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Pierre François Verhulst (28 October 1804, in
Brussels Brussels, officially the Brussels-Capital Region, (All text and all but one graphic show the English name as Brussels-Capital Region.) is a Communities, regions and language areas of Belgium#Regions, region of Belgium comprising #Municipalit ...
– 15 February 1849, in Brussels) was a Belgian
mathematician A mathematician is someone who uses an extensive knowledge of mathematics in their work, typically to solve mathematical problems. Mathematicians are concerned with numbers, data, quantity, mathematical structure, structure, space, Mathematica ...
and a doctor in
number theory Number theory is a branch of pure mathematics devoted primarily to the study of the integers and arithmetic functions. Number theorists study prime numbers as well as the properties of mathematical objects constructed from integers (for example ...
from the University of Ghent in 1825. He is best known for the logistic growth model.


Logistic equation

Verhulst developed the logistic function in a series of three papers between 1838 and 1847, based on research on modeling
population growth Population growth is the increase in the number of people in a population or dispersed group. The World population, global population has grown from 1 billion in 1800 to 8.2 billion in 2025. Actual global human population growth amounts to aroun ...
that he conducted in the mid 1830s, under the guidance of
Adolphe Quetelet Lambert Adolphe Jacques Quetelet FRSF or FRSE (; 22 February 1796 – 17 February 1874) was a Belgian- French astronomer, mathematician, statistician and sociologist who founded and directed the Brussels Observatory and was influential ...
; see for details. Verhulst published in the equation: : \frac = rN - \alpha N^2 where ''N''(''t'') represents number of individuals at time ''t'', ''r'' the intrinsic growth rate, and ''\alpha'' is the density-dependent crowding effect (also known as intraspecific competition). In this equation, the population equilibrium (sometimes referred to as the carrying capacity, ''K''), N^*, is : N^* = \frac . In he named the solution the
logistic curve A logistic function or logistic curve is a common S-shaped curve (sigmoid function, sigmoid curve) with the equation f(x) = \frac where The logistic function has domain the real numbers, the limit as x \to -\infty is 0, and the limit as x \ ...
. Later, Raymond Pearl and Lowell Reed popularized the equation, but with a presumed equilibrium, ''K'', as : \frac = r N \left(1 - \frac \right) where ''K'' sometimes represents the maximum number of individuals that the environment can support. In relation to the density-dependent crowding effect, \alpha = \frac. The Pearl-Reed logistic equation can be integrated exactly, and has solution : N(t) = \frac where ''C'' = 1/''N''(0) − 1/''K'' is determined by the initial condition ''N''(0). The solution can also be written as a weighted
harmonic mean In mathematics, the harmonic mean is a kind of average, one of the Pythagorean means. It is the most appropriate average for ratios and rate (mathematics), rates such as speeds, and is normally only used for positive arguments. The harmonic mean ...
of the initial condition and the carrying capacity, : \frac = \frac+ \frac. Although the continuous-time logistic equation is often compared to the logistic map because of similarity of form, it is actually more closely related to the Beverton–Holt model of fisheries recruitment. The concept of
R/K selection theory In ecology, selection theory relates to the selection of combinations of traits in an organism that trade off between quantity and quality of offspring. The focus on either an increased quantity of offspring at the expense of reduced individua ...
derives its name from the competing dynamics of
exponential growth Exponential growth occurs when a quantity grows as an exponential function of time. The quantity grows at a rate directly proportional to its present size. For example, when it is 3 times as big as it is now, it will be growing 3 times as fast ...
and carrying capacity introduced by the equations above.


See also

* Population dynamics * Logistic map * Logistic distribution


Works

* * * *


References

* ** Published as:


External links

* {{DEFAULTSORT:Verhulst, Pierre Francois 1804 births 1849 deaths Belgian mathematicians 19th-century Belgian male writers