Psephophorus Terrypratchetti
   HOME

TheInfoList



OR:

''Psephophorus'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
sea turtle Sea turtles (superfamily Chelonioidea), sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerh ...
that lived from the
Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
to the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
,
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, and New Zealand. It was first named by
Hermann von Meyer Christian Erich Hermann von Meyer (3 September 1801 – 2 April 1869), known as Hermann von Meyer, was a German palaeontologist. He was awarded the 1858 Wollaston medal by the Geological Society of London. Life He was born in Frankfurt am ...
in 1847, and contains seven species, ''P. polygonus'', ''P. calvertensis'', ''P. eocaenus'', ''P. oregonesis'', ''P. californiensis'', ''P. rupeliensis'',''Proceedings''; page 8. By the Zoological Society of London; published 1891. Retrieved on June 28th, 2008. ''P. scaldii'', and a species discovered in 1995, ''P. terrypratchetti''. ''Psephophorus'' is the only
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
dermochelyid turtle found in Europe. One species of ''Psephophorus'' could measure up to ten feet in length.


Discovery and identification

Von Meyer originally named ''Psephophorus'' in 1846. At first he was unable to identify the creature beyond its
dermal plate A plate in animal anatomy may refer to several things: Flat bones (examples: bony plates, dermal plates) of vertebrates * an appendage of the Stegosauria group of dinosaurs * articulated armoured plates covering the head of thorax of Placodermi ...
s, but when he later received a drawing he was able to describe the specimen, which was then in
Pressburg Bratislava (German: ''Pressburg'', Hungarian: ''Pozsony'') is the Capital city, capital and largest city of the Slovakia, Slovak Republic and the fourth largest of all List of cities and towns on the river Danube, cities on the river Danube. ...
, as a fragment of a
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
, which contained seventy
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
s. In 1879,
H. G. Seeley Harry Govier Seeley (18 February 1839 – 8 January 1909) was a British paleontologist. Early life Seeley was born in London on 18 February 1839, the second son of Richard Hovill Seeley, a goldsmith, and his second wife Mary Govier. When his fa ...
was asked to study the ''Psephophorus'' specimen by
Franz Ritter von Hauer Franz Ritter von Hauer, or Franz von Hauer (30 January 1822 – 20 March 1899) was an Austrian geologist. Biography Hauer was born in Vienna, the son of Joseph von Hauer (1778-1863), who was equally distinguished as a high Austrian official ...
, the Director of the Austro-Hungarian Imperial and Royal Geological Survey. Up until then, the specimen's identity had been undeterminable, with even Seeley describing it at first to seem like "the dermal covering of an
Edentate Xenarthra (; from Ancient Greek wikt:ξένος, ξένος, xénos, "foreign, alien" + wikt:ἄρθρον, ἄρθρον, árthron, "joint") is a Order (biology), superorder and major clade of Placentalia, placental mammals native to the America ...
closely allied to the
Armadillo Armadillos () are New World placental mammals in the order (biology), order Cingulata. They form part of the superorder Xenarthra, along with the anteaters and sloths. 21 extant species of armadillo have been described, some of which are dis ...
es." Seeley examined some bone fragments and concluded the specimen was that of a reptilian creature, furthermore a chelonid. It also proved to be more closely related to '' Sphargis'' than any other type in the
Chelonian Turtles are reptiles of the order Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), ...
order.


Relation to modern Leatherback sea turtles

For a long time, modern Leatherback sea turtles (''Dermochelys'') were believed to be descended directly from ''Psephophorus'', specifically the species ''P. polygonus''. However, a 1996 analysis by Wood et al. proved that most of the
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
in the two
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
were not connected, meaning ''Psephophorus'' could not be a direct ancestor of the modern leatherbacks. The
platelets Platelets or thrombocytes () are a part of blood whose function (along with the coagulation factors) is to react to bleeding from blood vessel injury by clumping to form a blood clot. Platelets have no cell nucleus; they are fragments of cyto ...
on ''Psephophorus'' are quite similar to those on '' Dermochelys'', despite differences in outer
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
and size. The platelet comprises an external compact layer and an internal zone of
cancellous bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, an ...
.


Species

*''Psephophorus polygonus'' is the type
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, and was discovered by von Meyer in 1846.
Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
of the species have shown bony, carapacial ridges, adjacent, small and polygonal bony ossicles which measure about 22 × 25 mm. When the ossicles extend further along the longitudinal axis, their size increases up to an average of 33 × 41 mm. *''Psephophorus calvertensis'' was first named by Palmer in 1909. It was so named for being found in the
Calvert Formation The Calvert Formation is a Formation (geology), geologic formation in Maryland, Virginia, and Delaware. It preserves fossils dating back to the early to middle Miocene, Miocene epoch of the Neogene Period (geology), period. It is one of the three ...
. ''P. calvertensis'' is a rarely found specimen. *''Psephophorus eocaenus'' was first named by Andrews in 1901. *''Psephophorus californiensis'' was first named by Gilmore in 1937. *''Psephophorus terrypratchetti'' was discovered in the 1990s by
Richard Köhler Richard is a male given name. It originates, via Old French, from Frankish language, Old Frankish and is a Compound (linguistics), compound of the words descending from Proto-Germanic language, Proto-Germanic ''*rīk-'' 'ruler, leader, king' an ...
in New Zealand and named in 1995 after the author
Terry Pratchett Sir Terence David John Pratchett (28 April 1948 – 12 March 2015) was an English author, humorist, and Satire, satirist, best known for the ''Discworld'' series of 41 comic fantasy novels published between 1983 and 2015, and for the Apocalyp ...
, who wrote a series of
fantasy books Fantasy literature is literature set in an imaginary universe, often but not always without any locations, events, or people from the real world. magic (paranormal), Magic, the supernatural and Legendary creature, magical creatures are common i ...
set on a world carried on the back of a giant turtle. This species reached in body length which is more than 80% bigger than ''P. eocaenus''.


See also

*
List of organisms named after famous people (born 1900–1949) In biological nomenclature, organisms often receive scientific names that honor a person. A taxon (e.g., species or genus; plural: taxa) named in honor of another entity is an eponymous taxon, and names specifically honoring a person or persons ...
* ''
Archelon ''Archelon'' is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring from head to tail and in body mass. It is known only from the Pierre Shale and ...
'' * '' Gigantatypus'' *
Largest prehistoric animals The largest prehistoric animals include both vertebrate and invertebrate species. Many of them are described below, along with their typical range of size (for the general dates of extinction, see the link to each). Many species mentioned might ...


References


Further reading

* * * *


External links

* * * * * * {{Taxonbar, from=Q7254102 Dermochelyidae Fossil taxa described in 1995 Terry Pratchett Oligocene turtles Miocene turtles Pliocene turtles Cenozoic reptiles of Europe Cenozoic reptiles of North America Oligocene genus first appearances Pliocene genus extinctions Prehistoric turtle genera Taxa named by Christian Erich Hermann von Meyer Extinct turtles