Photopic Negative Response
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Photopic vision is the
vision Vision, Visions, or The Vision may refer to: Perception Optical perception * Visual perception, the sense of sight * Visual system, the physical mechanism of eyesight * Computer vision, a field dealing with how computers can be made to gain und ...
of the
eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...
under well-lit conditions (
luminance Luminance is a photometric measure of the luminous intensity per unit area of light travelling in a given direction. It describes the amount of light that passes through, is emitted from, or is reflected from a particular area, and falls wit ...
levels from 10 to 108  cd/m2). In humans and many other animals, photopic vision allows
color perception Color vision, a feature of visual perception, is an ability to perceive differences between light composed of different frequencies independently of light intensity. Color perception is a part of the larger visual system and is mediated by a co ...
, mediated by
cone cells Cone cells or cones are photoreceptor cells in the retina of the vertebrate eye. Cones are active in daylight conditions and enable photopic vision, as opposed to rod cells, which are active in dim light and enable scotopic vision. Most vertebr ...
, and a significantly higher
visual acuity Visual acuity (VA) commonly refers to the clarity of visual perception, vision, but technically rates an animal's ability to recognize small details with precision. Visual acuity depends on optical and neural factors. Optical factors of the eye ...
and temporal resolution than available with
scotopic vision In the study of visual perception, scotopic vision (or scotopia) is the vision of the eye under low-light conditions. The term comes from the Greek ''skotos'', meaning 'darkness', and ''-opia'', meaning 'a condition of sight'. In the human eye, c ...
. The human eye uses three types of cones to sense light in three bands of color. The
biological pigment A biological pigment, also known simply as a pigment or biochrome, is a substance produced by living organisms that have a color resulting from selective Absorption (electromagnetic radiation), color absorption. Biological pigments include plant ...
s of the cones have maximum absorption values at wavelengths of about 420 nm (blue), 534 nm (bluish-green), and 564 nm (yellowish-green). The color of the pure signal of the cones could be described as violet, blue-green, and scarlet red, respectively, but, in their wavelengths of maximum absorption other cones are activated as well. The sensitivity ranges of the conecells overlap to provide vision throughout the
visible spectrum The visible spectrum is the spectral band, band of the electromagnetic spectrum that is visual perception, visible to the human eye. Electromagnetic radiation in this range of wavelengths is called ''visible light'' (or simply light). The optica ...
. The maximum
efficacy Efficacy is the ability to perform a task to a satisfactory or expected degree. The word comes from the same roots as '' effectiveness'', and it has often been used synonymously, although in pharmacology a distinction is now often made betwee ...
is 683 lm/W at a wavelength of 555 nm (green). By
definition A definition is a statement of the meaning of a term (a word, phrase, or other set of symbols). Definitions can be classified into two large categories: intensional definitions (which try to give the sense of a term), and extensional definitio ...
, light at a frequency of hertz has a luminous efficacy of 683 lm/W. The wavelengths for when a person is in photopic vary with the intensity of light. For the blue-green region (500 nm), 50% of the light reaches the image point of the
retina The retina (; or retinas) is the innermost, photosensitivity, light-sensitive layer of tissue (biology), tissue of the eye of most vertebrates and some Mollusca, molluscs. The optics of the eye create a focus (optics), focused two-dimensional ...
.
Adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the p ...
is much faster under photopic vision; it can occur in 5 minutes for photopic vision but it can take 30 minutes to transition from photopic to
scotopic In the study of visual perception, scotopic vision (or scotopia) is the vision of the eye under low-light conditions. The term comes from the Greek ''skotos'', meaning 'darkness', and ''-opia'', meaning 'a condition of sight'. In the human eye, co ...
. Most older adult humans lose photopic spatial contrast sensitivity. Adults in their 70s tend to require about 30–60% more contrast to detect high
spatial frequencies In mathematics, physics, and engineering, spatial frequency is a characteristic of any structure that is periodic across position in space. The spatial frequency is a measure of how often sinusoidal components (as determined by the Fourier tran ...
than adults in their 20s.{{cite journal, last=Burton, first=Kerri B., author2=Cynthia Owsley , author3=Michale E. Sloane , title=Aging and Neural Spatial Contrast Sensitivity: Photopic Vision, journal=Vision Research, date=4 June 1992, volume=33, issue=7, pages=939–949, doi=10.1016/0042-6989(93)90077-a, pmid=8506637, s2cid=26003597 The human eye uses scotopic vision under low-light conditions (luminance level 10−6 to 10−3.5 cd/m2), and
mesopic vision Mesopic vision, sometimes also called twilight vision, is a combination of photopic and scotopic vision under low-light (but not necessarily dark) conditions. Mesopic levels range approximately from 0.01 to 3.0  cd/m2 in luminance. Most ni ...
in intermediate conditions (luminance level 10−3 to 100.5 cd/m2).


See also

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Adaptation (eye) In visual physiology, adaptation is the ability of the retina of the eye to adjust to various levels of light. Natural night vision, or scotopic vision, is the ability to see under low-light conditions. In humans, rod cells are exclusively resp ...
*
Candela The candela (symbol: cd) is the unit of luminous intensity in the International System of Units (SI). It measures luminous power per unit solid angle emitted by a light source in a particular direction. Luminous intensity is analogous to radi ...
*
Cone cell Cone cells or cones are photoreceptor cells in the retina of the vertebrate eye. Cones are active in daylight conditions and enable photopic vision, as opposed to rod cells, which are active in dim light and enable scotopic vision. Most v ...
*
Contrast (vision) Contrast is the difference in luminance or color that makes an object (or its representation in an image or display) visible against a background of different luminance or color. The human visual system is more sensitive to contrast than to absol ...
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Mesopic vision Mesopic vision, sometimes also called twilight vision, is a combination of photopic and scotopic vision under low-light (but not necessarily dark) conditions. Mesopic levels range approximately from 0.01 to 3.0  cd/m2 in luminance. Most ni ...
*
Night vision Night vision is the ability to see in low-light conditions, either naturally with scotopic vision or through a night-vision device. Night vision requires both sufficient spectral range and sufficient intensity range. Humans have poor night v ...
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Purkinje effect The Purkinje effect or Purkinje phenomenon (; sometimes called the Purkinje shift, often pronounced ) is the tendency for the peak luminance sensitivity of the eye to shift toward the blue end of the color spectrum at low illumination (lighting), ...
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Photometry (optics) Photometry is a branch of optics that deals with measuring light Light, visible light, or visible radiation is electromagnetic radiation that can be visual perception, perceived by the human eye. Visible light spans the visible spectr ...
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Photosensitive ganglion cell Intrinsically photosensitive retinal ganglion cells (ipRGCs), also called photosensitive retinal ganglion cells (pRGC), or melanopsin-containing retinal ganglion cells (mRGCs), are a type of neuron in the retina of the mammalian eye. The presence ...
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Scotopic vision In the study of visual perception, scotopic vision (or scotopia) is the vision of the eye under low-light conditions. The term comes from the Greek ''skotos'', meaning 'darkness', and ''-opia'', meaning 'a condition of sight'. In the human eye, c ...


References

Eye Vision Color vision