''Pterostichus jaljaleensis'' J.Schmidt, 2009 * ''Pterostichus janbritoi'' Morvan, 1994 * ''Pterostichus juga'' J.Schmidt, 2006 * ''Pterostichus kadoudali'' Morvan, 1982 * ''Pterostichus kalo'' J.Schmidt, 2012 * ''Pterostichus kangchenjunga'' J.Schmidt, 2012 * ''Pterostichus keltiekus'' Morvan, 1981 * ''Pterostichus kleinfeldi'' Straneo, 1982 * ''Pterostichus kopetzi'' J.Schmidt, 2006 * ''Pterostichus leica'' J.Schmidt, 2012 * ''Pterostichus letensis'' Habu, 1973 * ''Pterostichus lugi'' Morvan, 1982 * ''Pterostichus makalu'' J.Schmidt, 2012 * ''Pterostichus matsumurai'' Habu, 1973 * ''Pterostichus mewakholensis'' J.Schmidt, 2012 * ''Pterostichus miehei'' J.Schmidt, 2006 * ''Pterostichus nepalensis'' Straneo, 1977 * ''Pterostichus nowitzkii'' (Tschitscherine, 1899) * ''Pterostichus olafi'' J.Schmidt, 2006 * ''Pterostichus perlamatus'' Straneo, 1989 * ''Pterostichus pseudoharmandi'' Morvan, 1981 * ''Pterostichus pseudoplatysma'' Straneo, 1982 * ''Pterostichus raraensis'' Morvan, 1980 * ''Pterostichus rugosiceps'' J.Schmidt, 2012 * ''Pterostichus sagarmatha'' J.Schmidt, 2012 * ''Pterostichus santostamangi'' J.Schmidt, 2006 * ''Pterostichus schrettenbrunneri'' J.Schmidt, 2012 * ''Pterostichus thanglaensis'' J.Schmidt, 2012 * ''Pterostichus tulobalu'' J.Schmidt, 2009 * ''Pterostichus weigeli'' J.Schmidt, 2006 * ''Pterostichus weiperti'' J.Schmidt, 2006 * ''Pterostichus wittmeri'' Morvan, 1978 Subgenus Pseudoferonina Ball, 1965 * ''Pterostichus amadeus'' Bousquet, 2012 * ''Pterostichus bousqueti'' Bergdahl in Bergdahl & Kavanaugh, 2011 * ''Pterostichus campbelli'' Bousquet, 1985 * ''Pterostichus humidulus'' (Van Dyke, 1943) * ''Pterostichus lanei'' Van Dyke, 1926 * ''Pterostichus lolo'' Bergdahl in Bergdahl & Kavanaugh, 2011 * ''Pterostichus shulli'' (Hatch, 1949) * ''Pterostichus smetanai'' Bousquet, 1985 * ''Pterostichus spathifer'' Bousquet, 1992 Subgenus Pseudohaptoderus Tschitscherine, 1888 * ''Pterostichus aemiliae'' Facchini & Sciaky, 2000 * ''Pterostichus jugivagus'' (Tschitscherine, 1898) * ''Pterostichus ssemenovi'' Tschitscherine, 1888 Subgenus Pseudomaseus Chaudoir, 1838 * ''Pterostichus aimaki'' Jedlicka, 1968 * ''Pterostichus ambigenus'' Bates, 1883 * ''Pterostichus anthracinus'' (Illiger, 1798) * ''Pterostichus basipunctatus'' Straneo, 1955 * ''Pterostichus carri'' Angus; Galian; Wrase & Chaladze, 2009 * ''Pterostichus chujoiellus'' Jedlicka, 1962 * ''Pterostichus fuscicornis'' (Reiche & Saulcy, 1855) * ''Pterostichus gracilis'' (Dejean, 1828) * ''Pterostichus hejkali'' B.Gueorguiev & Skoupy, 2011 * ''Pterostichus koslovi'' Solodovnikov, 2001 * ''Pterostichus luctuosus'' (Dejean, 1828) * ''Pterostichus michailovi'' Wrase, 1992 * ''Pterostichus minor'' (Gyllenhal, 1827) * ''Pterostichus mukdenensis'' Breit, 1933 * ''Pterostichus nigrita'' (Paykull, 1790) * ''Pterostichus oenotrius'' Ravizza, 1975 * ''Pterostichus piceolus'' (Chaudoir, 1850) * ''Pterostichus rhaeticus'' Heer, 1837 * ''Pterostichus rotundangulus'' A.Morawitz, 1862 * ''Pterostichus tenuis'' (Casey, 1924) * ''Pterostichus tsukubasanus'' Kasahara, 1988 Subgenus Pseudorambousekiella Schweiger, 1967 * ''Pterostichus montisdeorum'' Schweiger, 1967 Subgenus Pseudorites Ganglbauer, 1891 * ''Pterostichus nicaeensis'' (A. & G.B.Villa, 1835) Subgenus Pseudosteropus Chaudoir, 1838 * ''Pterostichus illigeri'' (Panzer, 1802) * ''Pterostichus schmidtii'' (Chaudoir, 1838) Subgenus Pterostichus Bonelli, 1810 * ''Pterostichus andreinii'' Dodero, 1922 * ''Pterostichus auratus'' Heer, 1837 * ''Pterostichus bischoffianus'' Jedlicka, 1936 * ''Pterostichus bruckii'' Schaum, 1859 * ''Pterostichus cantalicus'' (Chaudoir, 1868) * ''Pterostichus cristatus'' (L.Dufour, 1820) * ''Pterostichus devillei'' Puel, 1924 * ''Pterostichus dux'' L.Schaufuss, 1862 * ''Pterostichus epiroticus'' Csiki, 1930 * ''Pterostichus fasciatopunctatus'' (Creutzer, 1799) * ''Pterostichus focarilei'' Casale & Giachino, 1985 * ''Pterostichus funestes'' Csiki, 1930 * ''Pterostichus hagenbachii'' Sturm, 1824 * ''Pterostichus honnoratii'' (Dejean, 1828) * ''Pterostichus impressicollis'' (Fairmaire & Laboulbène, 1854) * ''Pterostichus lanista'' (Tschitscherine, 1898) * ''Pterostichus lasserrei'' (Dejean, 1828) * ''Pterostichus latifianus'' Apfelbeck, 1906 * ''Pterostichus lumensis'' Apfelbeck, 1905 * ''Pterostichus malissorum'' Apfelbeck, 1905 * ''Pterostichus meisteri'' Reitter, 1885 * ''Pterostichus merklii'' (J.Frivaldszky, 1879) * ''Pterostichus micans'' Heer, 1838 * ''Pterostichus pedemontanus'' Ganglbauer, 1891 * ''Pterostichus pentheri'' Apfelbeck, 1918 * ''Pterostichus phaeopus'' (Sainte-Claire Deville, 1903) * ''Pterostichus reiseri'' Ganglbauer, 1889 * ''Pterostichus rhilensis'' Rottenberg, 1874 * ''Pterostichus ruffoi'' Sciaky, 1984 * ''Pterostichus rufipes'' (Dejean, 1828) * ''Pterostichus rutilans'' (Dejean, 1828) * ''Pterostichus sacheri'' I.Frivaldszky von Frivald, 1865 * ''Pterostichus vagepunctatus'' Heer, 1837 * ''Pterostichus walteri'' Reitter, 1883 Subgenus Rambousekiella Knirsch, 1925 * ''Pterostichus ledenikensis'' (Knirsch, 1925) Subgenus Rhagadus Motschulsky, 1866 * ''Pterostichus brittoni'' Habu, 1958 * ''Pterostichus glabripennis'' Jedlicka, 1962 * ''Pterostichus harponifer'' Tanaka, 1987 * ''Pterostichus ishiii'' Morita; Kurosa & Mori, 2009 * ''Pterostichus kalhys'' (Motschulsky, 1866) * ''Pterostichus kimurai'' Morita, 1994 * ''Pterostichus komiyai'' Morita, 2007 * ''Pterostichus laevipunctatus'' Tschitscherine, 1889 * ''Pterostichus latemarginatus'' (Straneo, 1936) * ''Pterostichus microcephalus'' (Motschulsky, 1861) * ''Pterostichus modicellus'' (Tschitscherine, 1897) * ''Pterostichus mundatus'' Jedlicka, 1940 * ''Pterostichus nimbatidius'' (Chaudoir, 1878) * ''Pterostichus polygenus'' Bates, 1883 * ''Pterostichus satsumanus'' Habu, 1958 * ''Pterostichus solskyi'' (Chaudoir, 1878) * ''Pterostichus takaosanus'' Habu, 1958 * ''Pterostichus thorectes'' Bates, 1873 * ''Pterostichus thorectoides'' Jedlicka, 1958 Subgenus Sinoreophilus Sciaky, 1996 * ''Pterostichus potanini'' Tschitscherine, 1889 * ''Pterostichus scalptus'' Sciaky & Wrase, 1997 * ''Pterostichus strigosus'' Sciaky & Wrase, 1997 * ''Pterostichus validior'' Tschitscherine, 1889 Subgenus Sinosteropus Sciaky, 1994 * ''Pterostichus barbarae'' Sciaky & Facchini, 2003 * ''Pterostichus carinatus'' Sciaky & Facchini, 2003 * ''Pterostichus cathaicus'' Sciaky, 1994 * ''Pterostichus curvatus'' Sciaky & Facchini, 2003 * ''Pterostichus horstweiperti'' J.Schmidt & Guorguiev, 2013 * ''Pterostichus jaechianus'' Sciaky & Facchini, 2003 * ''Pterostichus jani'' Sciaky & Facchini, 2003 * ''Pterostichus latitemporis'' Sciaky & Wrase, 1997 * ''Pterostichus microbus'' Sciaky & Facchini, 2003 * ''Pterostichus perhoplites'' J.Schmidt & Tian, 2011 * ''Pterostichus pseudobarbarae'' Sciaky & Facchini, 2003 * ''Pterostichus pseudojugivagus'' J.Schmidt & Tian, 2011 * ''Pterostichus pseudorotundus'' Sciaky & Facchini, 2003 * ''Pterostichus pseudosinensis'' Sciaky & Facchini, 2003 * ''Pterostichus rotundus'' Sciaky, 1994 * ''Pterostichus schneideri'' Sciaky & Facchini, 2003 * ''Pterostichus scuticollis'' (Fairmaire, 1889) * ''Pterostichus septemtrionalis'' Sciaky & Facchini, 2003 * ''Pterostichus sinensis'' Jedlicka, 1962 * ''Pterostichus triangularis'' Sciaky & Facchini, 2003 * ''Pterostichus wrasei'' Sciaky & Facchini, 2003 Subgenus Sphodroferonia Kasahara & Y.Ito, 1987 * ''Pterostichus abaciformis'' Straneo, 1955 * ''Pterostichus arcuaticarinatus'' Kasahara, 1986 * ''Pterostichus audax'' (Tschitscherine, 1895) * ''Pterostichus bilobatus'' N.Ito, 2010 * ''Pterostichus bisulcitarsis'' N.Ito, 2012 * ''Pterostichus confusianus'' N.Ito, 2010 * ''Pterostichus constricticollis'' N.Ito, 2010 * ''Pterostichus esakii'' Ishida, 1959 * ''Pterostichus fenestratus'' Kasahara & Ohtani, 1989 * ''Pterostichus geojensis'' Sasakawa; Kim & Kubota, 2006 * ''Pterostichus gotoensis'' Kasahara & Matsumoto, 1990 * ''Pterostichus gracilitarsis'' N.Ito, 2010 * ''Pterostichus hikosanus'' Kasahara, 1994 * ''Pterostichus hisamatsui'' Ishida & Shibata, 1961 * ''Pterostichus imasakai'' Kasahara & Ohtani, 1988 * ''Pterostichus imitatorius'' N.Ito, 2010 * ''Pterostichus ishizuchiensis'' Kasahara, 1985 * ''Pterostichus itoi'' Kasahara, 1986 * ''Pterostichus kosakai'' Morita, 1998 * ''Pterostichus koujitanakai'' N.Ito, 2010 * ''Pterostichus kyushuensis'' Habu, 1955 * ''Pterostichus laeviceps'' N.Ito, 2010 * ''Pterostichus macrocephalus'' Habu, 1955 * ''Pterostichus marginellus'' Sasakawa; Kim & Kim, 2006 * ''Pterostichus masatakayoshidai'' N.Ito, 2010 * ''Pterostichus masidai'' Ishida, 1959 * ''Pterostichus masumotoi'' Tanaka; Morita & Suga, 1987 * ''Pterostichus miyamai'' Kasahara & Y.Ito, 1987 * ''Pterostichus mosaicus'' Sasakawa, 2005 * ''Pterostichus mucronatus'' Straneo, 1955 * ''Pterostichus multinodosus'' Sasakawa; Kim & Kim, 2006 * ''Pterostichus namedai'' Morita, 2019 * ''Pterostichus nasui'' Kasahara, 1993 * ''Pterostichus nishidai'' Nakane, 1989 * ''Pterostichus obscurimpressus'' N.Ito, 2010 * ''Pterostichus okiensis'' Nakane, 1989 * ''Pterostichus orionis'' Jedlicka, 1962 * ''Pterostichus pachinus'' Bates, 1883 * ''Pterostichus platyocularis'' N.Ito, 2010 * ''Pterostichus platypennis'' N.Ito, 2010 * ''Pterostichus plesiomorphus'' Nemoto, 1989 * ''Pterostichus prominens'' N.Ito, 2010 * ''Pterostichus pseudopachinus'' Nakane, 1963 * ''Pterostichus punctatulus'' N.Ito, 2010 * ''Pterostichus punctibasalis'' N.Ito, 2012 * ''Pterostichus raptor'' (Tschitscherine, 1901) * ''Pterostichus sakuragii'' N.Ito, 2010 * ''Pterostichus sculpturalis'' N.Ito, 2012 * ''Pterostichus scurra'' (Tschitscherine, 1901) * ''Pterostichus scurroides'' Jedlicka, 1958 * ''Pterostichus sediorcrus'' Sasakawa & Kim, 2006 * ''Pterostichus seunglaki'' Park & Kwon, 1996 * ''Pterostichus shiibanus'' Habu, 1958 * ''Pterostichus sphodriformis'' Bates, 1873 * ''Pterostichus strigipennis'' N.Ito, 2012 * ''Pterostichus subabacipennis'' N.Ito, 2012 * ''Pterostichus sulcatus'' N.Ito, 2010 * ''Pterostichus suruganus'' Straneo in Nakane, 1979 * ''Pterostichus symmetricus'' Straneo, 1955 * ''Pterostichus taradakensis'' Kasahara & Ohtani, 1988 * ''Pterostichus tosanus'' Kasahara & Y.Ito, 1997 * ''Pterostichus touzalinioides'' Sasakawa; Kim & Kim, 2007 * ''Pterostichus tsurugaensis'' Ishida, 1959 * ''Pterostichus variicornis'' Sasakawa; Kim & Kim, 2007 * ''Pterostichus yakushimanus'' Nakane & Ishida, 1961 * ''Pterostichus yamajii'' Kasahara, 1993 * ''Pterostichus yoshidai'' Kasahara, 1985 * ''Pterostichus yoshikawai'' Ishida, 1959 * ''Pterostichus yoshiyukiitoi'' N.Ito, 2010 Subgenus Steropanus Fairmaire, 1889 * ''Pterostichus aequus'' (Andrewes, 1937) * ''Pterostichus alveolatus'' Fedorenko, 2018 * ''Pterostichus asulcatus'' Fedorenko, 2018 * ''Pterostichus boriskataevi'' Fedorenko, 2020 * ''Pterostichus cavifrons'' Fedorenko, 2018 * ''Pterostichus deliciatus'' (Andrewes, 1937) * ''Pterostichus felix'' (Andrewes, 1937) * ''Pterostichus forticornis'' (Fairmaire, 1889) * ''Pterostichus fossifrons'' Fedorenko, 2020 * ''Pterostichus glymmiger'' (Andrewes, 1937) * ''Pterostichus infissus'' (Andrewes, 1937) * ''Pterostichus mengtzei'' Jedlicka, 1931 * ''Pterostichus obliteratus'' Fedorenko, 2018 * ''Pterostichus pseudoglymmiger'' Fedorenko, 2020 * ''Pterostichus pseudoviolaceus'' Fedorenko, 2018 * ''Pterostichus securipenis'' Fedorenko, 2020 * ''Pterostichus sulcatipennis'' Fedorenko, 2018 * ''Pterostichus thailandensis'' (Morvan, 1992) * ''Pterostichus thailandicus'' Straneo, 1989 * ''Pterostichus violaceus'' (Straneo, 1949) Subgenus Steropus Dejean, 1821 * ''Pterostichus catalonicus'' J.Daniel in K. & J.Daniel, 1906 * ''Pterostichus ebenus'' (Quensel, 1806) * ''Pterostichus ferreri'' (Español & Mateu, 1942) * ''Pterostichus galaecianus'' Lauffer, 1909 * ''Pterostichus gallega'' (Fairmaire, 1859) * ''Pterostichus ghilianii'' (Putzeys, 1845) * ''Pterostichus insidatrix'' (Piochard de la Brûlerie, 1872) * ''Pterostichus madidus'' (Fabricius, 1775) * ''Pterostichus rifensis'' Antoine, 1933 * ''Pterostichus validus'' (Dejean, 1828) Subgenus Tausternus Fedorenko, 2020 * ''Pterostichus hoii'' Fedorenko, 2020 Subgenus Tinautius Mateu, 1997 * ''Pterostichus exilis'' (Mateu, 2001) * ''Pterostichus troglophilus'' (Mateu, 1997) Subgenus Tschitscherinea Berg, 1898 * ''Pterostichus farkaci'' Sciaky, 1997 * ''Pterostichus filum'' (Tschitscherine, 1897) * ''Pterostichus krali'' Sciaky, 1997 * ''Pterostichus mulensis'' Sciaky, 1997 Subgenus Tubuliphallus Sciaky & Allegro, 2013 * ''Pterostichus megacephalus'' Sciaky & Allegro, 2013 Subgenus Unitrichus Sciaky, 1997 * ''Pterostichus platyops'' Sciaky, 1997 Subgenus Vietosteropus Fedorenko, 2017 * ''Pterostichus abramovi'' Fedorenko, 2017 * ''Pterostichus anichkini'' Fedorenko, 2017 * ''Pterostichus annamita'' (Straneo, 1939) * ''Pterostichus bidoupensis'' Fedorenko, 2017 * ''Pterostichus cavicollis'' Straneo, 1984 * ''Pterostichus chupanphan'' Fedorenko, 2017 * ''Pterostichus dalatensis'' Fedorenko, 2017 * ''Pterostichus gialaiensis'' Fedorenko, 2017 * ''Pterostichus honbaensis'' Fedorenko, 2017 * ''Pterostichus konchurang'' Fedorenko, 2017 * ''Pterostichus konplongensis'' Fedorenko, 2017 * ''Pterostichus kontumensis'' Fedorenko, 2017 * ''Pterostichus kuznetsovi'' Fedorenko, 2020 * ''Pterostichus ngoclinhensis'' Fedorenko, 2017 * ''Pterostichus ngokboci'' Fedorenko, 2017 * ''Pterostichus semiopacus'' Fedorenko, 2017 * ''Pterostichus sulcicollis'' Fedorenko, 2017 Subgenus Wraseiellus Shi & Sciaky, 2013 * ''Pterostichus andrewesi'' Jedlicka, 1931 * ''Pterostichus comatus'' Shi & Sciaky, 2013 * ''Pterostichus crassiapex'' Shi & Sciaky, 2013 * ''Pterostichus diversus'' (Fairmaire, 1886) * ''Pterostichus kambaiti'' (Andrewes, 1947) * ''Pterostichus meyeri'' Jedlicka, 1934 * ''Pterostichus pseudodiversus'' Shi & Sciaky, 2013 * ''Pterostichus stictopleurus'' (Fairmaire, 1889) Unassigned Subgenus * ''Pterostichus analis'' Jedlicka, 1963 * ''Pterostichus antepunctatus'' Straneo, 1949 * ''Pterostichus aristochroides'' Deuve, 2006 * ''Pterostichus chenpengi'' Li, 1992 * ''Pterostichus clepsydra'' Sciaky & Wrase, 1997 * ''Pterostichus deuvesianus'' Morvan, 1995 * ''Pterostichus georgi'' Straneo, 1963 * ''Pterostichus glabricollis'' Jedlicka, 1962 * ''Pterostichus gwervaenus'' Morvan, 1995 * ''Pterostichus heilongjiangensis'' Li, 1992 * ''Pterostichus imitatus'' Morvan, 1978 * ''Pterostichus jelepus'' Andrewes, 1932 * ''Pterostichus kiangsu'' Jedlicka, 1965 * ''Pterostichus kolosovi'' (Lutshnik, 1928) * ''Pterostichus leviculus'' Andrewes, 1933 * ''Pterostichus lidarus'' (Andrewes, 1937) * ''Pterostichus lukjanovitshi'' (Lutshnik, 1927) * ''Pterostichus malaisei'' Jedlicka, 1965 * ''Pterostichus migliaccioi'' Straneo, 1982 * ''Pterostichus orobius'' Fairmaire, 1861 * ''Pterostichus peilingi'' Jedlicka, 1937 * ''Pterostichus punctisternus'' Straneo, 1989 * ''Pterostichus quadriimpressus'' Straneo, 1983 * ''Pterostichus reuteri'' Wrase & J.Schmidt, 2006 * ''Pterostichus rolex'' Morvan, 1995 * ''Pterostichus sintanus'' Andrewes, 1927 * ''Pterostichus styx'' (Andrewes, 1937) * ''Pterostichus szekessyianus'' Sciaky, 1996 * ''Pterostichus tantillus'' (Fairmaire, 1889) * ''Pterostichus tonkinensis'' Straneo, 1980 * ''Pterostichus zhejiangensis'' Kirschenhofer, 1997 * †''Pterostichus abrogatus'' Scudder, 1890 * †''Pterostichus depletus'' Scudder, 1900 * †''Pterostichus destitutus'' Scudder, 1890 * †''Pterostichus destructus'' Scudder, 1890 * †''Pterostichus dormitans'' Scudder, 1890 * †''Pterostichus fernquisti'' Wickham, 1931 * †''Pterostichus fractus'' Scudder, 1890 * †''Pterostichus laevigatus'' Scudder, 1890 * †''Pterostichus minax'' (Oustalet, 1874) * †''Pterostichus minutulus'' Heer, 1862 * †''Pterostichus provincialis'' (Oustalet, 1874) * †''Pterostichus pumpellyi'' Scudder, 1900 * †''Pterostichus walcotti'' Scudder, 1900

References

{{Taxonbar, from=Q1540893 Pterostichus, Pterostichinae Taxa named by Franco Andrea Bonelli