Ptarmigans
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''Lagopus'' is a genus of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s in the
grouse Grouse are a group of birds from the order (biology), order Galliformes, in the family (biology), family Phasianidae. Grouse are presently assigned to the Tribe (biology), tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetr ...
subfamily commonly known as ptarmigans (). The genus contains four living species with numerous described subspecies, all living in
tundra In physical geography, a tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. There are three regions and associated types of tundra: #Arctic, Arctic, Alpine tundra, Alpine, and #Antarctic ...
or cold upland areas.


Taxonomy and etymology

The
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Lagopus'' was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
in 1760 with the
willow ptarmigan The willow ptarmigan ( ); ''Lagopus lagopus'') or willow grouse is a bird in the grouse subfamily Tetraoninae of the pheasant family Phasianidae. It is also known colloquially as awebo bird. The willow ptarmigan breeds in birch and other forests ...
(''Lagopus lagopus'') as the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
. The genus name ''Lagopus'' is derived from
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
(), meaning "
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...
,
rabbit Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
", + (), "foot", in reference to the feathered feet and toes typical of this cold-adapted group (such as the
snowshoe hare The snowshoe hare (''Lepus americanus''), also called the varying hare or snowshoe rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet. The animal's feet prevent it from sink ...
). The specific epithets ''muta'' and ''leucura'' were for a long time misspelt ''mutus'' and ''leucurus'', in the erroneous belief that the ending of ''Lagopus'' denotes
masculine gender In linguistics, a grammatical gender system is a specific form of a noun class system, where nouns are assigned to gender categories that are often not related to the real-world qualities of the entities denoted by those nouns. In languages wi ...
. However, as the Ancient Greek term is of feminine gender, and the specific epithet has to agree with that, the feminine ''muta'' and ''leucura'' are correct. The English name ''ptarmigan'' comes from the
Scottish Gaelic Scottish Gaelic (, ; Endonym and exonym, endonym: ), also known as Scots Gaelic or simply Gaelic, is a Celtic language native to the Gaels of Scotland. As a member of the Goidelic language, Goidelic branch of Celtic, Scottish Gaelic, alongs ...
name for ''L. muta'', ([]), meaning “croaker”, which refers to the bird’s frog-like call. The p- was added due to a mistaken belief in a Greek origin, as if the word were related to the Greek word (''pterón''), 'wing'.Lockwood, W. B. (1984). ''The Oxford Book of British Bird Names''. Oxford University Press .


Description

The four species are all sedentary specialists of cold regions. Willow ptarmigan is a circumpolar
boreal forest Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by pinophyta, coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. I ...
species, white-tailed ptarmigan is a North American alpine bird, and rock ptarmigan breeds in both
Arctic The Arctic (; . ) is the polar regions of Earth, polar region of Earth that surrounds the North Pole, lying within the Arctic Circle. The Arctic region, from the IERS Reference Meridian travelling east, consists of parts of northern Norway ( ...
and mountain habitats across
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
and North America. With the exception of the red grouse (until recently considered a subspecies of willow ptarmigan), all have a white winter plumage that helps them blend into the snowy background. Even their
remiges Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the ta ...
are white, while these feathers are black in almost all birds (even birds that are predominantly white, such as the
Bali myna The Bali myna (''Leucopsar rothschildi''), also known as Rothschild's mynah, Bali starling, or Bali mynah, locally known as jalak Bali, is a medium-sized (up to long), stocky myna, almost wholly white with a long, drooping crest, and black tip ...
) because
melanin Melanin (; ) is a family of biomolecules organized as oligomers or polymers, which among other functions provide the pigments of many organisms. Melanin pigments are produced in a specialized group of cells known as melanocytes. There are ...
makes them more resilient and thus improves flight performance. The ''Lagopus'' grouse apparently found it easier to escape
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s by not being seen than by flying away. These are hardy vegetarian birds, but
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s are also taken by the developing young. In all species except for the willow ptarmigan, the female takes all responsibility for nesting and caring for the chicks, as is typical with gamebirds.


Species

The genus contains four species:


Fossil record

Two
prehistoric Prehistory, also called pre-literary history, is the period of human history between the first known use of stone tools by hominins  million years ago and the beginning of recorded history with the invention of writing systems. The use o ...
species and two
paleosubspecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
are only known from
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s: * ''Lagopus atavus'' (
Early Pliocene Early may refer to: Places in the United States * Early, Iowa, a city * Early, Texas, a city * Early Branch, a stream in Missouri * Early County, Georgia * Fort Early, Georgia, an early 19th century fort Music * Early B, stage name of Jamaican d ...
of
Bulgaria Bulgaria, officially the Republic of Bulgaria, is a country in Southeast Europe. It is situated on the eastern portion of the Balkans directly south of the Danube river and west of the Black Sea. Bulgaria is bordered by Greece and Turkey t ...
? -
Late Pliocene Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * Late (The 77s album), ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudo ...
) * ''Lagopus balcanicus'' (Late Pliocene of
Varshets Varshets (, variously transliterated; ) is a spa town in Montana Province, northwestern Bulgaria. It is the administrative centre of the homonymous Varshets Municipality. As of December 2019, its population was 5,444. The town is located on th ...
, Bulgaria)Boev, Z. 1995. "Middle Villafranchian birds from Varshets (Western Balkan Range - Bulgaria)". In: Peters, D. (ed.). ''Acta palaeornithologica''. 3. Symposium SAPE. 5. Internat. Senckenberg-Konferenz 22–26 Juni 1992. - Courier Forschungsinstitut Senckenberg. Frankfurt a. M., 181: 259-269. * ''Lagopus lagopus noaillensis'' (
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
of W Europe) * ''Lagopus mutus correzensis'' (Pleistocene of W Europe)


References

* Madge, Steve; McGowan, Philip J. K. & Kirwan, Guy M. (2002): ''Pheasants, partidges and grouse: a guide to the pheasants, partridges, quails, grouse, guineafowl, buttonquails and sandgrouse of the world''.
Christopher Helm Christopher Alexander Roger Helm (born Dundee, 1 February 1937 – 20 January 2007) was a Scottish book publisher, notably of ornithology related titles, including the ''Helm Identification Guides''. Born in Dundee, he was raised in Forfar, wh ...
, London. .


External links

* {{Taxonbar, from=Q162029 Bird genera Provincial symbols of Newfoundland and Labrador