__NOTOC__ Portunoidea is a

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

crab Crabs are decapod crustaceans of the infraorder Brachyura (meaning "short tailed" in Greek language, Greek), which typically have a very short projecting tail-like abdomen#Arthropoda, abdomen, usually hidden entirely under the Thorax (arthropo ...

s that includes the family

Portunidae Portunidae is a family of crabs which contains the swimming crabs. Its members include well-known shoreline crabs such as the blue crab (''Callinectes sapidus'') and velvet crab ('' Necora puber''). Description Portunid crabs are characterised b ...

, the swimming crabs. Which other crab families are also placed here is a matter of some contention, and may be revised following

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

analyses.

Description

Their rather flat and smooth

carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...

is usually wider than long and of

hexagonal In geometry, a hexagon (from Greek , , meaning "six", and , , meaning "corner, angle") is a six-sided polygon. The total of the internal angles of any simple (non-self-intersecting) hexagon is 720°. Regular hexagon A regular hexagon is d ...

, subhexagonal,

rectangular In Euclidean plane geometry, a rectangle is a rectilinear convex polygon or a quadrilateral with four right angles. It can also be defined as: an equiangular quadrilateral, since equiangular means that all of its angles are equal (360°/4 = 90 ...

, or transversely

ovate Ovate may refer to: * Ovate (egg-shaped) leaves, tepals, or other botanical parts *Ovate, a type of prehistoric stone hand axe * Ovates, one of three ranks of membership in the Welsh Gorsedd * Vates or ovate, a term for ancient Celtic bards ...

shape. It is usually widest between the hindmost spines of the forward rim; there may be up to 9 pairs of these spines, with a few smaller ones right above the head, but they are missing altogether in some

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

. In some, the first

maxilliped An appendage (or outgrowth) is an external body part or natural prolongation that protrudes from an organism's body such as an arm or a leg. Protrusions from single-celled bacteria and archaea are known as cell-surface appendages or surface app ...

endopod The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, : ...

is lobed, forming the characteristic portunid lobe. The

cheliped A chela ()also called a claw, nipper, or pinceris a pincer-shaped organ at the end of certain limbs of some arthropods. The name comes from Ancient Greek , through Neo-Latin '. The plural form is chelae. Legs bearing a chela are called chelipeds ...

s are usually robust, and in some the last

pereiopod The anatomy of a decapod consists of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment – often called a somite – may possess one pair of appendages, although in various groups these m ...

pair has ovate dactyls. The sutures of the

sternum The sternum (: sternums or sterna) or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major bl ...

between segments 4 to 8 are usually incomplete, and in the

, the eighth

sternite The sternum (: sterna) is the ventral portion of a segment of an arthropod thorax or abdomen. In insects, the sterna are usually single, large sclerites, and external. However, they can sometimes be divided in two or more, in which case the su ...

is usually visible if seen from below and has a penial groove. In males, the

abdominal The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...

somites The somites (outdated term: primitive segments) are a set of bilaterally paired blocks of paraxial mesoderm that form in the embryonic stage of somitogenesis, along the head-to-tail axis in segmented animals. In vertebrates, somites subdivide in ...

are either all free or the third to fifth are fused, often retaining the sutures though. The first

gonopod Gonopods are specialized appendages of various arthropods used in reproduction or egg-laying. In males, they facilitate the transfer of sperm from male to female during mating, and thus are a type of intromittent organ. In crustaceans and millipe ...

is strongly curved, with a swollen and strongly hooked base.

Classification

Portunoidea are close relatives of the

Xanthoidea Xanthoidea is a Taxonomic rank, superfamily of crabs, comprising seven families. Formerly, a number of other families were included in Xanthoidea, but many of these have since been removed to other superfamilies. These include Carpilioidea, Eriph ...

, and the

families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...

Hexapodidae Hexapodidae is a family of crabs, the only family in the superfamily Hexapodoidea. It has traditionally been treated as a subfamily of the family Goneplacidae, and was originally described as a subfamily of Pinnotheridae. Its members can be d ...

and Mathildellidae, usually included there, are sometimes placed in the Portunoidea, while the deep-sea crabs ( Geryonidae) are usually placed in the Portunoidea but sometimes in the Xanthoidea. All Portunoidea live in the

ocean The ocean is the body of salt water that covers approximately 70.8% of Earth. The ocean is conventionally divided into large bodies of water, which are also referred to as ''oceans'' (the Pacific, Atlantic, Indian Ocean, Indian, Southern Ocean ...

, although the family Trichodactylidae, sometimes included here, live in

fresh water Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salt (chemistry), salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include ...

. There are thirteen families in the superfamily Portunoidea, four of which are extinct: It also contains the genus '' Garthopilumnus''.

References

External links

* {{Taxonbar, from=Q1337809 Crustacean superfamilies Taxa named by Constantine Samuel Rafinesque