Prosopium Williamsoni
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The mountain whitefish (''Prosopium williamsoni'') is one of the most widely distributed
salmonid Salmonidae (, ) is a family of ray-finned fish, the only extant member of the suborder Salmonoidei, consisting of 11 extant genera and over 200 species collectively known as "salmonids" or "salmonoids". The family includes salmon (both Atlantic a ...
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
of western
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. It is found from the
Mackenzie River The Mackenzie River (French: ; Slavey language, Slavey: ' èh tʃʰò literally ''big river''; Inuvialuktun: ' uËkpÉ‘k literally ''great river'') is a river in the Canadian Canadian boreal forest, boreal forest and tundra. It forms, ...
drainage in
Northwest Territories The Northwest Territories is a federal Provinces and territories of Canada, territory of Canada. At a land area of approximately and a 2021 census population of 41,070, it is the second-largest and the most populous of Provinces and territorie ...
, Canada through western Canada and the northwestern USA in the Pacific, Hudson Bay and upper Missouri River basins to the
Truckee River The Truckee River is a river in the U.S. states of California and Nevada. The river flows northeasterly and is long.U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map accessed October 20, 2012 Th ...
drainage in
Nevada Nevada ( ; ) is a landlocked state in the Western United States. It borders Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the seventh-most extensive, th ...
and Sevier River drainage in
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
.


Description

The body shape is superficially similar to the
cyprinid Cyprinidae is a Family (biology), family of freshwater fish commonly called the carp or minnow family, including the carps, the true minnows, and their relatives the barb (fish), barbs and barbel (fish), barbels, among others. Cyprinidae is the ...
s, although it is distinguished by having both the
adipose fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only b ...
and axillary process of
salmonids Salmonidae (, ) is a family of ray-finned fish, the only extant member of the suborder Salmonoidei, consisting of 11 extant genera and over 200 species collectively known as "salmonids" or "salmonoids". The family includes salmon (both Atlantic a ...
. The body is slender and nearly cylindrical in cross section, generally silver with a dusky olive-green shade dorsally. The scales possess pigmented borders, which are especially defined on the posterior end. Mountain whitefish possess a forked homocercal tail. The short head has a small mouth underneath the
snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, Rostrum (anatomy), rostrum, beak or proboscis. The wet furless surface around the nostrils of the n ...
. The short
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
has 12–13 rays, with 11–13 for the
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
, 10–12 for the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...
s, and 14–18 for the
pectoral fins Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only b ...
. Size has been recorded at up to 70 centimeters (28 inches) in length and a weight of 2.9 kilograms (6.4 lb).


Life history

The
spawning Spawn is the Egg cell, eggs and Spermatozoa, sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is ...
season is from October to early December, when water temperatures are 2–6 Â°C. Mountain whitefish congregate in large schools on fall spawning runs and seek out areas of coarse gravels or cobbles at depths of at least 75 cm (30 inches), typically in shallow areas of small tributaries or shorelines of lakes. Their non-adhesive eggs are scattered along the substrate. The eggs then develop slowly through the winter (6–10 weeks), hatching in the early spring, generally in March. Mountain whitefish reach reproductive maturity at approximately three years old, females can produce as many as 4,000 eggs annually. Mountain whitefish typically live between 7–9 years in the wild.


Feeding

Mountain whitefish are demersal feeders, stirring up the substrate with pectoral and tail fins to expose insect larvae and other invertebrates, including
snail A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...
s,
crayfish Crayfish are freshwater crustaceans belonging to the infraorder Astacidea, which also contains lobsters. Taxonomically, they are members of the superfamilies Astacoidea and Parastacoidea. They breathe through feather-like gills. Some spe ...
, and
amphipod Amphipoda () is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods () range in size from and are mostly detritivores or scavengers. There are more than 10,700 amphipod species cur ...
s. Their main feeding time is in the evening, but they will also take drifting prey during the day. The mountain whitefish frequently feeds in the lower strata of streams, but populations may rise to the surface to prey on hatching insects, including mayflies. Mountain whitefish fry are a common food source for other trout species.


Ecology


Habitat

The mountain whitefish are commonly found in mountain streams and lakes, favoring clear cold water and large deep pools of at least a meter's depth; the
Lake Tahoe Lake Tahoe (; Washo language, Washo: ''dáʔaw'') is a Fresh water, freshwater lake in the Sierra Nevada of the Western United States, straddling the border between California and Nevada. Lying at above sea level, Lake Tahoe is the largest a ...
population lives just above the bottom in deeper water.


Range

This species occurs throughout the western half of North America, as far north as the
Mackenzie River The Mackenzie River (French: ; Slavey language, Slavey: ' èh tʃʰò literally ''big river''; Inuvialuktun: ' uËkpÉ‘k literally ''great river'') is a river in the Canadian Canadian boreal forest, boreal forest and tundra. It forms, ...
(Canada) and the drainages of the
Hudson Bay Hudson Bay, sometimes called Hudson's Bay (usually historically), is a large body of Saline water, saltwater in northeastern Canada with a surface area of . It is located north of Ontario, west of Quebec, northeast of Manitoba, and southeast o ...
, in the
Columbia River The Columbia River (Upper Chinook language, Upper Chinook: ' or '; Sahaptin language, Sahaptin: ''Nch’i-Wàna'' or ''Nchi wana''; Sinixt dialect'' '') is the largest river in the Pacific Northwest region of North America. The river headwater ...
, upper
Missouri River The Missouri River is a river in the Central United States, Central and Mountain states, Mountain West regions of the United States. The nation's longest, it rises in the eastern Centennial Mountains of the Bitterroot Range of the Rocky Moun ...
, upper
Colorado River The Colorado River () is one of the principal rivers (along with the Rio Grande) in the Southwestern United States and in northern Mexico. The river, the List of longest rivers of the United States (by main stem), 5th longest in the United St ...
, Adirondack Lakes of New York, and Lake Champlain.


Migration

Mountain whitefish are non-
anadromous Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousa ...
fish and spend the majority of their lives in freshwater ecosystems. Relatively little is known about the migration patterns of whitefish, but it is believed that they do migrate for spawning and because of temperature fluctuations. In the
Methow River The Methow River ( ) is a tributary of the Columbia River in northern Washington in the United States. The river's watershed drains the eastern North Cascades, with a population of about 5,000 people. The Methow's watershed is characterized by ...
, a
tributary A tributary, or an ''affluent'', is a stream or river that flows into a larger stream (''main stem'' or ''"parent"''), river, or a lake. A tributary does not flow directly into a sea or ocean. Tributaries, and the main stem river into which they ...
of the Columbia River, mountain whitefish have been found migrating due to increases in shallower tributary water temperatures into the larger Columbia River. Mountain whitefish have also been known to spend the winter in the Columbia River which provides more suitable habitats for these fishes. Evidence shows that mountain whitefish in the Methow River are returning annually to the same
headwater The headwater of a river or stream is the geographical point of its beginning, specifically where surface runoff water begins to accumulate into a flowing channel of water. A river or stream into which one or many tributary rivers or streams flo ...
tributaries, which indicates migration for the primary purpose of returning to birth streams to spawn – similar to other anadromous salmonid species.


Aging fish

To better understand the conservation and management needs of mountain whitefish biologists seek to understand life history strategies of these fishes. Understanding the growth, age structure, and age at maturity of these fish can better inform limitations on which fish can be harvested and which should be released. If the age at which the fish matures is known, management agencies can estimate sizes of these fishes and inform anglers that no fish under this maturation size may be kept to ensure new generations. The age of these fish can be determined by counting annuli (growth rings) in fish scales, pectoral
fin ray Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only b ...
s, and
otolith An otolith (, ' ear + , ', a stone), also called otoconium, statolith, or statoconium, is a calcium carbonate structure in the saccule or utricle (ear), utricle of the inner ear, specifically in the vestibular system of vertebrates. The saccule ...
s (ear bones). Efficacy of these counting strategies can vary, but in combination they could provide vital data to ensure that mountain whitefish populations continue to stay at healthy levels. Some of the most effective ways to age mountain whitefish have been determined to be by examining scales for younger fishes or otoliths for older fishes.


Conservation

Mountain whitefish have a secure conservation status. In many of the larger intermountain Western rivers, mountain whitefish are the only native salmonid. Mountain whitefish typically occur in high abundance, in the
Snake River Basin The canyons">Snake River cutting through the plain leaves many canyons and Canyon#List of gorges">gorges, such as this one near Twin Falls, Idaho The Snake River Plain is a geologic feature located primarily within the U.S. state of Idaho. ...
in
Idaho Idaho ( ) is a landlocked U.S. state, state in the Pacific Northwest and Mountain states, Mountain West subregions of the Western United States. It borders Montana and Wyoming to the east, Nevada and Utah to the south, and Washington (state), ...
, mountain whitefish abundance was found to be 1,257/100 m. Historically, mountain whitefish are not popular
game fish Game fish, sport fish or quarry refer to popular fish species pursued by recreational fishing, recreational fishers (typically angling, anglers), and can be freshwater fish, freshwater or saltwater fish. Game fish can be fish as food, eaten aft ...
and have been thrown onto the banks when caught because they are 'trash fish.' Similar to other native salmonids, mountain whitefish have received backlash because anglers believe that they are competing for food and spawning resources for more popular introduced species, like
brook trout The brook trout (''Salvelinus fontinalis'') is a species of freshwater fish in the char genus ''Salvelinus'' of the salmon family Salmonidae native to Eastern North America in the United States and Canada. Two ecological forms of brook trout h ...
(Idaho). Due to common misconceptions, some native mountain whitefish populations have been decimated by both management agencies and anglers alike. The health of mountain whitefish, like other salmonids, can be a predictor of the health of the environment that they live in, so understanding and maintaining their populations can be beneficial for ecosystem prosperity.


References

{{Authority control Prosopium Freshwater fish of the United States Fish of Canada Fish of the Western United States Fauna of the Northwestern United States Fauna of California Fauna of the Great Basin Fauna of the Rocky Mountains Taxa named by Charles Frédéric Girard Fish described in 1856