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''Phrynoidis'' is a small genus of true toads, family
Bufonidae {{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae. Animals by common name Frogs ...
. They are found in Mainland Southeast Asia and the
Greater Sundas The Greater Sunda Islands (Indonesian language, Indonesian and Malay language, Malay: ''Kepulauan Sunda Besar'') are four tropical islands situated within Indonesian Archipelago, in the Pacific Ocean. The islands, Borneo, Java, Sulawesi and Sumat ...
. They are sometimes known as the rough toads or river toads.


Systematics

''Phrynoidis'' was included in ''
Bufo ''Bufo'' is a genus of true toads in the amphibian family Bufonidae. As traditionally defined, it was a wastebasket genus containing a large number of toads from much of the world, but following taxonomic reviews most of these have been moved ...
'' until 2006. Their
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
is the genus ''
Rentapia ''Rentapia'' is a genus of true toads, family Bufonidae. It is endemic to Southeast Asia and occurs in the Malay Peninsula (including extreme southern peninsular Thailand), Borneo, and Sumatra. It was erected in 2016 resolve the polyphyly of ''Pe ...
'' (formerly part of then
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...
''
Pedostibes The Malabar tree toad (''Pedostibes tuberculosus''), or warty Asian tree toad, is a species of toad found in forests along the Western Ghats of great Karnataka or Deccan. It is a small species and is found in wet tree hollows or leaf bases con ...
'').


Description

''Phrynoidis'' are large toads, with a maximum female snout–vent length of , depending on the species (males are smaller). Finger tips are dilated into keratinized, bulbous tips. Supernumerary palmar tubercles are present. The fingers have basal webbing. The
tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found ...
s have large oral disc that is as wide as the body. ''Phrynoidis'' are terrestrial
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
habitat generalists. They lay very large egg clutches.


Species

There are two species: * ''
Phrynoidis asper The Asian giant toad (''Phrynoidis asper''), sometimes referred to as the river toad, is a species of true toad native to Mainland Southeast Asia and the Greater Sundas. It is a medium-large toad, but it is easily confused with its larger relati ...
'' (Gravenhorst, 1829) * '' Phrynoidis juxtasper'' (Inger, 1964)


References

{{Taxonbar, from=Q2769608 Amphibian genera Amphibians of Asia Taxa named by Leopold Fitzinger