''Prolibytherium'' is an extinct genus of prolibytheriid

artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poste ...

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraff ...

native to Middle

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...

North

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

and

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...

, from around 16.9 to 15.97 million years ago. Fossils of ''Prolibytherium'' were found in the

Marada Formation The Mardaites () or al-Jarajima ( syr, ܡܪ̈ܕܝܐ; ar, ٱلْجَرَاجِمَة / ALA-LC: ''al-Jarājimah''), inhabited the highland regions of the Nur Mountains. The Mardaites were early Christians following either Miaphysitism or Monoth ...

Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Su ...

, Vihowa Formation of Pakistan, and the Moghara Formation of

Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Med ...

Description

The long creature would have superficially resembled an

okapi The okapi (; ''Okapia johnstoni''), also known as the forest giraffe, Congolese giraffe, or zebra giraffe, is an artiodactyl mammal that is endemic to the northeast Democratic Republic of the Congo in central Africa. It is the only species ...

or a deer. Unlike these, however, ''Prolibytherium'' displayed dramatic

sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...

, in that the male had a set of large,

leaf A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

-shaped

ossicone Ossicones are columnar or conical skin-covered bone structures on the heads of giraffes, male okapi, and some of their extinct relatives. Ossicones are distinguished from the superficially similar structures of horns and antlers by their uniqu ...

s with a width of , while the female had a set of slender, horn-like ossicones. The taxonomic status of ''Prolibytherium'' remains in flux. At one time, it was described as a relative of ''

Sivatherium ''Sivatherium'' ("Shiva's beast", from Shiva and ''therium'', Latinized form of Ancient Greek θηρίον -'' thēríon'') is an extinct genus of giraffids that ranged throughout Africa to the Indian subcontinent. The species ''Sivatherium giga ...

'' (as a precursor to "Libytherium maurusium" (''S. maurusium'')). Later, it would be regarded as a palaeomerycid, or either as a climacoceratid, or as a basal member of

Giraffoidea Giraffoidea is a superfamily that includes the families Climacoceratidae, Antilocapridae, and Giraffidae. The only extant members in the superfamily are the pronghorn, giraffe, and okapi. The Climacoceratidae are also placed in the superfami ...

. With the discovery and study of a female skull in 2010, ''Prolibytherium'' is tentatively confirmed as a climacoceratid. A recent study published in 2022 found it to be part of a separate family, Prolibytheriidae.

References

External links

* Prehistoric even-toed ungulate genera Mammal enigmatic taxa Miocene mammals of Africa Fossils of Egypt Fossils of Libya Miocene mammals of Asia Fossils of Pakistan Fossil taxa described in 1961 {{Paleo-eventoedungulate-stub