Populus Sect. Tacamahaca
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The balsam poplars (''Populus'' sect. ''Tacamahaca'') are a group of about 10 species of
poplars ''Populus'' is a genus of 25–30 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar (), aspen, and cottonwood. The we ...
, indigenous to
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
and eastern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, distinguished by the
balsam Balsam is the resinous exudate (or sap) which forms on certain kinds of trees and shrubs. Balsam (from Latin ''balsamum'' "gum of the balsam tree," ultimately from a Semitic source such as ) owes its name to the biblical Balm of Gilead. Ch ...
scent of their
bud In botany, a bud is an undeveloped or Plant embryogenesis, embryonic Shoot (botany), shoot and normally occurs in the axil of a leaf or at the tip of a Plant stem, stem. Once formed, a bud may remain for some time in a dormancy, dormant conditi ...
s, the whitish undersides of their
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
, and the leaf petiole being round (not flattened) in cross-section. They are large
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s, 30–60 m tall, with leaves with a rounded base, pointed apex, and a whitish
wax Waxes are a diverse class of organic compounds that are lipophilic, malleable solids near ambient temperatures. They include higher alkanes and lipids, typically with melting points above about 40 °C (104 °F), melting to give lo ...
y coating on the underside of the leaf; this latter distinguishes them from most other poplars. The name is derived from the pleasant balsam smell of the opening buds and leaves in spring, produced by a sticky gum on the buds which also helps protect the buds from
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
damage. The balsam poplars are light-demanding trees that require considerable moisture. Balsam poplars are tolerant of very cold conditions, occurring further north than other poplars except for the
aspen Aspen is a common name for certain tree species in the Populus sect. Populus, of the ''Populus'' (poplar) genus. Species These species are called aspens: * ''Populus adenopoda'' – Chinese aspen (China, south of ''P. tremula'') * ''Populus da ...
s. The poplars in Southern California are tolerant of 100 plus degree heat. They grow along dry washes and dry riverbed's. The dry washes and dry riverbeds will have flowing water when it rains sufficiently. Their leaves hang down and are at an edge to the sun. This may be another factor why they can take the high heat. Their leaves tremble in the slightest breeze like the quaking aspen


Species

The balsam poplar '' P. balsamifera'' (= ''P. tacamahaca'', ''P. candicans'') is a native of North America, where it grows on alluvial bottomlands in the northeastern United States and Canada. It grows to a height of 30 metres and has yellow-grey bark, thick and furrowed, and coloured blackish at the base of the trunk. The twigs are yellow-brown to brown, the buds covered with a layer of balsam resin. The flowers and fruit are very much like those of the white poplar ('' P. alba'') which is a relative of the
aspen Aspen is a common name for certain tree species in the Populus sect. Populus, of the ''Populus'' (poplar) genus. Species These species are called aspens: * ''Populus adenopoda'' – Chinese aspen (China, south of ''P. tremula'') * ''Populus da ...
s (''Populus'' sect. ''Populus''). The western balsam poplar, black cottonwood, or California poplar '' P. trichocarpa'' is native to western North America, from
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
south to northern
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
. It is the largest species of poplar, recorded to 65 m tall. It is also a very important species in
plant biology Botany, also called plant science, is the branch of natural science Natural science or empirical science is one of the branches of science concerned with the description, understanding and prediction of natural phenomena, based on empiri ...
. It was announced on 15 September 2006 in the journal
Science Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
that ''P. trichocarpa'' became the first tree species to have its entire
genome A genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding genes, other functional regions of the genome such as ...
sequenced In genetics and biochemistry, sequencing means to determine the primary structure (sometimes incorrectly called the primary sequence) of an unbranched biopolymer. Sequencing results in a symbolic linear depiction known as a sequence which succi ...
. In the mountains of interior western North America, it is replaced by the willow-leaved poplar or narrowleaf cottonwood, '' P. angustifolia''. ''P. trichocarpa'' is sometimes treated within ''P. balsamifera'' as ''P. balsamifera'' ssp. ''trichocarpa.'' Simon's poplar ('' P. simonii''), a native of northwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, is frequently planted as a shade tree in northern European cities. It is an attractive ornamental tree with whitish bark, and nearly rhombic, 6–10 cm long leaves which appear on the tree in early spring. Maximowicz' poplar or Japanese poplar ('' P. maximowiczii'') and the Ussuri poplar ('' P. ussuriensis'' = ''P. maximowiczii'' var. ''barbinervis'') are similar, occurring in northeastern China,
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...
, and eastern
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
; they have broader leaves. Another similar species is the laurel-leaf poplar ('' P. laurifolia'' from
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
), which differs from its relatives in narrower leaves shaped like a
bay laurel ''Laurus nobilis'' is an aromatic evergreen tree or large shrub with green, glabrous (smooth) leaves. It is in the flowering plant family Lauraceae. According to Flora Cretica (Kleinsteuber Books, 2024, ISBN 978-3-9818110-5-6) the stem can be 1 ...
leaf. Another putative member of this group is the Korean poplar ('' P. koreana''). Whether the northeast Asian Sichuan poplar ('' P. szechuanica'') and '' P. tristis'' belong here or with the
aspen Aspen is a common name for certain tree species in the Populus sect. Populus, of the ''Populus'' (poplar) genus. Species These species are called aspens: * ''Populus adenopoda'' – Chinese aspen (China, south of ''P. tremula'') * ''Populus da ...
s is not yet resolved. Likewise, the affiliation of '' P. cathayana'' and the Yunnan poplar ('' P. yunnanensis'') are in need of further study, though these may well be balsam poplars. There are other species not listed here. How to classify ''Populus'' into species, at least for Chinese populations, is not yet settled.


Use by humans

Balsam poplars are cultivated mainly in parks for their ornamental, light-coloured bark and pleasant scent in spring. Western balsam poplar is also planted as a
timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...
crop. The
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
is soft, very light in weight, but strong for its weight, coarse and fibrous, does not polish or plane easily, and is used for pallet boxes and other similar rough uses. Several hybrids between balsam poplars (particularly western balsam poplar) and the cottonwoods have also been developed for lumber production or for use as shade trees. These hybrids are selected for exceptionally fast growth and disease resistance. Poplars are also of potential use for
biofuel Biofuel is a fuel that is produced over a short time span from Biomass (energy), biomass, rather than by the very slow natural processes involved in the formation of fossil fuels such as oil. Biofuel can be produced from plants or from agricu ...
s because of their fast growth. Researchers are aiming to use genetic techniques to make poplars grow fatter, with a smaller canopy, so that more trees can be grown more quickly in a small space, and to make the plants contain a higher proportion of
cellulose Cellulose is an organic compound with the chemical formula, formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of glycosidic bond, β(1→4) linked glucose, D-glucose units. Cellulose is an important s ...
to
lignin Lignin is a class of complex organic polymers that form key structural materials in the support tissues of most plants. Lignins are particularly important in the formation of cell walls, especially in wood and bark, because they lend rigidit ...
. The increased cellulose content would make them easier to convert into sugars and ethanol for biofuel or for pulp in paper mills. The buds of various balsam poplars have long been combined with a lard base to make the vulnerary ointment
Balm of Gilead Balm of Gilead was a rare perfume used medicinally that was mentioned in the Hebrew Bible and named for the region of Gilead, where it was produced. The expression stems from William Tyndale's language in the King James Bible of 1611 and has ...
.Angier, Bradford. ''Field Guide to Medicinal Wild Plants''. Stackpole Books, 1978, p. 45.


See also

*
Balm of Gilead Balm of Gilead was a rare perfume used medicinally that was mentioned in the Hebrew Bible and named for the region of Gilead, where it was produced. The expression stems from William Tyndale's language in the King James Bible of 1611 and has ...
*
Protium heptaphyllum ''Protium heptaphyllum'', also known as almécega, breu, and almíscar, is a species of tree from South America commonly found in the areas of Brazil, the The Guianas, Guyanas, and Venezuela. The plant is typically harvested by locals for resin, f ...
, South-American tacamahaca.


References

{{Taxonbar, from=Q15700186 sect. Tacamahaca Medicinal plants of Asia Medicinal plants of North America Plant sections