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''Phragmipedium'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of the
Orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan plants that are found in almost every habitat on Eart ...
family (Orchidaceae) (Subfamily
Cypripedioideae Cypripedioideae is a subfamily of orchids commonly known as lady's slipper orchids, lady slipper orchids or slipper orchids. Cypripedioideae includes the genera ''Cypripedium, Mexipedium, Paphiopedilum, Phragmipedium'' and '' Selenipedium''. ...
) and the only genus comprised in the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Phragmipedieae and
subtribe Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants). The first use of this word dates back to the late 19th century. An example of ...
Phragmipediinae. The name of the genus is derived from the Greek ''phragma'', which means "division", and ''pedium'', which means "slipper" (referring to the pouch). It is abbreviated 'Phrag' in trade journals. About 20
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of these lady's slipper orchids are known from SW
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, Central and tropical
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. All ''Phragmipedium'' species are listed under Appendix I of the
Convention on International Trade in Endangered Species Convention may refer to: * Convention (norm), a custom or tradition, a standard of presentation or conduct ** Treaty, an agreement in international law ** Convention (political norm), uncodified legal or political tradition * Convention (meeting ...
(CITES), meaning that commercial international trade in wild-sourced specimens is prohibited, while non-commercial trade is regulated.


Taxonomy

The genus ''Phragmipedium '' is divided into several sections : * Phragmipedium : ''P. caudatum, P. exstaminodium, P. lindenii'' * Himantopetalum : ''P. caricinum, P. christiansenianum, P. pearcei, P. klotzscheanum, P. richteri, P. tetzlaffianum''. * Platypetalum : ''P. lindleyanum'' * Lorifolia : ''P. boissierianum, P. hirtzii, P. longifolium, P. vittatum'' * Micropetalum : ''P. besseae, P. besseae var. dalessandroi, P. fischeri, P. schlimii.'' * Schluckebieria : ''P. kovachii'' The exact number of species is still being discussed among specialists : O. Gruss recognizes 20 species, compared to the 15 species accepted by Lucile M. McCook (see References). Most ''Phragmipedium'' species are either
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
,
epiphytic An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
or lithophytic in habit. They show a unique shieldlike staminode, long, moustache-like petals and a 3-locular ovary. The large pouchlike
lip The lips are a horizontal pair of soft appendages attached to the jaws and are the most visible part of the mouth of many animals, including humans. Mammal lips are soft, movable and serve to facilitate the ingestion of food (e.g. sucklin ...
is curved inwards at the margins. The acute leaves attain a length of about 80 cm. The stem lacks pseudobulbs and grows about 80 cm high, showing 2 to 3 flowers. ''Phragmipedium besseae'' was first found in Peru by Elizabeth Locke Besse in 1981. Soon afterwards, the site was plundered and destroyed by orchid hunters. Luckily enough seed was preserved, to avert extinction. This orchid is unusual, because its flowers have a bright orange-red to almost strong salmon-red color (there is also a yellow variety), unseen in any lady's slipper orchid. The oval-shaped petals are wide. The narrow leaves are elliptic in shape. It has since been used extensively in hybridization. ''Phragmipedium caudatum'' is considered a complex, i.e. it could contain several species or subspecies, based on differences in flower size and color. This orchid with a short stem is semi-terrestrial, semi-lithophytic to epiphytic, depending on the substrate . The cream-colored flowers are laced with greenish stripes. The lateral spiraling, drooping petals are red-tinted and very long, even reaching the soil. They grow on wet, moss-covered hillsides. ''Phragmipedium lindleyanum'', named after
John Lindley John Lindley Fellow of the Royal Society, FRS (5 February 1799 – 1 November 1865) was an English botanist, gardener and orchidology, orchidologist. Early years Born in Old Catton, Catton, near Norwich, England, John Lindley was one of four c ...
, forms a rosette of five long linear leaves with a yellow margin, reaching a length of 50 cm. The erect raceme can grow as high as 1 m. It is many-flowered and sometimes branched at the basis. The flowers open in succession, giving the orchid a long blooming period. The hoary flowers are green with brown veins. The glabrous, pouchlike lip is yellow, with red veins. ''Phragmipedium longifolium'', described in 1852 by H.G. Reichenbach f. and J. v. Warscewicz, has long lanceolate leaves without yellow margin, growing to a length of 60 cm. The inflorescence reaches a length of 1 m, with about 10 flowers, opening in succession. The long lateral petals are purplish green. The rather small glabrous labellum has a green color. Allied genera include ''
Paphiopedilum ''Paphiopedilum'', often called the Venus slipper, is a genus of the lady slipper orchid subfamily '' Cypripedioideae'' of the flowering plant family Orchidaceae. The genus comprises some 80 accepted taxa including several natural hybrids. The ...
'', '' Selenipedium'', ''
Cypripedium ''Cypripedium'' is a genus of 58 species and nothospecies of hardy Orchidaceae, orchids; it is one of five genera that together compose the subfamily of lady's slipper orchids (Cypripedioideae). They are widespread across much of the Northern He ...
'' and the monotypic '' Mexipedium''. There are many interspecific hybrids. Rare crossings have been made between ''Phragmipedium'' and ''Paphiopedilum''. The genus ''Uropedium'' Lindl. is generally included in ''Phragmipedium''.


Species

The following species ordered by sections are recognized by
Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...
:


Natural Hybrids

The following Natural hybrids are recognized:


References

* Albert, V.Z. & B.Pettersson, ''Expansion of Genus Paphiopedilum Pfitzer to Include All Conduplicate-leaved Slipper Orchids'' (Cypripedioideae: Orchidaceae). Lindleyana 9(2) 133-139 (1994). * McCook, L. ''An Annotated Checklist of the Genus Phragmipedium'' - 28 p. - Special Publication of the Orchid Digest (1998). * Gruss, O. 2003. ''A checklist of the genus Phragmipedium''. Orchid Digest 67: 213–241. * Braem, G. J., Ohlund, S., and Quene, R. J. 2004. ''The real Phragmipedium warszewiczianum: a clarification of the Phragmipedium caudatum complex'' (Phragmipedium section Phragmipedium). Orquideologia 23(2): 87-136 * Pridgeon, A.M.; Cribb, P.J.; Chase, M.W. & F. N. Rasmussen (1999): Genera Orchidacearum Vol.1, Oxford U. Press.


External links

* {{Authority control Orchid genera Epiphytic orchids