The principle of faunal succession, also known as the law of faunal succession, is based on the observation that sedimentary rock

strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...

contain

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

ized

flora Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. The corresponding term for animals is ''fauna'', and for f ...

and

fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...

, and that these fossils succeed each other vertically in a specific, reliable order that can be identified over wide horizontal distances. A fossilized

Neanderthal Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinction, extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle Pleistocene, Middle to Late Plei ...

bone (less than 500,000 years old) will never be found in the same stratum as a fossilized

Megalosaurus ''Megalosaurus'' (meaning "great lizard", from Ancient Greek, Greek , ', meaning 'big', 'tall' or 'great' and , ', meaning 'lizard') is an extinct genus of large carnivorous theropod dinosaurs of the Middle Jurassic Epoch (Bathonian stage, 166 ...

(about 160 million years old), for example, because neanderthals and megalosaurs lived during different

geological periods The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronolo ...

, separated by millions of years. This allows for strata to be identified and dated by the fossils found within. This principle, which received its name from the English

geologist A geologist is a scientist who studies the structure, composition, and History of Earth, history of Earth. Geologists incorporate techniques from physics, chemistry, biology, mathematics, and geography to perform research in the Field research, ...

William Smith, is of great importance in determining the relative age of rocks and strata. The fossil content of rocks together with the

law of superposition The law of superposition is an axiom that forms one of the bases of the sciences of geology, archaeology, and other fields pertaining to geological stratigraphy. In its plainest form, it states that in undeformed stratigraphic sequences, the ...

helps to determine the time sequence in which sedimentary rocks were laid down.

Evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...

explains the observed faunal and floral succession preserved in rocks. Faunal succession was documented by Smith in England during the first decade of the 19th century, and concurrently in France by Cuvier (with the assistance of the mineralogist

Alexandre Brongniart Alexandre Brongniart (5 February 17707 October 1847) was a French chemist, mineralogist, geologist, paleontologist, and zoologist, who collaborated with Georges Cuvier on a study of the geology of the region around Paris. Observing fossil conten ...

). Archaic biological features and organisms are succeeded in the

fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

by more modern versions. For instance, paleontologists investigating the evolution of

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s predicted that

feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...

s would first be seen in primitive forms on flightless predecessor organisms such as feathered dinosaurs. This is precisely what has been discovered in the fossil record: simple feathers, incapable of supporting flight, are succeeded by increasingly large and complex feathers. In practice, the most useful diagnostic species are those with the fastest rate of species turnover and the widest distribution; their study is termed

biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. "Biostratigraphy." ''Oxford Reference: Dictionary of Biology ...

, the science of dating rocks by using the

s contained within them. In

Cenozoic The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...

strata, fossilized tests of

foraminifera Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are unicellular organism, single-celled organisms, members of a phylum or class (biology), class of Rhizarian protists characterized by streaming granular Ectoplasm (cell bio ...

are often used to determine faunal succession on a refined scale, each biostratigraphic unit ( biozone) being a geological stratum that is defined on the basis of its characteristic fossil

taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

. An outline microfaunal zonal scheme based on both foraminifera and

ostracod Ostracods, or ostracodes, are a Class (biology), class of the crustacean, Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 33,000 species (only 13,000 of which are extant taxon, extant) have been identified,Brandão, S.N.; Antoni ...

a was compiled by M. B. Hart (1972). Earlier fossil life forms are simpler than more recent forms, and more recent fossil forms are more similar to living forms (principle of faunal succession).''Evolutionary Analysis'', 4th Edition. p 61.

References

{{Geologic Principles Paleontological concepts and hypotheses Dating methods Stratigraphy de:Biostratigraphie#Das Prinzip der Fossilfolge

See also

References