''Prejanopterus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

pterodactyloid Pterodactyloidea ( ; derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contai ...

pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...

from the

Lower Cretaceous Lower may refer to: * ''Lower'' (album), 2025 album by Benjamin Booker * Lower (surname) * Lower Township, New Jersey *Lower Receiver (firearms) * Lower Wick Gloucestershire, England See also * Nizhny {{Disambiguation ...

(lower

Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...

age) layers of the Leza Formation of

La Rioja La Rioja () is an autonomous communities in Spain, autonomous community and provinces of Spain, province in Spain, in the north of the Iberian Peninsula. Its capital is Logroño. Other List of municipalities in La Rioja, cities and towns in the ...

Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...

Discovery and naming

In 1980 a fossil site was discovered near Yacimiento de Fuente Amarga on the western slopes of the Peña Isasa. In 1993 and 1994 ten fossiliferous blocks were recovered, containing numerous disarticulated fossil bones and bone fragments, among them those of a pterosaur. The

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

''Prejanopterus curvirostra'' was named and described in

2010 The year saw a multitude of natural and environmental disasters such as the 2010 Haiti earthquake, the Deepwater Horizon oil spill, and the 2010 Chile earthquake. The 2009 swine flu pandemic, swine flu pandemic which began the previous year ...

by Carolina Fuentes Vidarte and Manuel Meijide Calvo. The generic name is derived from the village of Préjano and a Latinised Greek ''pteron'', "wing". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

is derived from

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

''curvus'', "bent", and ''rostrum'', "snout", a reference to the distinctive bend to the left which was claimed to be present in all fossils of the snout. A later study indicates that "there is no genuine sideways bend" of the snout, "but a slight dorsal curvature". The same study emended the specific name to ''curvirostris'' as the feminine ''curvirostra'' had been incorrect. The

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

, F. A. 112, consists of a fragmented snout. Another snout, specimen F. A. 185, is the

paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...

. Numerous other elements of the skull and the

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

have been assigned to the species, mostly from the wing, the pelvis and the hindlimbs. A

humerus The humerus (; : humeri) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius (bone), radius and ulna, and consists of three sections. The humeral upper extrem ...

vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...

e are still lacking. The species represents the best preserved pterosaur known from Spain and the first named from the Early Cretaceous of that country. The bent snout features twenty pairs of small teeth with an oval cross-section. That the curvature is no preservation artefact, a ''

post mortem An autopsy (also referred to as post-mortem examination, obduction, necropsy, or autopsia cadaverum) is a surgical procedure that consists of a thorough examination of a corpse by dissection to determine the cause, mode, and manner of death; ...

'' distortion, is indicated by the fact that both known snouts show it. Remarkably, all lower jaws found are straight. The wing span was initially estimated at ; however, later studies indicate that the wing span of ''Prejanopterus'' was probably not much (if ever) in excess of .

Systematics

Vidarte and Calvo (2010) assigned ''Prejanopterus'' to

Pterodactyloidea Pterodactyloidea ( ; derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contai ...

. Using the

comparative method In linguistics, the comparative method is a technique for studying the development of languages by performing a feature-by-feature comparison of two or more languages with common descent from a shared ancestor and then extrapolating backwards ...

they established that no pterodactyloid group has obvious strong affinities to the species. In a later phylogenetic study Pereda-Suberbiola ''et al.'' (2012) recovered ''Prejanopterus'' as a pterodactylid, more precisely positioned between the genus ''

Pterodactylus ''Pterodactylus'' (from ) is a genus of extinct pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying reptile and one of the first prehis ...

'' and the group

Ctenochasmatoidea Archaeopterodactyloidea (meaning "ancient Pterodactyloidea") is an extinct clade of pterodactyloid pterosaurs that lived from the middle Late Jurassic to the latest Early Cretaceous periods (Kimmeridgian to Albian stages) of Africa, Asia, Europe a ...

. Witton (2013) saw the genus as a possible lonchodectid due to its unusual tooth socket morphology and long, low nature of the jaw.Witton, Mark P. (2013). ''Pterosaurs: Natural History, Evolution, Anatomy''

Cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

after Pereda-Suberbiola ''et al.'' (2012):

References

{{Portal bar, Paleontology, Spain Pterodactyloidea Early Cretaceous pterosaurs of Europe Fossil taxa described in 2010