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''Phyllodon'' (meaning "leaf tooth") was a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of small
ornithischian Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxford ...
-aged
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
Camadas de Guimarota Formation of
Leiria Leiria (; cel-x-proto, ɸlāryo) is a city and municipality in the Central Region of Portugal. It is the 2nd largest city in that same region, with a municipality population of 128,640 (as of 2021) in an area of . It is the seat of its own distr ...
,
Portugal Portugal, officially the Portuguese Republic, In recognized minority languages of Portugal: :* mwl, República Pertuesa is a country located on the Iberian Peninsula, in Southwestern Europe, and whose territory also includes the Macaronesian ...
and possibly also the
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. St ...
-aged Chipping Norton Limestone of
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...
.Listed as "cf. ''Phyllodon'' sp." in "10.3 Gloucestershire, England; 2. Chipping Norton Formation," in Weishampel, et al. (2004). Page 538. It may have been closely related to contemporaneous dinosaurs in North America. This genus is known from
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, te ...
and possibly partial lower jaws. The nam
is also in use
for a genus of modern
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta ('' sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and ...
, but this is not considered to be a problem because the two organisms are in two different
kingdoms Kingdom commonly refers to: * A monarchy ruled by a king or queen * Kingdom (biology), a category in biological taxonomy Kingdom may also refer to: Arts and media Television * ''Kingdom'' (British TV series), a 2007 British television drama s ...
.


History

''Phyllodon'' is based on MGSP G5, a partial
lower jaw In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
tooth recovered from a lignite marl in a mine near the city of
Leiria Leiria (; cel-x-proto, ɸlāryo) is a city and municipality in the Central Region of Portugal. It is the 2nd largest city in that same region, with a municipality population of 128,640 (as of 2021) in an area of . It is the seat of its own distr ...
. Richard Thulborn, who described the genus, added an upper beak tooth (MGSP G2). He regarded the new genus as a
hypsilophodont Hypsilophodontidae (or Hypsilophodontia) is a traditionally used family of ornithopod dinosaurs, generally considered invalid today. It historically included many small bodied bipedal neornithischian taxa from around the world, and spanning from ...
id, and presented a conjectural restoration of the tooth arrangement.
Peter Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosau ...
, reviewing Late Jurassic North American hypsilophodontids a few years later, found that the ''Phyllodon'' teeth best matched those of ''
Nanosaurus ''Nanosaurus'' ("small or dwarf lizard") is the name given to a genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic-age. Its fossils are known from the Morrison Formation of the south-wester ...
'', and agreed with a hypsilophodontid identity because the lower jaw tooth is asymmetric in front and back views. Sometime before 2004, possible ''Phyllodon'' remains were recovered from the Chipping Norton Limestone of
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...
. It is likely that these remains actually belonged to an indeterminate
ornithischian Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
separate from ''Phyllodon''."10.3 Gloucestershire, England; 2. Chipping Norton Formation," in Weishampel, et al. (2004). Page 538. Because of the sparse material, ''Phyllodon'' has often been tossed off as a
dubious Doubt is a mental state in which the mind remains suspended between two or more contradictory propositions, unable to be certain of any of them. Doubt on an emotional level is indecision between belief and disbelief. It may involve uncertainty ...
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
ornithopod of uncertain affinities. However, more material that might belong to this genus has been recovered from the original locality and described. Included in this material are over 120 more teeth from all parts of the jaw and four partial lower jaws with the teeth lost. Oliver Rauhut, who described the new material, tentatively identified the lower jaws as ''Phyllodon'' due to there being no other similar dinosaurs found at the locality. The teeth were very small (up to 3 millimeters across, or 0.1 inches) and possibly juvenile. He also found additional diagnostic characteristics for ''Phyllodon'' in the new material, including very tall
upper jaw The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
teeth, indicating that it could be a valid genus after all. After comparing it to other hypsilophodonts, he found that it best matched the roughly contemporaneous '' Drinker'' of the North American
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone, ...
, with various details suggesting that they were closely related. Similarly, Galton found its teeth to be similar to those of ''Drinker'' and ''Nanosaurus'' in his 2006 review.


Paleobiology

As a hypsilophodontid or other basal ornithopod, ''Phyllodon'' would have been a
bipedal Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
. Its size has not been estimated, but as most adult hypsilophodonts were long, this genus would probably have been of similar size. Its similarity to the North American ''Drinker'' and ''Nanosaurus'' is another piece of evidence linking Late Jurassic Portuguese dinosaur
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''biota''. Zoo ...
s with the contemporaneous Morrison Formation dinosaurs.


References

{{Taxonbar, from=Q2413478 Ornithischian genera Kimmeridgian life Late Jurassic dinosaurs of Europe Jurassic Portugal Fossils of Portugal Fossil taxa described in 1973