''Phytelephas seemannii'',
commonly called Panama ivory palm, is a species of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the family
Arecaceae
The Arecaceae () is a family (biology), family of perennial plant, perennial, flowering plants in the Monocotyledon, monocot order Arecales. Their growth form can be climbing palm, climbers, shrubs, tree-like and stemless plants, all commonly k ...
. It is one of the plants used for
vegetable ivory
Vegetable ivory or Phytelephas, tagua nut is a product made from the very hard white endosperm of the seeds of certain Arecaceae, palm trees. Vegetable ivory is named for its resemblance to ivory, animal ivory. Vegetable ivory is naturally white ...
.
Names
The
species epithet Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In Taxonomy (biology), taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany) ...
''seemannii'' honors botanist
Berthold Carl Seemann
Berthold Carl Seemann (25 February 1825, in Kingdom of Hanover – 10 October 1871, in Nicaragua) was a German botanist. He travelled widely and collected and described plants from the Pacific and South America.
In 1844 he travelled to the Unit ...
who collected some of the first specimens, including the
lectotype
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...
.
In
Spanish
Spanish might refer to:
* Items from or related to Spain:
**Spaniards are a nation and ethnic group indigenous to Spain
**Spanish language, spoken in Spain and many countries in the Americas
**Spanish cuisine
**Spanish history
**Spanish culture
...
it is called (), (), and
tagua
''Phytelephas'' is a genus containing six known species of dioecious palms (family Arecaceae), occurring from southern Panama along the Andes to Ecuador, Bolivia, Colombia, northwestern Brazil, and Peru. They are commonly known as ivory palms, ...
.
In
Colombian Spanish
Colombian Spanish () is a grouping of the varieties of Spanish spoken in Colombia. The term is of more geographical than linguistic relevance, since the dialects spoken in the various regions of Colombia are quite diverse. The speech of the nor ...
it is additionally known as .
In
Guna it is ,
or .
In both the
Quechua and
Choco languages
The Choco languages (also Chocoan, Chocó, Chokó) are a small family of Indigenous languages spread across Colombia and Panama.
Family division
Choco consists of six known branches, all but two of which are extinct.
*The Emberá languages ...
it is called .
Habitat
''Phytelephas seemannii'' is native to
Colombia
Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
and
Panama
Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
, with much of it growing in shaded areas by rivers in
lowland
Upland and lowland are conditional descriptions of a plain based on elevation above sea level. In studies of the ecology of freshwater rivers, habitats are classified as upland or lowland.
Definitions
Upland and lowland are portions of a ...
rainforest
Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
in Colombia's
Pacific/Chocó natural region
The Pacific/Chocó region is one of the five major natural regions of Colombia. Ecologically, this region belongs entirely to the Chocó Biogeographic Region and is considered a biodiversity hotspot. It also has areas with the highest rainfal ...
.
It is usually found at elevations from in semi-
deciduous
In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
forests.
Subspecies
''Phytelephas seemannii'' has two subspecies,
''P. s.'' ssp. ''brevipes'' and
''P. s.'' ssp. ''seemannii''.
''P. s.'' ssp. ''brevipes'' is
endemic
Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the upper
Mamoní Valley in Panama, at or below in elevation, and may be a hybrid of ''P. seemannii'' and ''P. macrocarpa''.
Description
''Phytelephas seemannii'' most closely resembles ''
Phytelephas macrocarpa
''Phytelephas macrocarpa'' is a single-stemmed, unarmed, reclining or erect palm from the extreme northern coastal regions of South America, growing to some 12 m tall. It has been introduced and cultivated in tropical regions all over the world. ...
''.
However, the former has
leaves
A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
that have fewer
pinnae which are larger.
Its trunk is also not upright but "creeping" and
decumbent.
The tree is generally less than tall, with
inflorescence
In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s below the mark.
Its
spathe
In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale.
Bracts are usually different from foliage leaves in size, color, shape or texture. They also look ...
s are double instead of in threes or fours.
On the male flowers are only 36
stamen
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filament ...
s and not the hundreds of other species.
The heads contain fewer fruits than other species, but inside are more nuts that are larger.
Typically each fruit has 5 seeds protected by a fibrous coat, and each inflorescence has up to 8 fruits.
Each tree can have dozens of inflorescences at a time.
In immature seeds, the
endosperm
The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the Embryo#Pla ...
is a liquid, like in a
coconut
The coconut tree (''Cocos nucifera'') is a member of the palm tree family (biology), family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, ...
, and then later it hardens as the fruit wall softens and deteriorates.
Ecology
Panama ivory palm trees flower after the end of the
dry season
The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The t ...
, between February and May.
The flowers are
pollinated by insects, specifically by two types of
rove beetle
The rove beetles are a family (biology), family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With over 66,000 species in thousand ...
s,
pollen-eating ''
Amazoncharis spp.'' and their predators in the genus ''
Xanthopygus''.
The ''Amazoncharis'' beetles hollow out egg chambers within the male
inflorescence
In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
, similar to how beetles in the related
subtribe
Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants). The first use of this word dates back to the late 19th century. An example of ...
''
Gyrophaenina'' do inside of
mushroom
A mushroom or toadstool is the fleshy, spore-bearing Sporocarp (fungi), fruiting body of a fungus, typically produced above ground on soil or another food source. ''Toadstool'' generally refers to a poisonous mushroom.
The standard for the n ...
s.
Squirrel
Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s and
agouti
The agouti (, ) or common agouti is any of several rodent species of the genus ''Dasyprocta''. They are native to Central America, northern and central South America, and the southern Lesser Antilles. Some species have also been introduced else ...
s will eat the fleshy inner
mesocarp
Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits.
Fruitlike structures may develop directly from the seed itself rather tha ...
surrounding the
endocarp
Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits.
Fruitlike structures may develop directly from the seed itself rather th ...
of the fruit, but do not eat the extremely hard
endosperm
The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the Embryo#Pla ...
.
The rock-hard endosperm also makes the seed immune from most
insect pests.
Seed dispersers include the Central American agouti (''
Dasyprocta punctata''),
and lowland paca (''
Agouti paca'').
Uses
The seeds are traded at a regional international level as
vegetable ivory
Vegetable ivory or Phytelephas, tagua nut is a product made from the very hard white endosperm of the seeds of certain Arecaceae, palm trees. Vegetable ivory is named for its resemblance to ivory, animal ivory. Vegetable ivory is naturally white ...
,
which is also called tagua. This commercial use is a threat to the species, but progress is being made on using more sustainable practices and conservation.
As the tree typically grows only tall, it fortunately was not chopped down to harvest the seeds as was done with ''
Phytelephas aequatorialis'' at the peak of tagua harvesting.
The jelly-like liquid in the immature seeds, which later turns into the vegetable ivory, is edible.
Occasionally in the
marketplace
A marketplace, market place, or just market, is a location where people regularly gather for the purchase and sale of provisions, livestock, and other goods. In different parts of the world, a marketplace may be described as a ''souk'' (from ...
s of
Guna Yala
Guna Yala, also known as Kuna Yala or by its former name San Blas, is a ''Comarca#Panama, comarca indígena'' (indigenous province) in northeast Panama. Guna Yala is home to the indigenous people known as the Guna people, Gunas. Its capital ...
the thin crust surrounding the ivory is sold as food.
In Colombia the fronds are sometimes used for
thatch
Thatching is the craft of building a roof with dry vegetation such as straw, Phragmites, water reed, Cyperaceae, sedge (''Cladium mariscus''), Juncus, rushes, Calluna, heather, or palm branches, layering the vegetation so as to shed water away fr ...
.
References
External links
*
*
{{Taxonbar, from=Q5438324
seemannii
Flora of Colombia
Flora of Panama
Conservation dependent plants
Ivory
Sculpture materials
Handicrafts
Plants described in 1912
Non-timber forest products
Taxonomy articles created by Polbot