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Family (, : ) is one of the eight major
hierarchical A hierarchy (from Greek: , from , 'president of sacred rites') is an arrangement of items (objects, names, values, categories, etc.) that are represented as being "above", "below", or "at the same level as" one another. Hierarchy is an importan ...
taxonomic rank In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of or ...
s in
Linnaean taxonomy Linnaean taxonomy can mean either of two related concepts: # The particular form of biological classification (taxonomy) set up by Carl Linnaeus, as set forth in his ''Systema Naturae'' (1735) and subsequent works. In the taxonomy of Linnaeus th ...
. It is classified between
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
and
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
. A family may be divided into
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zool ...
, which are intermediate ranks between the ranks of family and genus. The official family names are
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family
Juglandaceae The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have ...
, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active
taxonomist In biology, taxonomy () is the science, scientific study of naming, defining (Circumscription (taxonomy), circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxon, taxa (si ...
s. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community for extended periods. The continual publication of new data and diverse opinions plays a crucial role in facilitating adjustments and ultimately reaching a consensus over time.


Nomenclature

The naming of families is codified by various international bodies using the following suffixes: *In fungal, algal, and
botanical nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from taxonomy (biology), taxonomy. Plant taxonomy is concerned with grouping and classifying plants; Botany, botanical nomenclature then provides na ...
, the family names of plants, fungi, and algae end with the suffix " -aceae", except for a small number of historic but widely used names including
Compositae Asteraceae () is a large family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the order Asterales. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger fa ...
and
Gramineae Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivated i ...
. *In
zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
, the family names of animals end with the suffix " -idae".
/ref> Name changes at the family level are regulated by the codes of nomenclature. For botanical families, some traditional names like Palmae (
Arecaceae The Arecaceae () is a family (biology), family of perennial plant, perennial, flowering plants in the Monocotyledon, monocot order Arecales. Their growth form can be climbing palm, climbers, shrubs, tree-like and stemless plants, all commonly k ...
), Cruciferae (
Brassicaceae Brassicaceae () or (the older but equally valid) Cruciferae () is a medium-sized and economically important Family (biology), family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous pla ...
), and Leguminosae (
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
) are conserved alongside their standardized -aceae forms due to their historical significance and widespread use in the literature. Family names are typically formed from the stem of a
type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...
within the family. In zoology, when a valid family name is based on a genus that is later found to be a
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
, the family name may be maintained for stability if it was established before 1960. In botany, some family names that were found to be junior synonyms have been conserved due to their widespread use in the scientific literature. The family-group in zoological nomenclature includes several ranks: superfamily (-oidea), family (-idae),
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
(-inae), and
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
(-ini). Under the principle of coordination, a name established at any of these ranks can be moved to another rank while retaining its original authorship and date, requiring only a change in suffix to reflect its new rank. New family descriptions are relatively rare in taxonomy, occurring in fewer than one in a hundred taxonomic publications. Such descriptions typically result from either the discovery of organisms with unique combinations of characters that do not fit existing families, or from
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analyses that reveal the need for reclassification.


History

The taxonomic term ' was first used by French botanist
Pierre Magnol Pierre Magnol (8 June 1638 – 21 May 1715) was a French botanist. He was born in the city of Montpellier, where he lived and worked for most of his life. He became Professor of Botany and Director of the Royal Botanic Garden of Montpellier and h ...
in his ' (1689) where he called the seventy-six groups of plants he recognised in his tables families ('). The concept of rank at that time was not yet settled, and in the preface to the ' Magnol spoke of uniting his families into larger ', which is far from how the term is used today. In his work ''Philosophia Botanica'' published in 1751,
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
employed the term ''familia'' to categorize significant plant groups such as
trees In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only p ...
,
herb Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal purposes, or for fragrances. Culinary use typically distingu ...
s,
fern The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...
s, palms, and so on. Notably, he restricted the use of this term solely within the book's morphological section, where he delved into discussions regarding the vegetative and generative aspects of plants. Subsequently, in French botanical publications, from
Michel Adanson Michel Adanson (7 April 17273 August 1806) was an 18th-century French botanist and naturalist who traveled to Senegal to study flora and fauna. He proposed a "natural system" of taxonomy distinct from the binomial system forwarded by Linnaeus. ...
's ' (1763) and until the end of the 19th century, the word ' was used as a French equivalent of the Latin ' (or '). The family concept in botany was further developed by the French botanists
Antoine Laurent de Jussieu Antoine Laurent de Jussieu (; 12 April 1748 – 17 September 1836) was a French botanist, notable as the first to publish a natural classification of flowering plants; much of his system remains in use today. His classification was based on an e ...
and
Michel Adanson Michel Adanson (7 April 17273 August 1806) was an 18th-century French botanist and naturalist who traveled to Senegal to study flora and fauna. He proposed a "natural system" of taxonomy distinct from the binomial system forwarded by Linnaeus. ...
. Jussieu's 1789 ''Genera Plantarum'' divided plants into 100 'natural orders,' many of which correspond to modern plant families. However, the term 'family' did not become standardized in botanical usage until after the mid-nineteenth century. In
zoology Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...
, the family as a rank intermediate between order and genus was introduced by
Pierre André Latreille Pierre André Latreille (; 29 November 1762 – 6 February 1833) was a French zoology, zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained hi ...
in his ' (1796). He used families (some of them were not named) in some but not in all his orders of "insects" (which then included all
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s). The standardization of zoological family names began in the early nineteenth century. A significant development came in 1813 when William Kirby introduced the -idae suffix for animal family names, derived from the Greek 'eidos' meaning 'resemblance' or 'like'. The adoption of this naming convention helped establish families as an important taxonomic rank. By the mid-1800s, many of Linnaeus's broad genera were being elevated to family status to accommodate the rapidly growing number of newly discovered species. In nineteenth-century works such as the ' of
Augustin Pyramus de Candolle Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss people, Swiss botany, botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple ...
and the ' of
George Bentham George Bentham (22 September 1800 – 10 September 1884) was an English botanist, described by the weed botanist Duane Isely as "the premier systematic botanist of the nineteenth century". Born into a distinguished family, he initially studie ...
and
Joseph Dalton Hooker Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For 20 years he served as director of the Ro ...
this word ' was used for what now is given the rank of family.


Uses

Families serve as valuable units for evolutionary, paleontological, and genetic studies due to their relatively greater stability compared to lower taxonomic levels like genera and species. Families play a significant practical role in biological education and research. They provide an efficient framework for teaching taxonomy, as they group organisms with general similarities while remaining specific enough to be useful for identification purposes. For example, in botany, learning the characteristics of major plant families helps students identify related species across different geographic regions, since families often have worldwide distribution patterns. In many groups of organisms, families serve as the primary level for taxonomic identification keys, making them particularly valuable for
field guide A field guide is a book designed to help the reader identify wildlife (flora or fauna or funga) or other objects of natural occurrence (e.g. rocks and minerals). It is generally designed to be brought into the " field" or local area where suc ...
s and systematic work as they often represent readily recognizable groups of related organisms with shared characteristics. In ecological and
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
research, families frequently serve as the foundational level for identification in survey work and environmental studies. This is particularly useful because families often share life history traits or occupy similar
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of Resource (biology), resources an ...
s. Some families show strong correlations between their taxonomic grouping and ecological functions, though this relationship varies among different groups of organisms. The stability of family names has practical importance for applied biological work, though this stability faces ongoing challenges from new scientific findings. Modern
molecular A molecule is a group of two or more atoms that are held together by attractive forces known as chemical bonds; depending on context, the term may or may not include ions that satisfy this criterion. In quantum physics, organic chemistry, ...
studies and
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analyses continue to refine the understanding of family relationships, sometimes leading to reclassification. The impact of these changes varies among different groups of organisms – while some families remain well-defined and easily recognizable, others require revision as new evidence emerges about evolutionary relationships. This balance between maintaining nomenclatural stability and incorporating new scientific discoveries remains an active area of taxonomic practice.


See also

*
Systematics Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...
, the study of the diversity of living organisms *
Cladistics Cladistics ( ; from Ancient Greek 'branch') is an approach to Taxonomy (biology), biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesiz ...
, the classification of organisms by their order of branching in an evolutionary tree *
Virus classification Virus classification is the process of naming viruses and placing them into a taxonomy (biology), taxonomic system similar to the classification systems used for cell (biology), cellular organisms. Viruses are classified by phenotypic characteri ...
*
List of Anuran families This list of Anuran families shows all extant family (biology), families of Frog, Anura. Anura is an order of animals in the class Amphibia that includes frogs and toads. More than 5,000 species are described in the order. The living anurans are t ...
* List of Testudines families *
List of fish families This is a list of fish families sorted alphabetically by scientific name. There are 525 families in the list. __NOTOC__ A - B - C - D - E - F - G - H - I - J - K - L - M - N - O - P - R - S - T - U - V - W - X - Y - Z - ...
*
List of families of spiders Spider taxonomy is the part of taxonomy that is concerned with the science of naming, defining and classifying all spiders, members of the Araneae order of the arthropod class Arachnida, which has more than 52,700 described species. However, there ...


References


Bibliography

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External links

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