Prototaria
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''Prototaria'' is an extinct genus of
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant taxon, extant families Odobenidae (whose onl ...
that lived approximately 15.97 to 13.65 mya during the
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene epoch (geology), epoch made up of two Stage (stratigraphy), stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0. ...
in what is now
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
. It belonged to the family
Odobenidae Odobenidae is a family of pinnipeds, of which the only extant species is the walrus (''Odobenus rosmarus''). In the past, however, the group was much more diverse, and includes more than a dozen fossil genera. Taxonomy All genera, except '' Od ...
, the only extant species of which is the
walrus The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...
. Members of the genus ''Prototaria'' are believed to be the most basal imagotariine pinnipeds. Unlike their closest living relative, the walrus, members of ''Prototaria'' were primarily
piscivorous A piscivore () is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted rept ...
carnivores.


Description

''Prototaria'' were more similar in appearance to modern
fur seals Fur seals are any of nine species of pinnipeds belonging to the subfamily Arctocephalinae in the family Otariidae. They are much more closely related to sea lions than true seals, and share with them external ears ( pinnae), relatively long a ...
and
sea lions Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...
than walruses. They did not have long tusks as walruses do, and were more slender. Fossils ascribed to ''Prototaria'' exhibit some
autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...
, including: a large antorbital process and
orbit In celestial mechanics, an orbit (also known as orbital revolution) is the curved trajectory of an object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an artificial satellite around an ...
, and a relatively narrow intertemporal region of the skull. ''P. planicephala'' has a wider
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit ** podium * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ...
than ''P. primigena'' as well as more specialized
cheek teeth Cheek teeth or postcanines comprise the molar and premolar teeth in mammals. Cheek teeth are multicuspidate (having many folds or tubercles). Mammals have multicuspidate molars (three in placentals, four in marsupials, in each jaw quadrant) and ...
and a flatter
tympanic bulla The tympanic part of the temporal bone is a curved plate of bone lying below the squamous part of the temporal bone, in front of the mastoid process, and surrounding the external part of the ear canal. It originates as a separate bone (tympanic ...
.


Taxonomy

The type species of ''Prototaria'' is ''P. primigena''. Based on skeletal traits and various autapomorphies, Naoki Kohno concluded that ''P. planicephala'' is more derived than ''P. primigena''.


References

{{Taxonbar, from=Q64605261 Odobenids Prehistoric carnivoran genera Fossil taxa described in 1984