subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
of the plant bug family
Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs". Common names include plant bugs, leaf bugs, and g ...
. Species of this family are found worldwide.
In research published in 2013, the subfamily Phylinae was reorganized. The tribe Auricillocorini is now considered a synonym of Hallodapini, and the tribe Pronotocrepini a synonym of the subtribe Cremnorrhina. New tribes were created under the new or resurrected names Decomiini, Exocarpocorini, Cremnorrhini, Nasocorini, and Semiini. The revised subfamily consists of nine tribes containing around 400 genera and over 2000 species.
; Subfamily Phylinae Douglas & Scott, 1865
: Tribe
Cremnorrhinini
Cremnorrhinini is a tribe of plant bugs in the family Miridae. There are about 50 genera in Cremnorrhinini, all but five in the subtribe Cremnorrhinina.
Genera
These 50 genera belong to the tribe Cremnorrhinini:
; Tribe Cremnorrhinini Reuter, 1 ...
Decomiini
Decomiini is a tribe of plant bugs in the family Miridae. There are about six genera in Decomiini.
Genera
These six genera belong to the tribe Decomiini:
* '' Aurantiocoris'' Schuh & Schwartz, 2004 - Western Nearctic
* '' Decomia'' Poppius, 191 ...
Schuh & Menard, 2013
: Tribe
Exaeretini
Exaeretini is a tribe of plant bugs in the family Miridae. There are more than 20 genera in Exaeretini.
Genera
These 24 genera belong to the tribe Exaeretini:
* '' Anonychiella'' Reuter, 1912 - Palearctic
* '' Aphaenophyes'' Reuter, 1899 - Pal ...
Futon, 1875
: Tribe
Hallodapini
Hallodapini is a tribe of plant bugs in the family Miridae. There are more than 50 genera in Hallodapini.
Most species of this tribe are found in the old world, including Australia. Only one genus, Cyrtopeltocoris, is found in North America.
G ...
Van Duzee, 1916
: Tribe
Leucophoropterini
Leucophoropterini is a tribe of plant bugs in the family Miridae. There are more than 20 genera and over 100 species in Leucophoropterini. Most species of this tribe are found in eastern Asia and Australia, with a single genus, ''Tuxedo'', found ...
Nasocorini
Nasocorini is a tribe of plant bugs in the family Miridae. There are more than 50 genera in Nasocorini.
Genera
These 59 genera belong to the tribe Nasocorini:
* '' Adenostomocoris'' Schuh & Schwartz, 2004 - Southwest Nearctic
* '' Agrametra'' B ...
Reuter, 1883
: Tribe
Phylini
Phylini is a tribe of plant bugs in the family Miridae, based on the type genus ''Phylus''. There are at least 440 described species in Phylini.
Subtribes and selected genera
* Full list of Phylini genera here Keltoniina
Auth. Schuh & Menard, 2 ...
Douglas & Scott, 1865
:: Subtribe Keltoniina Schuh & Menard, 2013
:: Subtribe Oncotylina Douglas & Scott, 1865
:: Subtribe Phylina Douglas & Scott, 1865
: Tribe
Pilophorini
Pilophorini is a tribe (biology), tribe of Miridae, plant bugs. The type genus is ''Pilophorus'' (Hahn, 1826). Schuh's analysis indicates that the Pilophorini originated in tropical Gondwanaland and subsequently spread into the temperate Northern ...
Douglas & Scott, 1876
: Tribe
Semiini
Semiini is a tribe of plant bugs in the family Miridae. There are more than 30 genera in Semiini.
Genera
These 33 genera belong to the tribe Semiini:
; Subtribe Exocarpocorina Schuh & Menard, 2013
: '' Ampimpacoris'' Weirauch & Schuh, 2011 - Ne ...
Phylinae contains the following genera:
; Subfamily Phylinae Douglas & Scott, 1865
: Tribe
Cremnorrhinini
Cremnorrhinini is a tribe of plant bugs in the family Miridae. There are about 50 genera in Cremnorrhinini, all but five in the subtribe Cremnorrhinina.
Genera
These 50 genera belong to the tribe Cremnorrhinini:
; Tribe Cremnorrhinini Reuter, 1 ...
Amblytylus
''Amblytylus'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or ...
'' Fieber, 1858
::: Genus '' Brachyceratocoris'' Knight, 1968
::: Genus '' Calidroides'' Schwartz, 2005
::: Genus ''
Coquillettia
''Coquillettia'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs ...
'' Uhler, 1890
::: Genus '' Cremnorrhinus'' Reuter, 1880
::: Genus '' Dacota'' Uhler, 1872
::: Genus '' Denticulophallus'' Schuh, 1974
::: Genus '' Ethelastia'' Reuter, 1876
::: Genus '' Euderon'' Puton, 1888
::: Genus '' Excentricoris'' Carvalho, 1955
::: Genus '' Guentherocoris'' Schuh & Schwartz, 2004
::: Genus '' Harpocera'' Curtis, 1838
::: Genus '' Leutiola'' Wyniger, 2012
::: Genus '' Lopidodenus'' V. Putshkov, 1974
::: Genus ''
Lopus
''Lopus'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mi ...
'' Hahn, 1833
::: Genus '' Macrotylus'' Fieber, 1858
::: Genus '' Orectoderus'' Uhler, 1876
::: Genus '' Pachyxyphus'' Fieber, 1858
::: Genus '' Paralopus'' Wagner, 1957
::: Genus '' Pronotocrepis'' Knight, 1929
::: Genus '' Shendina'' Linnavuori, 1975
::: Genus '' Strophopoda'' Van Duzee, 1916
::: Genus '' Teleorhinus'' Uhler, 1890
::: Genus '' Ticua'' Wyniger, 2012
::: Genus '' Utopnia'' Reuter, 1881
::: Genus '' Zinjolopus'' Linnavuori, 1975
: Tribe
Decomiini
Decomiini is a tribe of plant bugs in the family Miridae. There are about six genera in Decomiini.
Genera
These six genera belong to the tribe Decomiini:
* '' Aurantiocoris'' Schuh & Schwartz, 2004 - Western Nearctic
* '' Decomia'' Poppius, 191 ...
Exaeretini
Exaeretini is a tribe of plant bugs in the family Miridae. There are more than 20 genera in Exaeretini.
Genera
These 24 genera belong to the tribe Exaeretini:
* '' Anonychiella'' Reuter, 1912 - Palearctic
* '' Aphaenophyes'' Reuter, 1899 - Pal ...
Yotvata
:''"Yotvat" is also the Hebrew name for Tiran Island''.
Yotvata ( ''yotváta'' (but commonly pronounced ''yotvetá''), ) is a kibbutz in the Aravah, Arava Valley in southern Israel. Located on the Highway 90 (Israel–Palestine), Highway 90's seg ...
'' Linnavuori, 1964
: Tribe
Hallodapini
Hallodapini is a tribe of plant bugs in the family Miridae. There are more than 50 genera in Hallodapini.
Most species of this tribe are found in the old world, including Australia. Only one genus, Cyrtopeltocoris, is found in North America.
G ...
Skukuza
Skukuza (formerly Reserve, Sabi Bridge and Sikhukhuza), a town in Mpumalanga located 57 km east of Hazyview at the confluence of the N'waswitshaka River, N'waswitshaka and Sabie Rivers, is the administrative headquarters of the Kruger Natio ...
Leucophoropterini
Leucophoropterini is a tribe of plant bugs in the family Miridae. There are more than 20 genera and over 100 species in Leucophoropterini. Most species of this tribe are found in eastern Asia and Australia, with a single genus, ''Tuxedo'', found ...
Sejanus
Lucius Aelius Sejanus ( – 18 October AD 31), commonly known as Sejanus (), was a Roman soldier and confidant of the Roman Emperor Tiberius. Of the Equites class by birth, Sejanus rose to power as prefect of the Praetorian Guard, the imperia ...
Tuxedo
Black tie is a semi-formal Western dress code for evening events, originating in British and North American conventions for attire in the 19th century. In British English, the dress code is often referred to synecdochically by its principal ...
'' Schuh, 2001
: Tribe
Nasocorini
Nasocorini is a tribe of plant bugs in the family Miridae. There are more than 50 genera in Nasocorini.
Genera
These 59 genera belong to the tribe Nasocorini:
* '' Adenostomocoris'' Schuh & Schwartz, 2004 - Southwest Nearctic
* '' Agrametra'' B ...
Campylomma
''Campylomma'' is a genus of bugs in the family Miridae and tribe Nasocorini.Henry TJ, Froeschner RC (1988). ''Catalog of the Heteroptera, True Bugs of Canada and the Continental United States''. Brill Academic Publishers.
Species
The following ...
'' Reuter, 1878
:: Genus '' Chinacapsus'' Wagner, 1961
:: Genus ''
Chlamydatus
''Chlamydatus'' is a genus of plant bugs in the family Miridae. There are more than 30 described species in ''Chlamydatus''.
Species
These 32 species belong to the genus ''Chlamydatus'':
* ''Chlamydatus acanthioides'' (J.Sahlberg, 1875)
* ''Chl ...
Phylini
Phylini is a tribe of plant bugs in the family Miridae, based on the type genus ''Phylus''. There are at least 440 described species in Phylini.
Subtribes and selected genera
* Full list of Phylini genera here Keltoniina
Auth. Schuh & Menard, 2 ...
Europiella
''Europiella'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs o ...
'' Reuter, 1909
::: Genus '' Eurycolpus'' Reuter, 1875
::: Genus '' Galbinocoris'' Weirauch, 2006
::: Genus '' Glaucopterum'' Wagner, 1963
::: Genus '' Hamatophylus'' Weirauch, 2006
::: Genus ''
Hoplomachus
A hoplomachus (pl. hoplomachi) (hoplon meaning "equipment" in Greek) was a type of gladiator in ancient Rome, armed to resemble a Greek hoplite (soldier with heavy armor and helmet, a small, round, concave shield, a spear and a sword). The hopl ...
Oncotylus
''Oncotylus'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or ...
Phyllopidea
''Phyllopidea'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs ...
'' Knight, 1919
::: Genus '' Phymatopsallus'' Knight
::: Genus '' Piceophylus'' Schwartz & Schuh, 1999
::: Genus '' Pinophylus'' Schwartz & Schuh, 1999
::: Genus '' Placochilus'' Fieber, 1858
::: Genus ''
Plagiognathus
''Plagiognathus'' is a genus of jumping tree bugs in the family Miridae. There are more than 120 described species in ''Plagiognathus''.
''Plagiognathus'' species
These 123 species belong to the genus ''Plagiognathus'':
* ''Plagiognathus alasha ...
'' Fieber, 1858
::: Genus ''
Plesiodema
''Plesiodema'' is a genus of true bugs belonging to the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as cap ...
'' Reuter, 1875
::: Genus '' Pleuroxonotus'' Reuter, 1903
::: Genus '' Pronototropis'' Reuter, 1879
::: Genus ''
Psallodema
''Psallodema'' is a genus of mostly European capsid bugs in the tribe Phylini, erected by V.G. Putshkov in 1970.
The species ''Psallodema fieberi'' is recorded from northern Europe including the British Isles
The British Isles are an arch ...
'' V. Putshkov, 1970
::: Genus '' Psallomorpha'' Duwal Yasunaga, & Lee, 2010
::: Genus '' Psallovius'' T. Henry
::: Genus '' Pygovepres'' Weirauch, 2006
::: Genus '' Quercophylus'' Weirauch, 2006
::: Genus '' Quernocoris'' Weirauch, 2006
::: Genus '' Ranzovius'' Distant, 1893
::: Genus ''
Rhinocapsus
''Rhinocapsus'' is a genus of North American plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to ...
Sthenaropsidea
''Sthenaropsidea'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bu ...
'' Henry & Schuh, 2002
::: Genus '' Stictopsallus'' Schuh, 2006
::: Genus '' Stirophylus'' Eckerlein & Wagner, 1965
::: Genus '' Stoebea'' Schuh, 1974
::: Genus '' Tapirula'' Carapezza, 1997
::: Genus ''
Tinicephalus
''Tinicephalus'' is a genus of mostly European Miridae, capsid bugs in the Tribe (biology), tribe Phylini, erected by Franz Xaver Fieber in 1858. The species ''Tinicephalus hortulanus'' is recorded from northern Europe including the British Is ...
'' Fieber, 1858
::: Genus '' Tragiscocoris'' Fieber, 1861
::: Genus '' Vanduzeephylus'' Schuh & Schwartz, 2004
::: Genus '' Vesperocoris'' Weirauch, 2006
::: Genus '' Viscacoris'' Weirauch, 2009
::: Genus '' Zophocnemis'' Kerzhner, 1962
:: Subtribe Phylina Douglas & Scott, 1865
::: Genus '' Adelphophylus'' Wagner, 1959
::: Genus '' Agraptocoris'' Reuter, 1903
::: Genus '' Alnopsallus'' Duwal, Yasunaga, & Lee, 2010
::: Genus '' Alvarengamiris'' Carvalho, 1991
::: Genus '' Amazonophilus'' Carvalho & Costa, 1993
::: Genus '' Anapsallus'' Odhiambo, 1960
::: Genus '' Anomalocornis'' Carvalho & Wygodzinsky, 1945
::: Genus '' Arlemiris'' Carvalho, 1984
::: Genus '' Bicurvicoris'' Carvalho & Schaffner, 1973
::: Genus '' Botocudomiris'' Carvalho, 1979
::: Genus '' Brachycranella'' Reuter, 1905
::: Genus '' Conostethus'' Fieber, 1858
::: Genus '' Crassicornus'' Carvalho, 1945
::: Genus '' Darectagela'' Linnavuori, 1975
::: Genus '' Darfuromma'' Linnavuori, 1975
::: Genus '' Dignaia'' Linnavuori, 1975
::: Genus '' Dominiquella'' Linnavuori, 1983
::: Genus '' Ectagela'' Schmidt, K., 1939
::: Genus '' Ellacapsus'' Yasunaga, 2013
::: Genus '' Eremophylus'' Yasunaga, 2001
::: Genus '' Farsiana'' Linnavuori, 1998
::: Genus '' Gediocoris'' Wagner, 1964
::: Genus '' Ghazalocoris'' Linnavuori, 1975
::: Genus '' Gressittocapsus'' Schuh, 1984
::: Genus '' Icodema'' Reuter, 1875
::: Genus '' Indatractus'' Linnavuori, 1975
::: Genus '' Izyaius'' Schwartz, 2006
::: Genus '' Juniperia'' Linnavuori, 1965
::: Genus '' Knightensis'' Schaffner, 1978
::: Genus '' Lalyocoris'' Linnavuori, 1993
::: Genus '' Lasiolabopella'' Schuh, 1974
::: Genus ''
Lepidargyrus
''Lepidargyrus'' is a genus of plant bugs in the family Miridae. There are about 14 described species in ''Lepidargyrus''.
Species
These 14 species belong to the genus ''Lepidargyrus'':
* '' Lepidargyrus ancorifer'' (Fieber, 1858)
* '' Lepidarg ...
'' Muminov, 1962
::: Genus '' Lepidocapsus'' Poppius, 1914
::: Genus '' Leptoxanthus'' Reuter, 1905
::: Genus '' Liviopsallus'' Carapezza, 1982
::: Genus '' Mendozaphylus'' Carvalho & Carpintero, 1991
::: Genus '' Millerimiris'' Carvalho, 1951
::: Genus '' Mixtecamiris'' Carvalho & Schaffner, 1973
::: Genus '' Natalophylus'' Schuh, 1974
::: Genus '' Nubaia'' Linnavuori, 1975
::: Genus '' Orthonotus'' Stephens, 1829
::: Genus '' Parafulvius'' Carvalho, 1954
::: Genus '' Paravoruchia'' Wagner, 1959
::: Genus '' Phylus'' Hahn, 1831
::: Genus '' Plagiognathidea'' Poppius, 1914
::: Genus '' Platyscytisca'' Costa & Henry, 1999
::: Genus '' Platyscytus'' Reuter, 1907
::: Genus '' Porophoroptera'' Carvalho & Gross, 1982
::: Genus '' Psallus'' Fieber, 1858
::: Genus '' Roudairea'' Puton & Reuter, 1886
::: Genus '' Sasajiophylus'' Yasunaga, 2001
::: Genus '' Somalocoris'' Linnavuori, 1975
::: Genus '' Sthenarus'' Fieber, 1858
::: Genus '' Stibaromma'' Odhiambo, 1961
::: Genus '' Tibiopilus'' Carvalho & Costa, 1993
::: Genus '' Trevessa'' China, 1924
::: Genus '' Villaverdea'' Carvalho, 1990
::: Genus '' Widdringtoniola'' Schuh, 1974
::: Genus '' Zakanocoris'' V. Putshkov, 1970
::: Genus '' Zanchiophylus'' Duwal, Yasunaga, & Lee, 2010
: Tribe
Pilophorini
Pilophorini is a tribe (biology), tribe of Miridae, plant bugs. The type genus is ''Pilophorus'' (Hahn, 1826). Schuh's analysis indicates that the Pilophorini originated in tropical Gondwanaland and subsequently spread into the temperate Northern ...
Douglas & Scott, 1876
:: Genus ''
Alepidiella
''Alepidiella'' is a genus of plant bugs in the family Miridae
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs ...
'' Poppius, 1914
:: Genus '' Aloea'' Linnavuori, 1975
:: Genus '' Dilatops'' Weirauch, 2006
:: Genus '' Druthmarus'' Distant, 1909
:: Genus '' Ethatractus'' Linnavuori, 1975
:: Genus ''
Hypseloecus
''Hypseloecus'' is a genus of capsid bugs in the tribe Pilophorini, erected by Odo Reuter in 1891. Records are mostly from Europe, with some in Africa, Asia and Australia.Lasiolabops'' Poppius, 1914
:: Genus '' Neoambonea'' Schuh, 1974
:: Genus '' Parambonea'' Schuh, 1974
:: Genus '' Parasthenaridea'' Miller, 1937
:: Genus '' Pherolepis'' Kulik, 1968
:: Genus '' Pilophorus'' Hahn, 1826
:: Genus '' Pseudambonea'' Schuh, 1974
:: Genus '' Randallophorus'' Henry, 2013
:: Genus '' Spinolosus'' Zou, 1985
:: Genus '' Sthenaridea'' Reuter, 1885
: Tribe
Semiini
Semiini is a tribe of plant bugs in the family Miridae. There are more than 30 genera in Semiini.
Genera
These 33 genera belong to the tribe Semiini:
; Subtribe Exocarpocorina Schuh & Menard, 2013
: '' Ampimpacoris'' Weirauch & Schuh, 2011 - Ne ...
Knight, 1923
:: Subtribe Exocarpocorina Schuh & Menard, 2013
::: Genus '' Ampimpacoris'' Weirauch & Schuh, 2011
::: Genus '' Ancoraphylus'' Weirauch, 2007
::: Genus '' Araucanophylus'' Carvalho, 1984
::: Genus '' Basileobius'' Eyles & Schuh, 2003
::: Genus '' Chiloephylus'' Carvalho, 1984
::: Genus '' Cyrtodiridius'' Eyles & Schuh, 2003
::: Genus '' Exocarpocoris'' Weirauch, 2007
::: Genus '' Gonzalezinus'' Carvalho, 1981
::: Genus '' Halormus'' Eyles & Schuh, 2003
::: Genus '' Harpagophylus'' Schuh & Weirauch, 2010
::: Genus '' Jiwarli'' Soto & Weirauch, 2007
::: Genus ''
Leptidolon
''Leptidolon'' is a genus of plant bug
The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs". Common ...
'' Reuter, 1904
::: Genus '' Mecenopa'' Eyles & Schuh, 2003
::: Genus '' Melaleucoides'' Schuh & Weirauch, 2010
::: Genus '' Pimeleocoris'' Eyles & Schuh, 2003
::: Genus '' Polyozus'' Eyles & Schuh, 2003
::: Genus '' Thryptomenomiris'' Schuh & Weirauch, 2010
::: Genus '' Wallabicoris'' Schuh & Pedraza, 2010
::: Genus '' Xiphoidellus'' Weirauch & Schuh, 2011
::: Genus '' Xiphoides'' Eyles & Schuh, 2003
:: Subtribe Semiina Knight, 1923
::: Genus '' Criocoris'' Fieber, 1858
::: Genus '' Hoplomachidea'' Reuter, 1909
::: Genus '' Karoocapsus'' Schuh, 1974
::: Genus '' Monocris'' V. Putshkov, 1974
::: Genus '' Monospatha'' Eyles & Schuh 2003
::: Genus '' Myrmicopsella'' Poppius, 1914
::: Genus '' Nicholia'' Knight, 1929
::: Genus '' Parapseudosthenarus'' Schuh, 1974
::: Genus '' Pseudosthenarus'' Poppius, 1914
::: Genus '' Rakula'' Odhiambo, 1967
::: Genus '' Semium'' Reuter, 1876
::: Genus ''
Thoth
Thoth (from , borrowed from , , the reflex of " eis like the ibis") is an ancient Egyptian deity. In art, he was often depicted as a man with the head of an African sacred ibis, ibis or a baboon, animals sacred to him. His feminine count ...
'' Linnavuori, 1993
::: Genus '' Tytthus'' Fieber, 1864