Prozostrodontia is a

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

cynodont Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 Megaannum, mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extin ...

s including

mammaliaform Mammaliaformes ("mammalian forms") is a clade of synapsid tetrapods that includes the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts during the Late Triassic. It is defined a ...

s and their closest relatives such as

Tritheledontidae Tritheledontidae, the tritheledontids or ictidosaurs, is an extinct family of small to medium-sized (about 10 to 20 cm long) cynodonts. They were highly mammal-like, specialized cynodonts, although they still retained a few reptile-like ...

and

Tritylodontidae Tritylodontidae ("three-knob teeth", named after the shape of their cheek teeth) is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy, and some d ...

. It was erected as a

node-based taxon Phylogenetic nomenclature is a method of nomenclature for taxon, taxa in biology that uses phylogenetics, phylogenetic definitions for taxon names as explained below. This contrasts with Biological classification, the traditional method, by which ...

by Liu and Olsen (2010) and defined as the least inclusive clade containing ''

Prozostrodon brasiliensis ''Prozostrodon'' is an extinct genus of probainognathian cynodonts that was closely related to mammals. The remains were found in Brazil and are dated to the Carnian age of the Late Triassic. The holotype has an estimated skull length of , ind ...

'', ''

Tritylodon langaevus ''Tritylodon'' (from the Greek for "three-cusped tooth") is an extinct genus of tritylodonts, one of the most advanced group of cynodont therapsids. They lived in the Early Jurassic and possibly Late Triassic periods along with dinosaurs. They ...

'', ''

Pachygenelus monus ''Pachygenelus'' is a genus of extinct cynodont. Fossils have been found from the Karoo basin in South Africa and date back to the Early Jurassic. The genus was named in 1913 on the basis of a partial lower jaw found from South Africa, with the ...

'', and ''

Mus musculus The house mouse (''Mus musculus'') is a small mammal of the rodent family Muridae, characteristically having a pointed snout, large rounded ears, and a long and almost hairless tail. It is one of the most abundant species of the genus ''Mus (genu ...

'' (the house mouse). Prozostrodontia is diagnosed by several characters, including: * Reduced

prefrontal Prefrontal may refer to: *Prefrontal bone, a skull bone in some tetrapods *Prefrontal cortex, a region of the brain of a mammal *Prefrontal scales The prefrontal scales on snakes and other reptiles are the scales adjacent and anterior to the fr ...

and

postorbital bone The ''postorbital'' is one of the bones in vertebrate skulls which forms a portion of the dermal skull roof and, sometimes, a ring about the orbit. Generally, it is located behind the postfrontal and posteriorly to the orbital fenestra. In some ...

s, with the disappearance of a strut of bone called the

postorbital bar The postorbital bar (or postorbital bone) is a bony arched structure that connects the frontal bone of the skull to the zygomatic arch, which runs laterally around the eye socket. It is a trait that only occurs in mammalian taxa, such as most strep ...

separating the

eye socket In anatomy, the orbit is the cavity or socket/hole of the skull in which the eye and its appendages are situated. "Orbit" can refer to the bony socket, or it can also be used to imply the contents. In the adult human, the volume of the orbit is ...

from the temporal region * Unfused

symphysis A symphysis (, : symphyses) is a fibrocartilaginous fusion between two bones. It is a type of cartilaginous joint, specifically a secondary cartilaginous joint. # A symphysis is an amphiarthrosis, a slightly movable joint. # A growing together o ...

between the

dentary bone In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

s in the lower jaw * The presence of a small hole within the

called the

sphenopalatine foramen The sphenopalatine foramen is a foramen of the skull that connects the nasal cavity and the pterygopalatine fossa. It gives passage to the sphenopalatine artery, nasopalatine nerve, and the superior nasal nerve (all passing from the pterygopala ...

* A long

sagittal crest A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are excepti ...

extending to the rearmost part of the lambdoidal crest at the back of the skull *

Neural spine Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...

s of the

dorsal vertebrae In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebra (anatomy), vertebrae of intermediate size between the ce ...

angled backward * A convex-shaped

iliac crest The crest of the ilium (or iliac crest) is the superior border of the wing of ilium and the superolateral margin of the greater pelvis. Structure The iliac crest stretches posteriorly from the anterior superior iliac spine (ASIS) to the posterio ...

and a reduced posterior iliac spine on the hip * An

acetabular notch The acetabular notch is a deep notch in the inferior portion of the rim of the acetabulum The acetabulum (; : acetabula), also called the cotyloid cavity, is a wikt:concave, concave surface of the pelvis. The femur head, head of the femur mee ...

on the

ischium The ischium (; : is ...

(a groove in the hip socket) * The position of a small projection called the

lesser trochanter In human anatomy, the lesser trochanter is a conical, posteromedial, bony projection from the shaft of the femur. It serves as the principal insertion site of the iliopsoas muscle. Structure The lesser trochanter is a conical posteromedial p ...

close to the head of the

femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...

Prozostrodontia includes tritylodontids, which have traditionally been placed within the more primitive cynodont group

Cynognathia Cynognathia ("dog jaw") is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus '' Cynognathus'' and the herbivorous or omnivorous gomphodonts such as traversodontids. Cynogn ...

as distant mammal relatives. It also includes Tritheledontidae, which has long been placed close to mammals. Most previous studies considered Tritheledontidae a valid

monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...

grouping, meaning it was a true

including all the descendants of a single common ancestor, but Liu and Olsen (2010) found Tritheledontidae to be a

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

series of basal prozostrodontians. Below is a

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

from Liu and Olsen (2010) showing the

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

position of Prozostrodontia:

References

{{Taxonbar, from=Q5852697 Carnian first appearances Extant Late Triassic first appearances