''Propalaeotherium'' was an early
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of perissodactyl
endemic
Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Europe
Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and
Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
during the
early Eocene
In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...
. There are currently six recognised species within the genus, with ''P. isselanum'' as the
type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
(named by
Georges Cuvier
Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
in 1824).
Taxonomy

''Propalaeotherium'' was named by
Paul Gervais
Paul Gervais (full name: François Louis Paul Gervais) (26 September 1816 – 10 February 1879) was a French palaeontologist and entomologist.
Biography
Gervais was born in Paris, where he obtained the diplomas of doctor of science and of medic ...
; its name means "before ''
Palaeotherium
''Palaeotherium'' is an extinct genus of Equoidea, equoid that lived in Europe and possibly the Middle East from the Middle Eocene to the Early Oligocene. It is the type genus of the Palaeotheriidae, a group exclusive to the Paleogene, Palaeogen ...
''". It was considered a member of
Palaeotheriidae
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals that inhabited Europe, with less abundant remains also known from Asia, from the mid-Eocene to the early Oligocene. They are classified in Equoidea, along with the livin ...
by Hooker (1986). A 2004 study found it to be an equid instead. A 2016 study lumped the genus back within the
Palaeotheriidae
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals that inhabited Europe, with less abundant remains also known from Asia, from the mid-Eocene to the early Oligocene. They are classified in Equoidea, along with the livin ...
.
The species ''P. parvulum'' and ''P. messelensis'' have been alternately assigned to the equid genus ''
Eurohippus
''Eurohippus'' is an extinct genus of equoid ungulate
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with t ...
''.
Description
''Propalaeotherium'' was a small animal, ranging from 30–60 cm at the shoulder (2.9 to 5.9
hands
A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the koala (which has two opposable thumbs on each "han ...
), and weighing just . Despite being an early form of
horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
, it looked similar to small
tapirs
Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
. It had no
hooves
The hoof (: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits; the ruminants with ...
, but instead several small nail-like hooflets. The well-preserved
Messel fossils showed their
herbivory
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
, specifically their preference to eat
berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...
and leaf matter picked up from the forest floor.
See also
*
Evolution of the horse
The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling '' Eohippus'' into the modern horse. Paleozoologists have been able to piece ...
*
List of prehistoric mammals
This is an incomplete list of prehistoric mammals. It does not include List of mammals, extant mammals or List of extinct mammals, recently extinct mammals. For extinct primate species, see: list of fossil primates.Mikko's Phylogeny Archiv
Ma ...
References
External links
BBC Online Science and Nature
{{Taxonbar, from=Q133295
Palaeotheriidae
Eocene horses
Eocene genus extinctions
Eocene mammals of Europe
Prehistoric placental genera
Eocene mammals of Asia
Fossil taxa described in 1849