The Philippine narrowmouth toad or the truncate-toed chorus frog (''Kaloula conjuncta'') is a species of

frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...

in the family

Microhylidae The Microhylidae, commonly known as narrow-mouthed frogs, are a geographically widespread family (biology), family of frogs. The 683 species are in 57 genera and 11 subfamilies. Evolution A molecular phylogenetic study by van der Meijden, et al. ...

. It is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...

. Its natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s are subtropical or tropical dry

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

s, subtropical or tropical moist lowland forests, subtropical or tropical moist

shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...

, subtropical or tropical seasonally wet or flooded lowland

grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...

s, intermittent rivers, freshwater

lake A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...

s, intermittent freshwater lakes, freshwater

marsh In ecology, a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p More in genera ...

es, intermittent freshwater marshes,

arable land Arable land (from the , "able to be ploughed") is any land capable of being ploughed and used to grow crops.''Oxford English Dictionary'', "arable, ''adj''. and ''n.''" Oxford University Press (Oxford), 2013. Alternatively, for the purposes of a ...

, pastureland,

plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...

s, rural gardens, heavily degraded former forest, water storage areas,

pond A pond is a small, still, land-based body of water formed by pooling inside a depression (geology), depression, either naturally or artificiality, artificially. A pond is smaller than a lake and there are no official criteria distinguishing ...

s, irrigated land, and seasonally flooded agricultural land. It is threatened by

habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

Range

''Kaloula conjuncta'' occurs throughout the Philippines, and has been recorded on the islands of

Negros Negros (, , ) is the fourth largest and third most populous island in the Philippines, with a total land area of . The coastal zone of the southern part of Negros is identified as a site of highest marine biodiversity importance in the Coral Tr ...

Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and a total population of 4,542,926, as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City of Il ...

Guimaras Guimaras (), officially the Province of Guimaras (; ), is an island province in the Philippines located in the Western Visayas region. The capital is Jordan, while the largest local government unit is the municipality of Buenavista. The provin ...

Cebu Cebu ( ; ), officially the Province of Cebu (; ), is a province of the Philippines located in the Central Visayas region, and consists of a main island and 167 surrounding islands and islets. The coastal zone of Cebu is identified as a ...

Luzon Luzon ( , ) is the largest and most populous List of islands in the Philippines, island in the Philippines. Located in the northern portion of the List of islands of the Philippines, Philippine archipelago, it is the economic and political ce ...

, Polillo,

Marinduque Marinduque (; ), officially the Province of Marinduque, is an island province in the Philippines located in Southwestern Tagalog Region or Mimaropa, formerly designated as Region IV-B. Its capital is the municipality of Boac, the most popul ...

Mindanao Mindanao ( ) is the List of islands of the Philippines, second-largest island in the Philippines, after Luzon, and List of islands by population, seventh-most populous island in the world. Located in the southern region of the archipelago, the ...

Leyte Leyte ( ) is an island in the Visayas group of islands in the Philippines. It is eighth-largest and sixth-most populous island in the Philippines, with a total population of 2,626,970 as of 2020 census. Since the accessibility of land has been ...

Patnanungan Patnanungan, officially the Municipality of Patnanungan (), is a municipality in the province of Quezon, Philippines. According to the 2020 census, it has a population of 15,052 people. History Patnanungan was once a barrio part of Polillo, da ...

Jomalig Jomalig ( ), officially the Municipality of Jomalig (), is a municipality in the province of Quezon, Philippines. According to the 2020 census, it has a population of 7,667 people. It is the least populated municipality in the province. It is p ...

Catanduanes Catanduanes (; ), officially the Province of Catanduanes (), is an island province located in the Bicol Region of Luzon in the Philippines. It is the 12th-largest island in the Philippines, and lies to the east of Camarines Sur, across the M ...

, Tablas,

Romblon Romblon (, , ), officially the Province of Romblon, is an archipelagic province of the Philippines located in the Mimaropa region. Its main components include Romblon, an archipelagic municipality of the same name that also serves as the provi ...

Sibuyan Sibuyan is a crescent-shaped island, the second largest in an archipelago comprising Romblon, Romblon Province, Philippines. Located in the namesake Sibuyan Sea, it has an area of and has a total population of 62,815, as of the 2020 census. Th ...

, and possibly

Mindoro Mindoro is the seventh largest and eighth-most populous island in the Philippines. With a total land area of 10,571 km2 ( 4,082 sq.mi ), it has a population of 1,408,454, as of the 2020 census. It is located off the southwestern coast of ...

Subspecies

There are currently 4 recognized subspecies of ''K. conjuncta'' (Inger 1954). Divergent varieties have been observed in

Eastern Visayas Eastern Visayas (; ; ; ) is an administrative region in the Philippines, designated as Region VIII. It consists of three main islands: Samar, Leyte, and Biliran. The region has six provinces: Biliran, Leyte, Northern Samar, Samar, Easte ...

(Samar-Leyte), and

.Blackburn, D. C., Siler, C. D., Diesmos, A. C., McGuire, J. A., Cannatella, D. C. and Brown, R. M. (2013), An adaptive radiation of frogs in a southeast Asian island archipelago. Evolution, 67: 2631–2646. The Sibuyan variety is most closely related to ''K. conjuncta negrosensis''. *''K. conjuncta conjuncta'':

*''K. conjuncta negrosensis'':

Visayas The Visayas ( ), or the Visayan Islands (Bisayan languages, Visayan: ''Kabisay-an'', ; Filipino language, Filipino: ''Kabisayaan'' ), are one of the three Island groups of the Philippines, principal geographical divisions of the Philippines, a ...

*''K. conjuncta meridionalis'':

(a small scansorial species with widely expanded digital disks) *''K. conjuncta stickeli'':

Samar Samar ( ) is the third-largest and seventh-most populous island in the Philippines, with a total population of 1,909,537 as of the 2020 census. It is located in the eastern Visayas, which are in the central Philippines. The island is divided in ...

(possibly a hybrid between ''K. picta'' and ''K. conjuncta meridionalis'')

References

Kaloula Amphibians of the Philippines Amphibians described in 1863 Taxa named by Wilhelm Peters Taxonomy articles created by Polbot {{Microhylidae-stub