Pheropsophus
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''Pheropsophus'' is a genus of
ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it ...
s in the family
Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal ...
. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxonomic revision of the North and Middle American species (Carabidae: Brachinida).


Species

These 163 species are members of the genus ''Pheropsophus'' * '' Pheropsophus abbreviatus'' Arrow, 1901 (sub-Saharan Africa) * '' Pheropsophus abyssinicus'' Alluaud, 1916 (Ethiopia) * '' Pheropsophus acutecostatus'' Fairmaire, 1892 (Madagascar) * '' Pheropsophus adrianae'' Giachino, 2005 (Indonesia and New Guinea) * '' Pheropsophus aequinoctialis'' (Linnaeus, 1763) (Mexico, Central, and South America) * '' Pheropsophus africanus'' (Dejean, 1825) (North Africa and Southwest Asia) * ''
Pheropsophus agnatus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Chaudoir, 1876 (Asia) * '' Pheropsophus akhaensis'' Kirschenhofer, 2010 (Thailand) * '' Pheropsophus alexandrae'' Giachino, 2003 (Australia) * '' Pheropsophus amnicola'' Darlington, 1968 (New Guinea) * '' Pheropsophus andrewesi'' Jedlicka, 1964 (India) * '' Pheropsophus androyanus'' Alluaud, 1932 * '' Pheropsophus angolensis'' (Erichson, 1843) (Sub-Saharan Africa) * ''
Pheropsophus aptinoides ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Chaudoir, 1876 (India) * '' Pheropsophus aptinomorphus'' Heller, 1910 (Indonesia and New Guinea) * ''
Pheropsophus arabicus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Arrow, 1901 (Middle East) * '' Pheropsophus arcanus'' (Erichson, 1843) (Sub-Saharan Africa) * ''
Pheropsophus assimilis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Chaudoir, 1876 (East and Southeast Asia) * ''
Pheropsophus azoulayi ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Lassalle & Schnell, 2018 (Philippines) * ''
Pheropsophus baehri ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Giachino, 2005 (Indonesia and New Guinea) * ''
Pheropsophus baliothorax ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Heller, 1910 (Indonesia and New Guinea) * ''
Pheropsophus balkei ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Giachino, 2005 (Indonesia and New Guinea) * ''
Pheropsophus basiguttatus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Chaudoir, 1876 (Sub-Saharan Africa) * ''
Pheropsophus beauvoisi ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' (Dejean, 1825) * ''
Pheropsophus beckeri ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Jedlicka, 1930 (China and temperate Asia) * ''
Pheropsophus bequaerti ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Burgeon, 1937 (Sub-Saharan Africa) * ''
Pheropsophus bidoupensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxon ...
'' Fedorenko, 2013 (Vietnam) * '' Pheropsophus bimaculatus'' (Linnaeus, 1771) (South Asia) * '' Pheropsophus bipartitus'' Fairmaire, 1869 (Madagascar) * '' Pheropsophus biplagiatus'' Chaudoir, 1876 (Mexico) * '' Pheropsophus bisulcatus'' Chaudoir, 1843 (Africa) * ''
Pheropsophus bohemani ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus ''Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxonomi ...
'' Chaudoir, 1876 (Africa) * '' Pheropsophus borkuanus'' Bruneau de Miré, 1990 (Chad) * '' Pheropsophus brussinoi'' Giachino, 2005 (Indonesia and New Guinea) * '' Pheropsophus canis'' Darlington, 1968 (New Guinea) * '' Pheropsophus capensis'' Chaudoir, 1876 (Sub-Saharan Africa) * '' Pheropsophus capitatus'' Jedlicka, 1935 (Philippines) * '' Pheropsophus cardoni'' Maindron, 1898 (India) * '' Pheropsophus catoirei'' (Dejean, 1825) (Southern Asia) * '' Pheropsophus catulus'' Darlington, 1968 (New Guinea) * '' Pheropsophus chaudoiri'' Arrow, 1901 (Indomalaya) * '' Pheropsophus cincticollis'' (LaFerté-Sénectère, 1850) (Sub-Saharan Africa) * '' Pheropsophus cinctus'' (Gory, 1833) (Africa and the Middle-East) * '' Pheropsophus claudiae'' Giachino, 2005 (New Guinea) * '' Pheropsophus congoensis'' Arrow, 1901 (Sub-Saharan Africa) * '' Pheropsophus consularis'' (Schmidt-Goebel, 1846) (Indomalaya) * '' Pheropsophus darwini'' Giachino, 2003 (Australia) * '' Pheropsophus debauvii'' (Guérin-Méneville, 1838) (South America) * '' Pheropsophus delmastroi'' Giachino, 2005 (Indonesia and New Guinea) * ''
Pheropsophus devagiriensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Venugopal & Sabu, 2019 (India) * '' Pheropsophus dilatatus'' Burgeon, 1936 (Ghana) * '' Pheropsophus dimidiatus'' Arrow, 1901 (Sub-Saharan Africa) * ''
Pheropsophus dissolutus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Andrewes, 1923 (Asia) * ''
Pheropsophus dongnaiensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus ''Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxonomi ...
'' (Fedorenko, 2021) (Vietnam) * '' Pheropsophus dregei'' Chaudoir, 1876 (Sub-Saharan Africa) * '' Pheropsophus dux'' Chaudoir, 1876 (Ethiopia) * '' Pheropsophus ecuadorensis'' Hubenthal, 1911 (Ecuador) * '' Pheropsophus emarginatus'' Chaudoir, 1876 (Asia, Philippines) * ''
Pheropsophus erjanii ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2017 (Indonesia) * '' Pheropsophus exiguus'' Arrow, 1901 (the Democratic Republic of the Congo) * '' Pheropsophus fastigiatus'' (Linnaeus, 1764) (Sub-Saharan Africa) * ''
Pheropsophus fimbriatus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Chaudoir, 1876 (Indomalaya) * '' Pheropsophus fulminans'' (Fabricius, 1801) * '' Pheropsophus fumigatus'' (Dejean, 1825) (Philippines) * '' Pheropsophus galloi'' Giachino, 2005 (Indonesia and New Guinea) * '' Pheropsophus gironieri'' Eydoux & Souleyet, 1839 (Indonesia and Philippines) * ''
Pheropsophus glabricollis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' (Fedorenko, 2020) (Vietnam) * '' Pheropsophus globulicollis'' Hubenthal, 1918 * '' Pheropsophus gracilis'' Arrow, 1901 * '' Pheropsophus gregoryi'' Giachino, 2003 (Australia) * '' Pheropsophus guanxiensis'' Kirschenhofer, 2010 (China and Vietnam) * '' Pheropsophus guineensis'' Chaudoir, 1876 * '' Pheropsophus halteri'' (Chaudoir, 1837) * '' Pheropsophus hassenteufeli'' Straneo, 1960 (Philippines) * '' Pheropsophus heathi'' Arrow, 1901 (Myanmar) * '' Pheropsophus hilaris'' (Fabricius, 1798) (Southern Asia) * '' Pheropsophus hispanicus'' (Dejean, 1824) (North Africa and the Mediterranean) * '' Pheropsophus humeralis'' Chaudoir, 1843 (the Comoro Islands, Madagascar, and Seychelles) * '' Pheropsophus impressicollis'' LaFerté-Sénectère, 1850 * ''
Pheropsophus indicus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Venugopal & Sabu, 2019 (India) * '' Pheropsophus infantulus'' Bates, 1892 (Myanmar, Thailand, and Vietnam) * '' Pheropsophus iranicus'' Reitter, 1919 (Iran) * ''
Pheropsophus jakli ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2015 (Indonesia) * ''
Pheropsophus jandai ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2019 (New Guinea and Papua) * '' Pheropsophus javanus'' (Dejean, 1825) (Palearctic) * '' Pheropsophus jessoensis'' A.Morawitz, 1862 (Eastern Asia) * '' Pheropsophus jurinei'' (Dejean, 1825) (Africa) * ''
Pheropsophus kabelkai ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2015 (Indonesia) * '' Pheropsophus katangensis'' Burgeon, 1937 (the Democratic Republic of the Congo) * '' Pheropsophus kersteni'' Gerstaecker, 1867 (Africa) * '' Pheropsophus kolbei'' Hubenthal, 1914 * '' Pheropsophus krichna'' (Maindron, 1906) (India) * '' Pheropsophus lafertei'' Arrow, 1901 * '' Pheropsophus langenhani'' Hubenthal, 1911 (Brazil) * '' Pheropsophus laticostis'' Chaudoir, 1876 * '' Pheropsophus lineifrons'' Chaudoir, 1850 (India and Pakistan) * ''
Pheropsophus lissoderus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Chaudoir, 1850 (Indomalaya) * '' Pheropsophus livingstoni'' Arrow, 1901 (Sub-Saharan Africa) * '' Pheropsophus longipennis'' Chaudoir, 1843 * ''
Pheropsophus lumawigi ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2019 (Philippines) * '' Pheropsophus maculicollis'' Hubenthal, 1914 (the Democratic Republic of the Congo) * '' Pheropsophus madagascariensis'' (Dejean, 1831) (Madagascar) * '' Pheropsophus malaisei'' Landin, 1955 (Myanmar) * '' Pheropsophus marginatus'' (Dejean, 1825) (Africa) * '' Pheropsophus marginicollis'' Motschulsky, 1854 (China) * '' Pheropsophus marginipennis'' (Laporte, 1835) * '' Pheropsophus mashunus'' Péringuey, 1896 (Sub-Saharan Africa) * '' Pheropsophus minahassae'' Heller, 1903 (Indonesia) * '' Pheropsophus minor'' Murray, 1857 (Sub-Saharan Africa) * ''
Pheropsophus montanus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' (Fedorenko, 2020) (Vietnam) * '' Pheropsophus nanodes'' Bates, 1892 (China and India) * '' Pheropsophus nebulosus'' Chaudoir, 1876 (Asia) * ''
Pheropsophus nepalensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Kirschenhofer, 2010 (Nepal) * ''
Pheropsophus ngoclinhensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus ''Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxonomi ...
'' Fedorenko, 2013 (Vietnam) * '' Pheropsophus nigerrimus'' Jedlicka, 1935 (Philippines) * '' Pheropsophus nigricollis'' Arrow, 1901 (India) * '' Pheropsophus nigriventris'' Chaudoir, 1878 (Africa) * '' Pheropsophus nyasae'' Arrow, 1901 * ''
Pheropsophus obiensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2014 (Indonesia) * '' Pheropsophus obliquatus'' (J.Thomson, 1858) * ''
Pheropsophus obliteratus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus ''Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxonomi ...
'' Fedorenko, 2013 (Vietnam) * '' Pheropsophus occipitalis'' (W.S.MacLeay, 1825) (Asia) * '' Pheropsophus pallidepunctatus'' Arrow, 1901 * '' Pheropsophus palmarum'' Chaudoir, 1876 (Ivory Coast) * '' Pheropsophus parallelus'' (Dejean, 1825) (Africa) * '' Pheropsophus pauliani'' Colas, 1944 (Cameroon) * '' Pheropsophus pedes'' Darlington, 1968 (Indonesia and New Guinea) * '' Pheropsophus perroti'' Arrow, 1901 (Madagascar) * '' Pheropsophus planti'' Chaudoir, 1876 (Mozambique and South Africa) * '' Pheropsophus platycephalus'' Reichardt, 1967 (Mexico) * '' Pheropsophus prophylax'' Heller, 1903 (India) * '' Pheropsophus raffrayi'' Chaudoir, 1878 (Africa) * '' Pheropsophus recticollis'' Arrow, 1901 (the Democratic Republic of the Congo) * '' Pheropsophus reductus'' Colas, 1944 * '' Pheropsophus riedeli'' Giachino, 2005 (Indonesia and New Guinea) * ''
Pheropsophus rivierii ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' (Demay, 1838) (South America) * '' Pheropsophus rolex'' Morvan, 1995 (Nepal) * '' Pheropsophus rufimembris'' Fairmaire, 1901 * ''
Pheropsophus sangirensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2017 (Indonesia) * '' Pheropsophus scythropus'' Andrewes, 1923 (South Asia) * '' Pheropsophus senegalensis'' (Dejean, 1825) (Africa) * '' Pheropsophus siamensis'' Chaudoir, 1876 (Southeast Asia) * ''
Pheropsophus similis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Fedorenko, 2013 (Vietnam) * '' Pheropsophus sobrinus'' (Dejean, 1826) (South Asia) * ''
Pheropsophus sonae ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2015 (Nigeria) * ''
Pheropsophus stenoderus ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus ''Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxonomi ...
'' Chaudoir, 1850 (Indomalaya) * '' Pheropsophus stenopterus'' Chaudoir, 1878 * '' Pheropsophus straneoi'' Giachino, 2003 (Australia) * ''
Pheropsophus sumatrensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' (Fedorenko, 2021) (Indonesia) * ''
Pheropsophus taclobanensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Lassalle & Schnell, 2019 (Philippines) * ''
Pheropsophus talaudensis ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Hrdlicka, 2014 (Indonesia) * '' Pheropsophus tamdaoensis'' Kirschenhofer, 2010 (Vietnam) * '' Pheropsophus transvaalensis'' Péringuey, 1896 (South Africa and Zimbabwe) * '' Pheropsophus tripustulatus'' (Fabricius, 1792) (India and Thailand) * '' Pheropsophus tristis'' Arrow, 1901 (Africa) * '' Pheropsophus ubomboensis'' Barker, 1919 (South Africa) * '' Pheropsophus uelensis'' Burgeon, 1937 (the Democratic Republic of the Congo) * '' Pheropsophus uniformis'' Hubenthal, 1914 (Africa) * '' Pheropsophus verticalis'' (Dejean, 1825) (Australia, Oceania) * ''
Pheropsophus windjanae ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' Baehr, 2012 (Australia) * '' Pheropsophus wolfi'' Giachino, 2005 (Indonesia and New Guinea) * ''
Pheropsophus worthingtoni ''Pheropsophus'' is a genus of ground beetles in the family Carabidae. It is considered a sister group of the genus '' Pheropsophidius'' within the subtribe Pheropsophina.Erwin, T. L. (1969). A reclassification of bombardier beetles and a taxono ...
'' * '' Pheropsophus yunnanensis'' Kirschenhofer, 2010 (China)


References


External links

* {{Taxonbar, from=Q1757457 Brachininae Carabidae genera