Plectocomiopsis
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''Plectocomiopsis'' is a dioecious
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the
palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae ** List of Arecaceae genera **Palm oil * Several other plants known as "palm" Palm or Palms may also refer to: Music ...
family found in
Indochina Mainland Southeast Asia (historically known as Indochina and the Indochinese Peninsula) is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to th ...
,
Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
,
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
and
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
.Kew World Checklist of Selected Plant Families
/ref>
Hapaxanthic Monocarpic plants are those that flower and set seeds only once, and then die. The term is derived from Greek (', "single" + , "fruit" or "grain"), and was first used by Alphonse de Candolle. Other terms with the same meaning are ''hapaxanth'' a ...
and armed with spines, they are a climbing
rattan Rattan, also spelled ratan (from Malay language, Malay: ''rotan''), is the name for roughly 600 species of Old World climbing palms belonging to subfamily Calamoideae. The greatest diversity of rattan palm species and genera are in the clos ...
, closely related to the ''
Myrialepis ''Myrialepis'' is a monotypic genus of flowering plant in the palm family. The single species, ''Myrialepis paradoxa'', is native to Southeast Asia.Riffle, Robert L. and Craft, Paul (2003) ''An Encyclopedia of Cultivated Palms''. Portland: Timber ...
'' palms.Uhl, Natalie W. and Dransfield, John (1987) ''Genera Palmarum - A classification of palms based on the work of Harold E. Moore''. Lawrence, Kansas: Allen Press. / The name is
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
for "similar to ''
Plectocomia ''Plectocomia'' is a genus of flowering plant in the family Arecaceae native to China, the Himalayas, and Southeast Asia. Plants are dioecious, with male and female flowers produced on separate individuals. Species The genus contains the follow ...
''", another close relative.Riffle, Robert L. and Craft, Paul (2003) ''An Encyclopedia of Cultivated Palms''. Portland: Timber Press. /


Description

The stems are small, densely clustering, spiny, and high climbing with long internodes and conspicuous scars. Young
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are undivided or with few segments, in maturity becoming pinnate and cirrate with a tubular, unarmed or sparsely armed, scaly leaf sheath. Ocreas present, entire or becoming tattered. The petiole, when present, and the proximal end of the rachis are deeply channeled and spiny; the cirrus and distal end of the rachis are armed with regularly arranged reflexed climbing spines. The single fold, lanceolate leaflets may be few to numerous, usually with armed margins and caducous scales, with conspicuous midribs and transverse veinlets. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
is produced at the top of the stem amongst the most distal, often reduced leaves, axis adnate to the internode and emerging from the leaf sheath mouth. The peduncle is short, the prophyll is tubular and two-keeled, peduncular bracts usually absent, and the rachis is much longer than the peduncle. The rachis bracts are tubular and more or less distichous, each subtending a horizontal or pendulous first order branch which features basal, tubular bracts with triangular limbs carrying monopodial
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
clusters. In ''P. corneri'' the inflorescence is branched to three orders instead of two, with flowers borne on the axes of all orders. The male flowers are arranged in clusters of up to 32, each flower held in a cuplike rachilla bract with a two-keeled bracteole. The calyx is thick, leathery and tubular, with three lobes, and abaxially covered in scaly trichomes; the corolla is similar, with two distal splits forming three triangle shaped lobes, also bearing scales. The six stamens are laterally fused forming a tube which is tipped by six free, reflexed filaments with short, oblong anthers. The
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
is disulcate and elliptic with tectate exine; pistillode minute. Female flowers are usually borne in clusters of two to four but may be solitary, and, like the male's, are carried in cuplike
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
s with two-keeled
bracteole In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also look ...
s, with the occasional barely developed second bracteole. The
calyx CALYX, Inc. is a non-profit publisher of art and literature by women founded in 1976 based in Corvallis, Oregon. CALYX publishes both '' CALYX, A Journal of Art and Literature by Women'' twice a year and CALYX Books, which publishes one to three ...
is also thick and leathery, thrice lobed and scaly, usually persistent into fruit; corolla leathery, scaly, with three lobes. The staminodial ring bears six short lobes and sagging, empty
anther The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s;
gynoecium Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
egg-shaped to cylindrical, the apical stigmas becoming scaly nearing antithesis. Three incomplete
locule A locule (: locules) or loculus (; : loculi) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usually refers to a chamber within an ovary ...
s are present, each bearing one antropous, basally attached
ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...
. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
has one, rarely two
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s, covered in persistent
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...
whorls, and stigmatic apical remains. The
epicarp Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Fruitlike structures may develop directly from the seed itself rather than ...
is matted in irregular vertical rows of reflexed scales, with a thin
mesocarp Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Fruitlike structures may develop directly from the seed itself rather tha ...
and an undifferentiated
endocarp Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Fruitlike structures may develop directly from the seed itself rather th ...
. The
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
is basally attached, spherical, usually depressed, with a thick sarcoesta, a homogeneous endosperm and a basal
embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...
.


Distribution and habitat

''Plectocomiopsis triquetra'' and ''P. wrayi'' are low land
peat swamp forest Peat swamp forests are tropical and subtropical moist broadleaf forests, tropical moist forests where waterlogged soil prevents dead leaves and wood from fully decomposing. Over time, this creates a thick layer of acidic peat. Large areas of th ...
inhabitants, the former in Borneo, the latter in Malaysia. ''P. corneri'' and ''P. mira'' are Dipterocarp
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
dwellers to 700 m, and ''P. geminiflora'' is distributed through a variety of tropical forest types to 1200 m.


Cultivation and uses

The ''Plectocomiopsis'' species are not usually cultivated, being extremely spiny and requiring particular tropical conditions. In their natural range they are used in basketry, however their fragile nature prevents them from being a common rattan source. The palm heart of ''P. geminiflora'', despite its bitter taste, is regarded as a delicacy in
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
; villagers regard some of the others as edible and some as poisonous.


References


External links


''Plectocomiopsis'' on NPGS/GRINGBIF portalFairchild Guide to Palms: ''Plectocomiopsis''Naturekind.org images
* {{Taxonbar, from=Q144325 Calamoideae Arecaceae genera Taxa named by Odoardo Beccari Dioecious plants