Pied Starling
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The pied starling or African pied starling (''Lamprotornis bicolor'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
endemic to
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
,
Lesotho Lesotho, formally the Kingdom of Lesotho and formerly known as Basutoland, is a landlocked country in Southern Africa. Entirely surrounded by South Africa, it is the largest of only three sovereign enclave and exclave, enclaves in the world, t ...
and
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
. It is common in most of its range, but largely absent from the arid northwest and the eastern lowlands of South Africa. It is found in open habitats such as grassland,
karoo The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe Khoemana (also known as !Orakobab or Korana) word is a semidesert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its extent is ...
scrub, thornbush and agricultural land, and often associates with farm animals.


Taxonomy

The pied starling was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
in 1789 by the German naturalist
Johann Friedrich Gmelin Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist. Education Johann Friedrich Gmelin was born as the eldest son of Philipp F ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
's ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. He placed it with the thrushes in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Turdus ''Turdus'' is a genus of medium-sized mostly insectivorous or omnivorous birds in the wider thrush family, Turdidae. The genus name ''Turdus'' is Latin for 'thrush'. Most of the species are called thrushes; the term thrush is also used for man ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Turdus bicolor''. Gmelin based his description on "Le merle brun du cap de Bonne Espérance" that had been described in 1775 by the French polymath
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French Natural history, naturalist, mathematician, and cosmology, cosmologist. He held the position of ''intendant'' (director) at the ''Jardin du Roi'', now ca ...
in his book ''Histoire Naturelle des Oiseaux''. The pied starling is now placed in the genus ''
Lamprotornis ''Lamprotornis'' is a large genus of glossy starlings all of which occur in Africa south of the Sahara. They have glossy blue or green upper parts, which is due to hollow melanin granules arranged in a single layer near the feather barbule's sur ...
'' that was introduced in 1820 by the Dutch zoologist
Coenraad Jacob Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch people, Dutch patrician, Zoology, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dutch Republic. Fro ...
. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.


Description

The adult of this long
starling Starlings are small to medium-sized passerine (perching) birds known for the often dark, glossy iridescent sheen of their plumage; their complex vocalizations including mimicking; and their distinctive, often elaborate swarming behavior, know ...
has mainly dully glossed black plumage except for a white lower belly and undertail. It has a white iris and yellow lower mandible. The sexes are alike, but the juvenile has unglossed plumage, a brown iris and a dull yellow lower mandible. There are no subspecies. This species has a number of calls, but the most familiar is a ''skeer kerrra kerrra''. There is also a soft warbling song.


Behaviour


Breeding

The pied starling usually nests in tunnels in river banks, but will use holes in buildings, straw bales or natural tree holes. There is a record of a nest being constructed in a wrecked ship from the shore. The nest is lined with a wide variety of plant material and items of human waste such as paper and rope. The female typically lays four eggs, though clutches of two to six are known. The eggs are blue-green, immaculate or with some red spots. The female alone incubates for , and the chicks fledge in another . Both parents feed the chicks, assisted by helpers, which are typically subadult or unmated birds. This cooperative breeding is reinforced by mutual allofeeding between adults, behaviour reinforced by the bright
gape The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, grasping, and holding (in probing for food, eating, manipulating and ca ...
, a feature normally lost in adults of most bird species. This starling is commonly double-brooded. It may be parasitised by the
great spotted cuckoo The great spotted cuckoo (''Clamator glandarius'') is a member of the cuckoo order of birds, the Cuculiformes, which also includes the roadrunners, the anis and the coucals. It is widely spread throughout Africa and the Mediterranean Basin. I ...
and
greater honeyguide The greater honeyguide (''Indicator indicator'') is a bird in the family honeyguide, Indicatoridae, Floristic kingdom#Paleotropical Kingdom, paleotropical near passerine birds related to the woodpeckers. Its English language, English and scientif ...
.


Roosting

The pied starling is gregarious and when not breeding will form large flocks, sometimes numbering more than . Its roosts may be shared by
lesser kestrel The lesser kestrel (''Falco naumanni'') is a small falcon. This species breeds from the Mediterranean across Afghanistan and Central Asia, to China and Mongolia. It is a summer bird migration, migrant, wintering in Africa and Pakistan and someti ...
s or
wattled starling The wattled starling (''Creatophora cinerea'') is a nomadic resident bird in eastern and southern Africa. It is a species of grassland, open woodland, and cultivation. This is the only African starling that appears to show affinities with the As ...
s. It will feed with
European starling The common starling (''Sturnus vulgaris''), also known simply as the starling in Great Britain and Ireland, and as European starling in North America, is a medium-sized passerine bird in the starling family, Sturnidae. It is about long and ha ...
s, but they rarely roost together.


Feeding

Like other starlings, the pied starling is an
omnivore An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize t ...
, taking a wide range of
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s, seeds and berries, but its diet is mainly
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, including many
ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s and
termite Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...
s. It will take
figs The fig is the edible fruit of ''Ficus carica'', a species of tree or shrub in the flowering plant family Moraceae, native to the Mediterranean region, together with western and southern Asia. It has been cultivated since ancient times and i ...
from gardens and some human food discards. Foraging is frequently near livestock, with birds feeding on insects disturbed by the animals and also perching on cattle or sheep to remove
ectoparasites Parasitism is a close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The en ...
. The pied starling has sometimes been seen as a pest when it takes soft fruit such as grapes or figs, and was also itself considered good eating. However, it is little persecuted at present.


Status

This species has a large range, estimated at . The population size has not been quantified, but it is believed to be large as the species is described as 'common' in at least parts of its range. It is not believed to approach the thresholds for the population decline criterion of the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
(i.e. declining more than 30% in ten years or three generations). For these reasons, the species is evaluated as Least Concern.


References

{{Taxonbar, from=Q3180337


External links

*(African) Pied starling
Species text in The Atlas of Southern African Birds
pied starling Birds of Southern Africa pied starling pied starling