''Proardea'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

heron Herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 75 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genus ''Botaurus'' are referred to as bi ...

, containing two

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

, ''Proardea amissa'' ("lost proto-heron") and ''Proardea? deschutteri'' from the Borgloon Formation of

Belgium Belgium, officially the Kingdom of Belgium, is a country in Northwestern Europe. Situated in a coastal lowland region known as the Low Countries, it is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeas ...

. It stood about 70 cm (2 ft 4 in) tall and was very similar to a modern

in shape. The species is known from rather fragmentary

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s in the area of

Quercy Quercy (; , locally ) is a former province of France located in the country's southwest, bounded on the north by Limousin, on the west by Périgord and Agenais, on the south by Gascony and Languedoc, and on the east by Rouergue and Auverg ...

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

; dated remains are from Pech Desse, a Late

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

locality, but the original fossil, a single right

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...

(

MNHN The French National Museum of Natural History ( ; abbr. MNHN) is the national natural history museum of France and a of higher education part of Sorbonne University. The main museum, with four galleries, is located in Paris, France, within the ...

QU-15720), isn't precisely dated and may have come from deposits as early as Late

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

in age.

''Proardea'' was apparently closely related to the true herons and egrets (Ardeinae). As these genera are only known from the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

onwards, ''Proardea'' possibly was a direct ancestor of today's herons and/or egrets. However, the Miocene genus '' Proardeola'' is closely related, or perhaps even synonymous, with ''Proardea''; the former's single species ''Proardeola walkeri'' may thus be ''Proardea walkeri'' or even identical with ''P. amissa''. The bird described as '' Ardea aurelianensis'' may also be identical with ''P. amissa'', which would in that case become known as ''Proardea aurelianensis''. Supposed other species of ''Proardea'', ''P. perplexa'' and ''P. similis'', are synonyms of the

ibis The ibis () (collective plural ibises; classical plurals ibides and ibes) are a group of long-legged wading birds in the family Threskiornithidae that inhabit wetlands, forests and plains. "Ibis" derives from the Latin and Ancient Greek word f ...

'' Geronticus perplexus'' and the phasianid '' Miogallus altus'', respectively.

References

Further reading