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The Miridae are a large and diverse insect family at one time known by the taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs".
Common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contra ...
s include plant bugs, leaf bugs, and grass bugs. It is the largest family of true bugs belonging to the suborder Heteroptera; it includes over 10,000 known species, and new ones are being described constantly. Most widely known mirids are species that are notorious agricultural pests that pierce plant tissues, feed on the sap, and sometimes transmit viral plant diseases. Some species however, are predatory.


Description

Miridae are small, terrestrial insects, usually oval-shaped or elongate and measuring less than in length. Many of them have a hunched look, because of the shape of the prothorax, which carries the head bent down. Some are brightly coloured and attractively patterned, others drab or dark, most being inconspicuous. Some genera are ant mimics at certain stages of life. The Miridae do not have any ocelli. Their rostrum has four segments. One useful feature in identifying members of the family is the presence of a cuneus; it is the triangular tip of the corium, the firm, horny part of the forewing, the hemelytron. The cuneus is visible in nearly all Miridae, and only in a few other Hemiptera, notably the family Anthocoridae, which are not much like the Miridae in other ways. The tarsi almost always have three segments.


Some mirid species

* Lygus bugs (''Lygus'' spp.), including the tarnished and western tarnished plant bugs, are serious pests in the
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor p ...
, strawberry, and alfalfa industries. * ''
Stenotus binotatus ''Stenotus binotatus'' is a species of plant bug, originally from Europe, but now also established across North America and New Zealand. It is long, yellowish, with darker markings on the pronotum and forewings. It feeds on various grasses, an ...
'', a minor pest of cereal crops, especially
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeolog ...
* Apple dimpling bug ('' Campylomma liebknechti'') damages
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus '' Malus''. The tree originated in Central Asia, where its wild ances ...
blossoms and small growing fruits. *
Mosquito bug The genus ''Helopeltis'', also known as mosquito bugs, is a group of Heteropterans in the family Miridae (capsid bugs) and tribe Dicyphini. They include important pests of various crops, including cacao, cashew, cotton and tea. Now in a different ...
s ''
Helopeltis The genus ''Helopeltis'', also known as mosquito bugs, is a group of Heteropterans in the family Miridae (capsid bugs) and tribe Dicyphini. They include important pests of various crops, including cacao, cashew, cotton and tea. Now in a different ...
'' and ''
Afropeltis The genus ''Helopeltis'', also known as mosquito bugs, is a group of Heteropterans in the family Miridae (capsid bugs) and tribe Dicyphini. They include important pests of various crops, including cacao, cashew, cotton and tea. Now in a different ...
'' spp. – that infest various crops including tea,
cacao Cacao is the seed from which cocoa and chocolate are made, from Spanish cacao, an adaptation of Nahuatl cacaua, the root form of cacahuatl ("bean of the cocoa-tree"). It may also refer to: Plants *''Theobroma cacao'', a tropical evergreen tree ** ...
and
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor p ...
*
Honeylocust plant bug ''Blepharidopterus chlorionis'', known commonly as the honeylocust plant bug and under the synonym ''Diaphnocoris chlorionis'', is a hemipteran in the family Miridae. They are a major pest of the honey locust tree (''Gleditsia triacanthos''), a ...
(''Diaphnocoris chlorionis'') damages foliage on honeylocust trees. * Green mirid (''Creontiades dilutus'') damages many types of field crops. *
Potato capsid ''Closterotomus norvegicus'' (also known as the potato capsid) is a species of bugs belonging to the family Miridae, subfamily Mirinae. It can be found feeding on nettle, clover, and cannabis, as well as Compositae, potatoes, carrots and ...
(''Closterotomus norwegicus'') is a noted pest of
potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Un ...
and
clover Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus h ...
plants in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 List of islands of New Zealand, smaller islands. It is the ...
. * ''
Deraeocoris nebulosus ''Deraeocoris nebulosus'' is a species of plant bug in the family Miridae. It is found in North America. References Further reading * External links * Articles created by Qbugbot Insects described in 1872 Deraeocorini Hemipter ...
'' prefers other insects to plants in its diet, and has been used as a biocontrol agent against
mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...
s and scale insects. * '' Dicyphus hesperus'' sucks sap from various plants and preys on whitefly and red spider mites and can be used in biological pest control.


Systematics

This family includes a large number of
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
, many of which are still unknown, distributed in more than 1300 genera. The taxonomic tree includes the following subfamilies and numerous tribes: * Bryocorinae **
Bryocorini ''Bryocorinae'' is a subfamily of bugs in the family Miridae. Tribes and genera ''BioLib''BioLib.cz
Bryocorinae Baerenspr ...
** Dicyphini **
Eccritotarsini Eccritotarsini is a tribe of plant bugs in the family Miridae. There are about 14 genera and at least 40 described species in Eccritotarsini. Genera These 14 genera belong to the tribe Eccritotarsini: * '' Aguayomiris'' Maldonado Caprilles, 1 ...
*
Cylapinae Cylapinae is a subfamily of plant bug. Bugs in this group tend to forage actively on fungus covered rotten logs in humid tropical forests. Overview of genera * '' Adcylapocoris'' * '' Afrobothriomiris'' * '' Afrofulvius'' * '' Afrovannius'' * ...
** Cylapini ** Fulviini *
Deraeocorinae Deraeocorinae is a subfamily of plant bugs in the family Miridae. There are more than 40 genera and around 500 described species in Deraeocorinae. Genera These 41 genera belong to the subfamily Deraeocorinae: * '' Acutifromiris'' Hernandez & S ...
** Clivinematini **
Deraeocorini Deraeocorini is a tribe of plant bugs in the family Miridae. There are at least 19 genera and more than 340 described species in Deraeocorini. Genera These 19 genera belong to the tribe Deraeocorini: * '' Acutifromiris'' Hernandez & Stonedahl ...
** Hyaliodini ** Saturniomirini ** Surinamellini ** Termatophylini * Isometopinae ** Diphlebini ** Isometopini * Mirinae ** Herdoniini ** Hyalopeplini ** Mirini ** Pithanini ** Restheniini **
Stenodemini Stenodemini is a tribe of plant bugs in the family Miridae. There are more than 60 described species in Stenodemini. Genera ''BioLib'' includes: # '' Acetropis'' Fieber, 1858 # '' Acomocera'' Eyles, 1975 # ''Actinocoris'' Reuter, 1878 # '' Ast ...
*
Orthotylinae Orthotylinae is a subfamily of plant bugs in the family Miridae. There are at least 650 described species and at least 70 genera in Orthotylinae. The Orthotylinae have traditionally been diagnosed by their possession of lamellate, apically diverg ...
**
Ceratocapsini Ceratocapsini is a tribe of plant bugs in the family Miridae. There are about 7 genera and at least 80 described species in Ceratocapsini. Genera These seven genera belong to the tribe Ceratocapsini: * '' Ceratocapsus'' Reuter, 1876 * '' Pamill ...
**
Halticini Halticini is a tribe of plant bugs in the family Miridae. Genera ''BioLib'' includes the following: # '' Acratheus'' Distant, 1910 # '' Anapomella'' V.G. Putshkov, 1961 # '' Anapus'' Stål, 1858 # '' Barbarosia'' Kiyak, 1995 # '' Chorosomella'' ...
**
Orthotylini Orthotylini is a tribe of plant bugs in the family Miridae. There are more than 230 described genera in Orthotylini. Selected genera The following genera belong to the tribe Orthotylini: * '' Acaciacoris'' Schaffner, 1977 * '' Aoplonema'' Knigh ...
*
Phylinae Phylinae is a subfamily of the plant bug family Miridae. Species of this family are found worldwide. In research published in 2013, the subfamily Phylinae was reorganized. The tribe Auricillocorini is now considered a synonym of Hallodapini, an ...
**
Hallodapini Hallodapini is a tribe of plant bugs in the family Miridae. There are more than 50 genera in Hallodapini. Most species of this tribe are found in the old world, including Australia. Only one genus, Cyrtopeltocoris, is found in North America. G ...
**
Leucophoropterini Leucophoropterini is a tribe of plant bugs in the family Miridae. There are more than 20 genera and over 100 species in Leucophoropterini. Most species of this tribe are found in eastern Asia and Australia, with a single genus, ''Tuxedo'', found ...
** Phylini **
Pilophorini Pilophorini is a tribe (biology), tribe of Miridae, plant bug. The type genus is ''Pilophorus'' (Hahn, 1826). Schuh's analysis indicates that the Pilophorini originated in tropical Gondwanaland and subsequently spread into the temperate Northern ...


Psallopinae

Auth.: Schuh, 1976 # '' Isometopsallops'' Herczek & Popov, 1992 # '' Psallops'' Usinger, 1946 # †'' Cylapopsallops'' Popov & Herczek, 2006 # †'' Epigonopsallops'' Herczek & Popov, 2009


Genera ''

Incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''

''BioLib'' includes: # '' Amulacoris'' Carvalho & China, 1959 # '' Anniessa'' Kirkaldy, 1903 # '' Auchus'' Distant, 1893 # '' Bahiarmiris'' Carvalho, 1977 # '' Brasiliocarnus'' Kerzhner & Schuh, 1995 # '' Carmelinus'' Carvalho & Gomes, 1972 # '' Carmelus'' Drake & Harris, 1932 # '' Chaetophylidea'' Knight, 1968 # '' Charitides'' Kerzhner, 1962 # '' Colimacoris'' Schaffner & Carvalho, 1985 # '' Cylapocerus'' Carvalho & Fontes, 1968 # '' Duckecylapus'' Carvalho, 1982 # '' Englemania'' Carvalho, 1985 # '' Eurycipitia'' Reuter, 1905 # '' Faliscomiris'' Kerzhner & Schuh, 1998 # '' Fuscus'' Distant, 1884 # '' Guerrerocoris'' Carvalho & China, 1959 # ''Gunhadya'' - monotypic '' Gunhadya rubrofasciata'' Distant, 1920 # '' Heterocoris'' Guérin-Ménéville in Sagra, 1857 # '' Knightocoris'' Carvalho & China, 1951 # '' Leonomiris'' Kerzhner & Schuh, 1998 # '' Macrotyloides'' Van Duzee, 1916 # '' Merinocapsus'' Knight, 1968 # '' Mircarvalhoia'' Kerzhner & Schuh, 1998 # '' Montagneria'' Akingbohungbe, 1978 # '' Muirmiris'' Carvalho, 1983 # '' Myochroocoris'' Reuter, 1909 # '' Nesosylphas'' Kirkaldy, 1908 # '' Notolobus'' Reuter, 1908 # '' Nymannus'' Distant, 1904 # '' Paracoriscus'' Kerzhner & Schuh, 1998 # '' Paraguayna'' Carvalho, 1986 # '' Prodomopsis'' TBD # ''
Prodomus A portico is a porch leading to the entrance of a building, or extended as a colonnade, with a roof structure over a walkway, supported by columns or enclosed by walls. This idea was widely used in ancient Greece and has influenced many cult ...
'' TBD # '' Pseudobryocoris'' Distant, 1884 # '' Pygophorisca'' Carvalho & Wallerstein, 1978 # '' Rayeria'' TBD # '' Rewafulvia'' Carvalho, 1972 # '' Rhynacloa'' Reuter # '' Rondonisca'' Carvalho & Costa, 1994 # '' Rondonoides'' Carvalho & Costa, 1994 # '' Rondonotylus'' Carvalho & Costa, 1994 # '' Spanogonicus'' Berg # '' Sthenaridia'' TBD # ''
Zoilus Zoilus ( el, Ζωΐλος ''Zoilos''; c. 400320 BC) was a Greek grammarian, Cynic philosopher, and literary critic from Amphipolis in Eastern Macedonia, then known as Thrace. He took the name Homeromastix (Ὁμηρομάστιξ "Homer whippe ...
'' Distant, 1884


References


Further reading

* *
Google books preview


External links


PBI Plant Bug Index

Link to Al Wheeler's book on the biology of mirids

Research on mirids of Southern California at UC Riverside


on the
University of Florida The University of Florida (Florida or UF) is a public land-grant research university in Gainesville, Florida. It is a senior member of the State University System of Florida, traces its origins to 1853, and has operated continuously on its ...
/ Institute of Food and Agricultural Sciences ''Featured Creatures'' website {{Taxonbar, from=Q864705 Heteroptera families Articles containing video clips Taxa named by Carl Wilhelm Hahn