Pilina Unguis
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''Pilina unguis'' is an extinct species of Paleozoic
Silurian The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of t ...
monoplacophora Shell of Monoplacophora Monoplacophora , meaning "bearing one plate", is a polyphyletic class of molluscs with a cap-like shell, inhabiting deep sea environments. Extant representatives were not recognized as such until 1952; previously they wer ...
n. It was first named as ''Tryblidium unguis'' and described by Gustaf Lindström in
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
from the Silurian deposits of
Gotland Gotland (; ; ''Gutland'' in Gutnish), also historically spelled Gottland or Gothland (), is Sweden's largest island. It is also a Provinces of Sweden, province/Counties of Sweden, county (Swedish län), Municipalities of Sweden, municipality, a ...
in Sweden, in 1880.


Shell description

The length of the shell is 59–67 mm, width 47–51 mm and the height of the shell is 15–16 mm. With its shell size up to 67 mm was at least twice larger than any recent known living monoplacophoran species.Moore J. & Raith Overhill R. 2006. ''An introduction to the invertebrates'', Second Edition. Cambridge, Cambridge University. Press, 319 pp.
Page 126
The
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses Science Biology * Seashell, a hard outer layer of a marine ani ...
has an obovate outline, anteriorly acuminate, posteriorly expanded, with the greatest width somewhat behind the median, transverse axis of the shell. It is regularly and moderately convex, being the most elevated near the median transverse axis. The
apex The apex is the highest point of something. The word may also refer to: Arts and media Fictional entities * Apex (comics) A-Bomb Abomination Absorbing Man Abraxas Abyss Abyss is the name of two characters appearing in Ameri ...
is close to the anterior margin and only very little prominent. The surface of the shell is covered with thread-fine ornamental lines parallel to the regular concentric lines of growth, somewhat interrupted by deeper sulci. It is generally even, excepting some wavy, irregular, shallow, longitudinal furrows. The
aperture In optics, the aperture of an optical system (including a system consisting of a single lens) is the hole or opening that primarily limits light propagated through the system. More specifically, the entrance pupil as the front side image o ...
is oval with comparatively thin margins, somewhat reflexed outwards. When seen from the side it forms a moderately elevated arch, being highest near the median line of the shell. The central space of the interior surface is surrounded by an oval ring of six pairs of
muscular MUSCULAR (DS-200B), located in the United Kingdom, is the name of a surveillance program jointly operated by Britain's Government Communications Headquarters (GCHQ) and the U.S. National Security Agency (NSA) that was revealed by documents release ...
scars, open anteriorly. The posterior ones are narrow and elongate, the middle ones are transversally broad and much enlarged towards the margin of the shell. The surface of these pairs is smooth or only partially scrobiculated. The uppermost pair is more complicated and consists, as in the kindred species, of a more shallow, elongated interior portion, and of a larger exterior one, from the inner corner of which there projects a narrow sinuous groove, directed obliquely upwards and leaving a small, smooth space between itself and the opposite similar one. The lower, interior part of this upper, muscular scar is pearshaped, wide below, with a narrow, stalklike neck upwards and its surface is finely reticulated by shallow pits and intervening ridges. Some dark, narrow streaks are directed from the enclosed central space of the shell towards the interstices between the muscular scars. The whole central part of the shell inside the scars and to their outer edges is of dark color, while the outer border is lighter. Near the apex there is on the inside a little oval depression or pit, which quite resembles a scar, filled up, as if there had been a foramen. Lindström considered this as the mark of the outlines of the initial shell, where now the apex is seen on the outside. The shell is very thin, scarcely exceeding 0.5 mm in thickness and composed of thin, glossy lamellas, which are not perforated by any parasite.


Distribution

This species has nearly the same geographical distribution as '' Tryblidium reticulatum''. It has been found in several specimens in
Fårö Fårö () or in Gutnish is a Baltic Sea island just north of the island of Gotland, itself off mainland Sweden's southeastern coast. It is the second-largest island in the county and it is a popular summer resort. It has its own language, Fårö ...
at Lauterhorn, Lansa and
Norsholm Norsholm () is a Urban areas in Sweden, locality situated in Norrköping Municipality, Östergötland County, Sweden with 615 inhabitants in 2010. It lies around 15  kilometres southwest of Norrköping. Norsholm lies on the European route E4, ...
, in Svarvare Huk, the canal near Westöös Änge in Hall,
Väskinde Väskinde () is a urban areas in Sweden, locality on the Swedish island of Gotland. Väskinde is also the name of the larger populated area, ''socken'' (not to be confused with Parishes of the Church of Sweden, parish). It comprises the same area ...
, Häftingsklint,
Lummelunda Lummelunda (sometimes referred to as Lummelunda and Etebols) is a populated area, a socken (not to be confused with Parishes of the Church of Sweden, parish), on the Swedish island of Gotland. It comprises the same area as the Registration district ...
, Stora Vede in Follingbo, the limestone cliffs near
Visby Visby () is an urban areas in Sweden, urban area in Sweden and the seat of Gotland Municipality in Gotland County on the island of Gotland with 24,330 inhabitants . Visby is also the episcopal see for the Diocese of Visby. The Hanseatic League, ...
and Kyrkberget in Visby. It has only been found in the strata ''b'' and ''c'' or the upper and lower
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
and never in the shale of ''a''. This is rather unexpected as the same species or at least a nearly allied variety, as mentioned above, has been found in the Lower Silurian strata of
Estonia Estonia, officially the Republic of Estonia, is a country in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the Baltic Sea across from Sweden, to the south by Latvia, and to the east by Ru ...
at
Borkholm Porkuni () is a village in Tapa Parish, Lääne-Viru County, in northern Estonia. The settlement is located around the Lake Porkuni, which is the source of the Valgejõgi River. In 1944, the Battle of Porkuni was fought in the area. Porkuni cas ...
and also in the Upper Silurian at Koik, in the »Jördensche Schicht». These are only a little more elongate and not so enlarged as the specimens from Gotland.


References

This article incorporates public domain text from reference.Lindström G. 1884
''On the Silurian Gastropoda and Pteropoda of Gotland''
Stockholm, 250 pp. Page
56-57Plate 1
figure 33-37
plate 19
figure 2.


External links


drawing
{{Taxonbar, from=Q7194032 Prehistoric monoplacophorans