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The pin-tailed sandgrouse (''Pterocles alchata'') is a medium large
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the
sandgrouse Sandgrouse is the common name for Pteroclidae (), a family (biology), family of sixteen species of bird, members of the order Pterocliformes (). They are traditionally placed in two Genus, genera. The two central Asian species are classified as ...
family. It has a small, pigeon-like head and neck and a sturdy, compact body. It has long pointed wings, which are white underneath, a long tail and a fast direct flight. Flocks fly to watering holes at dawn. The call is a loud ''kattar-kattar''. This gregarious species breeds on dry open treeless plains and similar habitats. Its nest is a ground scrape into which two or three cream-coloured eggs with cryptic markings are laid. Both sexes incubate the eggs. The pin-tailed sandgrouse is about long. Its head and upperparts are yellowish-green. The underparts are white with a chestnut breast band separating the belly from the green neck. Sexes are somewhat similar, but the female is better camouflaged and has a shorter tail than the male. There are two subspecies; ''P. a. alchata'' breeds in southern Europe and ''P. a. caudacutus'' breeds in northwestern Africa, the Middle East and southeastern Asia. It is a partial migrant, with some
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
n birds moving to the
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
and northern
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
in winter. Males of the eastern race have duller underparts than the
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
an birds, and the females have white, rather than yellow, wing coverts.


Taxonomy

There are two subspecies. The nominate race, ''P. a. alchata'', breeds in
Iberia The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, compri ...
and southern
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
, and the eastern form ''P. a. caudacutus'' (
Gmelin Gmelin may refer to: * Karl Christian Gmelin, Carl Christian Gmelin (1762–1837), German botanist, author of ''Flora Badensis, Alsatica et confinium regionum cis- et transrhenania'' (1806) * Charles Gmelin (1872–1950), British Olympic athlete * ...
, 1774) is found in northwest
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, and from southeast
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
east to
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...
. The species name ''alchata'' is thought to be derived from the Arabic name ''al-gata القطا'' which is derived from the call of the bird.


Description

The pin-tailed sandgrouse is a robust, medium-sized bird about in length. The general colouring is cryptic, a blend of barred and flecked olive green, brown, buff, yellow, grey and black. The underparts and the feathered legs are dull white. left, Pin-tailed sandgrouse The sexes are different and the male's plumage varies with the time of year. In the breeding season, the male has the crown, most of the neck, the back and under-wing coverts a yellowish-green colour with dull yellow spots in the shoulder region. The cheeks are also yellow with a narrow black line extending from the beak, through the eye to the nape of the neck. The irises are brown and the beak is slatey grey. There is a black patch on the throat immediately below the beak and below this, there is a broad, reddish-brown band round the breast, bordered by a thin black stripe above and below the band. The outer wing coverts are chestnut edged with black and white and the primaries are black with pale edges which gives both the leading and trailing edges of the wing the appearance of a black rim in flight. The rump and the tail are distinctly barred in black and brownish-yellow and the streamers on the central tail feathers are slate-grey. Outside the breeding season, all the upper parts, including the crown and cheeks, are barred in black and brownish-yellow and the throat loses its black patch, becoming whitish. The female is generally similar to the male but the colours are duller. The cheeks and neck are golden-buff and lack the greenish tinge of the male. There is a black stripe running through the eye. The chin is white and there is an additional yellow-buff band across the breast with a broad black stripe above, another thinner one about a third of the way down and a further narrow black stripe at the base. The back and wings are grey, barred with black. The rump and the tail (which has shorter central feathers than the male) are similar in colouration to the male but have finer black barrings. In flight, the pin-tailed sandgrouse can be identified by its bright, white underparts and under-wing coverts, and the long feathers in the centre part of its tail. It is usually silent when on the ground but in the air communicates with other birds with a frequently uttered, loud "kattar-kattar", a nasal "ga-ga-ga" and a low-pitched "gang gang".


Distribution and habitat

The pin-tailed sandgrouse breeds in North Africa and the Middle East, Turkey, Iran, Iraq and Kazakhstan. In Europe it breeds in Spain, Portugal and the southern part of France. Eastern populations, particularly those from Kazakhstan, migrate to Pakistan and parts of northern India in winter. The pin-tailed sandgrouse inhabits open areas of stony land, semi arid areas at the edge of deserts, treeless plains and occasionally dried-out mud flats. In winter it may visit ploughed or fallow land but prefers sandy soils and is much less reliant on vegetation cover than the black-bellied sandgrouse (''Pterocles orientalis'') which has a similar range. It does not occur at elevations above about .


Behaviour

The pin-tailed sandgrouse often feeds in groups and gathers regularly in large numbers at waterholes, to which the birds fly soon after dawn. During the day they disperse to forage for seeds, which are the main part of their diet, and also eat buds, green shoots and leaves. They favour leguminous seeds and also eat seeds of knotweed (''Polygonum''),
buckwheat Buckwheat (''Fagopyrum esculentum'') or common buckwheat is a flowering plant in the knotweed family Polygonaceae cultivated for its grain-like seeds and as a cover crop. Buckwheat originated around the 6th millennium BCE in the region of what ...
(''Fagopyrum''), ''
Salicornia ''Salicornia'' is a genus of succulent, halophytic (salt tolerant) flowering plants in the family Amaranthaceae that grow in salt marshes, on beaches, and among mangroves. ''Salicornia'' species are native to North America, Europe, central Asia, ...
'', '' Artemisia'', camelthorn (''Alhagi''),
rock rose Rock rose, rock-rose, and rockrose are common names of various plants, including: * Cistaceae * ''Cistus'' * ''Halimium'' * ''Helianthemum'' * ''Pavonia lasiopetala'' * ''Phemeranthus'' * ''Portulaca grandiflora'' {{Plant common name ...
(''Helianthemum'') and asphodel (''Asphodelus''). In cultivated areas they take grain and the seeds of leguminous crops. Beetles and other small invertebrates are sometimes eaten and grit is consumed to help grind up the contents of the crop.


Breeding

The pin-tailed sandgrouse nests in a slight depression on the ground in the open. Two or three eggs are laid at intervals of two days, creamy-brown spotted with darker brown, reddish-brown and grey. Both parents share the task of incubation which lasts from nineteen to twenty five days. The male starts his shift about an hour before sunset and the female takes over after she has been to the waterhole in the morning. The young are
precocial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
and leave the nest soon after they have hatched. Both parents care for them but only the male is involved in bringing them water, absorbed by the feathers on his breast. The chicks are able to feed themselves by the age of a week and can fly by the time they are four weeks old. They are dependent on their parents for two months and attain their adult plumage at about four months. There is normally a single brood each year but if the eggs are destroyed or removed, more eggs may be laid.


Status

The pin-tailed sandgrouse is rated as being of "Least Concern" by
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
. This is because it has a large range and the population, estimated to be somewhere between 130,000 and 1,500,000 individuals, is believed to be stable. The range may be expanding in south eastern Turkey, possibly as a result of changes in climate. However, the populations in parts of Europe are decreasing due to changes in agricultural production. The bird remains common in Spain but in the only remaining locality in southern France where it is found, at La Crau, only about 170 pairs remained in 2002, while there were fewer than 100 pairs of birds breeding in Portugal in the same year.


References

{{Taxonbar, from=Q753741 pin-tailed sandgrouse Birds of North Africa Birds of Central Asia Birds of West Asia pin-tailed sandgrouse pin-tailed sandgrouse