Ptenopus Garrulus
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''Ptenopus garrulus'', also known commonly as the common barking gecko and the whistling gecko, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Gekkonidae Gekkonidae (the common geckos) is the largest family of geckos, containing over 950 described species in 62 genera. The Gekkonidae contain many of the most widespread gecko species, including house geckos (''Hemidactylus''), the tokay gecko (''Ge ...
. The species is native to southern Africa. There are two recognized
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
.


Description

The
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
coloration of ''P. garrulus'' varies from reddish brown to grayish yellow to match the local substrate. Ventrally, it is white. The toes are strongly fringed. It has 110–190 rows of scales around the body at midbody. The subspecies ''P.g. garrulus'' has more than 160 rows, but the subspecies ''P. g. maculatus'' has fewer than 160 rows. Adults usually have a snout-to-vent length (SVL) of , which is relatively small for the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
.


Geographic range

''P. garrulus'' is found in
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
,
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
, and
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
.


Habitat

The preferred natural
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s of ''P. garrulus'' are
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
,
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
, and
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
, at altitudes from sea level to .


Behavior

''P. garrulus'' is
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
and
fossorial A fossorial animal () is one that is adapted to digging and which lives primarily (but not solely) underground. Examples of fossorial vertebrates are Mole (animal), moles, badgers, naked mole-rats, meerkats, armadillos, wombats, and mole salamand ...
. Males vocalize from the entrances of their burrows at dusk and on moonlit nights.


Diet

''P. garrulus'' preys upon ants, beetles, and termites. Branch, Bill (2004). ''Field Guide to Snakes and other Reptiles of Southern Africa''. Third Revised Edition, Second Impression. Sanibel Island, Florida: Ralph Curtis Books. 399 pp. . (''Ptenopus garrulus'', pp. 265–266).


Reproduction

''P. garrulus'' is
oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...
.


Subspecies

Two subspecies are recognized as being valid, including the
nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
. *''Ptenopus garrulus garrulus'' *''Ptenopus garrulus maculatus'' ''
Nota bene ( ; plural: ) is the Latin language, Latin phrase meaning ''note well''. In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
other than ''Ptenopus''.


Gallery

Common Barking Gecko (Ptenopus garrulus) barking (6854290978).jpg, Barking from lair Ptenopus garrulus garrulus.jpg,
Nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
,
''P. g. garrulus'' Ptenopus garrulus maculatus01.jpg, ''P. g. maculatus''


References


Further reading

* Boulenger GA (1885). ''Catalogue of the Lizards in the British Museum (Natural History). Second Edition. Volume I. Geckonidæ'' .... London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xii + 436 pp. + Plates I–XXXII. (''Ptenopus garrulus'', new combination, p. 15 + Plate II, figures 2, 2a–2d). * Gray JE (1866). "Descriptions of Two New Genera of Lizards from Damaraland". ''Proceedings of the Zoological Society of London'' 1865: 640–642 + Plate XXXVIII. (''Ptenopus maculatus'', new species, p. 640 + Plate XXXVIII, figure 1). * Smith A (1849). ''Illustrations of the Zoology of South Africa; Consisting Chiefly of Figures and Descriptions of the Objects of Natural History Collected during an Expedition into the Interior of South Africa, in the Years 1834, 1835, and 1836; Fitted out by "The Cape of Good Hope Association for Exploring Central Africa:" Together with a Summary of African Zoology, and an Inquiry into the Geographical Ranges of Species in that Quarter of the Globe. Reptilia.'' olume III London: Lords Commissioners of her Majesty's Treasury. (Smith, Elder and Co., printers). 78 Plates + unnumbered pages of text + Appendix pp. 1–28. (''Stenodactylus garrulus'', new species, Appendix p. 6). (in English and Latin). Ptenopus Reptiles described in 1849 {{Gekkonidae-stub