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''Populus deltoides'', the eastern cottonwood or necklace poplar, is a cottonwood poplar native to North America, growing throughout the eastern, central, and southwestern
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as well as the southern Canadian prairies, the southernmost part of eastern
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, and northeastern
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.


Description

''Populus deltoides'' is a large
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to tall and with a trunk up to diameter, one of the largest North American
hardwood Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes fro ...
trees. The
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
is silvery-white, smooth or lightly fissured when young, becoming dark gray and deeply fissured on old trees. The twigs are grayish-yellow and stout, with large triangular leaf scars. The winter buds are slender, pointed, long, yellowish brown, and resinous. It is one of the fastest growing trees in North America. In Mississippi River bottoms, height growth of per year for a few years has been seen. Sustained height growth of height growth and diameter growth per year for 25 years is common. The leaves are large, deltoid (triangular), long and broad with a truncated (flattened) base and a petiole long. The leaf is very coarsely toothed, the teeth are curved and gland tipped, and the petiole is flat; they are dark green in the summer and turn yellow in the fall (but many cottonwoods in dry locations drop their leaves early from the combination of drought and leaf rust, making their fall color dull or absent). Due to the flat stem of the leaf, the leaf has the tendency to shake from even the slightest breeze. This is one of the identifying characteristics. It is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
, with the
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
s (
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind- pollinated ( anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arra ...
s) produced on single-sex trees in early spring. The male (pollen) catkins are reddish-purple and long; the female catkins are green, long at pollination, maturing long with several seed capsules (samaras) in early summer, which split open to release the numerous small
seeds A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...
attached to
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor p ...
-like strands. A single tree may release 40 million seeds a season.USGS Aquatic and Wetland Vascular Plants of the Northern Great Plains: A successful, simple, reproducible, high frequency micropropagation protocol has been described by Yadav Rakesh et al., 200
''Populus deltoides''
v-Plants (Chicago Herbarium)
''Populus deltoides''


Variation

The species is divided into three subspecies or up to five varieties. The subspecies classification is as follows: *''Populus deltoides'' subsp. ''deltoides'', eastern cottonwood is found in southeastern Canada (the south of
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central C ...
and
Quebec Quebec ( ; )According to the Government of Canada, Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is ...
) and the eastern United States (throughout, west to
North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the indigenous Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north and by the U.S. states of Minnesota to the east, S ...
to
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
). *''P. d. monilifera'' (Aiton) Eckenw., the plains cottonwood (syn. ''P. deltoides'' var. ''occidentalis'' Rydb.; ''P. sargentii'' Dode) ranges from southcentral Canada (southern
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Saskatchewan Saskatchewan ( ; ) is a province in western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on the south by the U.S. states of Montana and North ...
, and
Manitoba , image_map = Manitoba in Canada 2.svg , map_alt = Map showing Manitoba's location in the centre of Southern Canada , Label_map = yes , coordinates = , capital = Win ...
) to the central United States and south to northern
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
and
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
. It is the state tree of
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the sou ...
. *''P. d. wislizeni'' (S.Watson) Eckenw., the Rio Grande cottonwood (syn. ''P. wislizeni'' (S.Watson) Sarg.; ''P. fremontii'' var. ''wislizeni'' S.Watson) grows from southern
Colorado Colorado (, other variants) is a state in the Mountain states, Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the wes ...
south through
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
to northeastern Mexico (
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, San Luis Potosi), and west to
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a U.S. state, state in the Southwestern United States. It is the list of U.S. states and territories by area, 6th largest and the list of U.S. states and territories by population, 14 ...
(presence in
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
, listed by GRIN, is doubtful, not included in the Jepson ''Flora of California''Jepson Flora
''Populus''
clicking 'next taxon' through the genus shows no entry for this taxon)
). (Note: Some sources mistakenly spell the epithet "''wislizenii''". Correct spelling is with one "i", per ICN article 60C.2.)


Ecology

It needs bare soil and full sun for successful germination and establishment; in natural conditions, it usually grows near
river A river is a natural flowing watercourse, usually freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the ...
s, with mud banks left after
flood A flood is an overflow of water ( or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are an area of study of the discipline hydrol ...
s providing ideal conditions for seedling germination; human soil cultivation has allowed it to increase its range away from such habitats. Unlike related species such as quaking aspen, it does not propagate through clonal colonies, but will resprout readily when cut down. The leaves serve as food for
caterpillar Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
s of various Lepidoptera.


Uses

The wood of eastern cottonwood is typical of the ''Populus'' family in its softness, weighing just . It is utilized for things like plywood and interior parts of furniture. General Custer fed his horses and mules the bark during the 1868–69 winter campaign against Native American tribes south of Arkansas. According to
Charles Goodnight Charles Goodnight (March 5, 1836 – December 12, 1929), also known as Charlie Goodnight, was a rancher in the American West. In 1955, he was inducted into the Hall of Great Westerners of the National Cowboy & Western Heritage Museum. Early ...
, cowboys afflicted with gastric disorders would make an astringent tea from the inner bark.


Oldest and largest

Eastern cottonwoods typically live 70–100 years, but they have the potential to live 200–400 years in ideal conditions. The
Balmville Tree The Balmville Tree was an old-growth eastern cottonwood growing at the intersection of River Road, Balmville Road and Commonwealth Avenue in Balmville, New York, a hamlet within the Town of Newburgh. It was the oldest tree of that species in th ...
(felled in 2015 at approximately 316 years old) was the oldest eastern cottonwood in the United States. *The US national champion ''Populus deltoides'' var. ''deltoides'' is located in Beatrice, Nebraska and measures tall, wide. *The US national champion ''Populus deltoides'' var. ''monilifera'' is located in Ravalli County, Montana and measures tall, wide. *The US national champion ''Populus deltoides'' var. ''wislizeni'' is located in
Bernalillo County, New Mexico Bernalillo County () is the most populous county in the U.S. state of New Mexico.Bernalillo C ...
and measures tall, wide. The largest recorded cottonwood tree in the world is the Frimley Park tree located in Hastings, New Zealand and measures tall, wide and in girth. This cottonwood was planted in the 1870s.


Symbolism

Calling the cottonwood tree "the pioneer of the prairie", the Kansas state legislature designated the cottonwood the official state tree of
Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to ...
in 1937. It became the state tree of
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the sou ...
in 1947, and that of
Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the so ...
since 1972.


See also

* ''Populus'' section ''Aigeiros''


References


External links

*
Video footage of Eastern Cottonwood
{{Authority control deltoides Symbols of Kansas Symbols of Nebraska Symbols of Wyoming Trees of Alberta Trees of humid continental climate Trees of Mexico Trees of Ontario Trees of Saskatchewan Trees of Eastern Canada Trees of the Eastern United States Trees of the Great Lakes region (North America) Trees of the North-Central United States Trees of the Northeastern United States Trees of the Northwestern United States Trees of the Plains-Midwest (United States) Trees of the South-Central United States Trees of the Southeastern United States Trees of the Southwestern United States Trees of the United States