''Pseudolarix amabilis'' is a

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

conifer Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...

ous tree in the pine family

Pinaceae The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly know ...

. The species is commonly known as golden larch, but being more closely related to '' Keteleeria'', ''

Abies Firs (''Abies'') are a genus of 48–56 species of evergreen coniferous trees in the family Pinaceae. They are found on mountains throughout much of North and Central America, Europe, Asia, and North Africa. The genus is most closely related to ...

'' and ''

Cedrus ''Cedrus'', common English name cedar, is a genus of coniferous trees in the plant family Pinaceae (subfamily Abietoideae). They are native to the mountains of the western Himalayas and the Mediterranean region, occurring at altitudes of 1,500� ...

'', is not a true

larch Larches are deciduous conifers in the genus ''Larix'', of the family Pinaceae (subfamily Laricoideae). Growing from tall, they are native to much of the cooler temperate northern hemisphere, on lowlands in the north and high on mountains furt ...

(''Larix''). ''P. amabilis'' is

native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and enterta ...

to eastern China, occurring in small areas in the mountains of southern

Anhui Anhui , (; formerly romanized as Anhwei) is a landlocked province of the People's Republic of China, part of the East China region. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze River ...

Zhejiang Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by ...

Fujian Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its c ...

Jiangxi Jiangxi (; ; formerly romanized as Kiangsi or Chianghsi) is a landlocked province in the east of the People's Republic of China. Its major cities include Nanchang and Jiujiang. Spanning from the banks of the Yangtze river in the north into h ...

Hunan Hunan (, ; ) is a landlocked province of the People's Republic of China, part of the South Central China region. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangxi ...

Hubei Hubei (; ; alternately Hupeh) is a landlocked province of the People's Republic of China, and is part of the Central China region. The name of the province means "north of the lake", referring to its position north of Dongting Lake. The pr ...

and eastern

Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of th ...

, at altitudes of . The earliest known occurrences are of

compression fossil A compression fossil is a fossil preserved in sedimentary rock that has undergone physical compression. While it is uncommon to find animals preserved as good compression fossils, it is very common to find plants preserved this way. The reason f ...

s found in the

Ypresian In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian i ...

Allenby Formation The Allenby formation is a sedimentary rock formation in British Columbia which was deposited during the Ypresian stage of the Early Eocene. It consists of conglomerates, sandstones with interbedded shales and coal. The shales contain an abund ...

and

mummified A mummy is a dead human or an animal whose soft tissues and organs have been preserved by either intentional or accidental exposure to chemicals, extreme cold, very low humidity, or lack of air, so that the recovered body does not decay fu ...

fossils found in the

Late Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "d ...

Buchanan Lake Formation on

Axel Heiberg Island Axel Heiberg Island ( iu, ᐅᒥᖕᒪᑦ ᓄᓈᑦ, ) is an uninhabited island in the Qikiqtaaluk Region, Nunavut, Canada. Located in the Arctic Ocean, it is the 32nd largest island in the world and Canada's seventh largest island. Accordin ...

Growth

It is a

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...

coniferous tree reaching tall, with a broad conic crown. The shoots are

dimorphic Dimorphism or dimorphic may refer to: Science * Dimorphic root systems, plant roots with two distinctive forms for two separate functions * Sexual dimorphism, a phenotypic difference between males and females of the same species * Nuclear dimorp ...

, with long shoots and short shoots similar to a larch, though the short shoots are not so markedly short, lengthening about 5 mm annually. The leaves are bright green, 3–6 cm long and 2–3 mm broad, with two glaucous

stoma In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bo ...

tal bands on the underside; they turn a brilliant golden yellow before falling in the autumn, hence the common name "golden larch". The leaves are arranged spirally, widely spaced on long shoots, and in a dense whorl on the short shoots. The

cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines co ...

are distinctive, superficially resembling small

globe artichoke The globe artichoke (''Cynara cardunculus'' var. ''scolymus'' ),Rottenberg, A., and D. Zohary, 1996: "The wild ancestry of the cultivated artichoke." Genet. Res. Crop Evol. 43, 53–58. also known by the names French artichoke and green articho ...

s, 4–7 cm long and 4–6 cm broad, with pointed triangular scales; they mature about 7 months after pollination, when (like fir and cedar cones) they disintegrate to release the winged seeds. The male cones, as in ''Keteleeria'', are produced in umbels of several together in one bud.

Characteristics

The golden larch is an attractive ornamental tree for parks and large garden. Unlike the true larches, it is tolerant of summer heat and humidity, growing successfully in the southeastern United States where most larches and firs do not succeed. In Europe growth is most successful in the Mediterranean region with notable specimens in northern Italy; further north in the United Kingdom it will grow, but only very slowly due to the cooler summers. This plant has gained the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (No ...

Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...

Gallery

File:PseudolarixAmabilis.jpg File:PseudolarixAmabilisBark.jpg, Closeup of bark File:Pseudolarix amabilis seeds, by Omar Hoftun.jpg,

Seeds A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...

without wings File:Pseudolarix amabilis pollen cones.jpg, Pollen cones File:Pseudolarix amabilis female cones.jpg, Maturing female cones File:Pseudolarix amabilis in fall.jpg, Needles and mature cone in fall File:Pseudolarix amabilis needles.jpg, Needle detail in fall File:Golden Larch Bonsai Forest.JPG,

Bonsai Bonsai ( ja, 盆栽, , tray planting, ) is the Japanese art of growing and training miniature trees in pots, developed from the traditional Chinese art form of ''penjing''. Unlike ''penjing'', which utilizes traditional techniques to produce ...

example

References

External links

''Pseudolarix amabilis'' images at the Arnold Arboretum of Harvard University Plant Image Database
*Friedman, William (Ned)
"Golden larches never disappoint."
''Posts from the Collection'', Arnold Arboretum of Harvard University website, 18 November 2019. Accessed 13 May 2020. *Rose, Nancy
"Not All Conifers are Evergreen."
Arnold Arboretum of Harvard University website, 6 January 2016. Accessed 13 May 2020. *Friedman, William (Ned)
"Leaf shadows on Peter Hill."
''Posts from the Collection,'' Arnold Arboretum of Harvard University website, 18 November 2019. Accessed 13 May 2020.
''Pseudolarix'' in the ''Flora of China''Article in Harvard University Bulletin of Popular Information (1919)
{{Taxonbar, from1=Q1458175, from2=Q17359075 Pinaceae Endemic flora of China Trees of China Medicinal plants of Asia Plants used in traditional Chinese medicine Extant Eocene first appearances Endangered flora of Asia Deciduous conifers Buchanan Lake Formation Allenby Formation