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''Phalaris aquatica'', known by the common names bulbous canary-grass and Harding grass, is a species of grass in the genus ''
Phalaris Phalaris () was the tyrant of Akragas (now Agrigento) in Sicily in Magna Graecia, from approximately 570 to 554 BC. History Phalaris was renowned for his excessive cruelty. Among his alleged atrocities is cannibalism: he was said to have eaten ...
'' of the family
Poaceae Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivate ...
.


Description

It is an erect, waist-high, stout perennial
bunch grass Tussock grasses or bunch grasses are a group of grass species in the family Poaceae. They usually grow as singular plants in clumps, tufts, hummocks, or bunches, rather than forming a sod or lawn, in meadows, grasslands, and prairies. As perennia ...
, with grayish to bluish green leaves. Flowering heads are dense, spike-like, and usually long. It is slow to develop from seed, but can form large bunches after several years. ''
Phalaris arundinacea ''Phalaris arundinacea'', or reed canary grass, is a tall, perennial bunchgrass that commonly forms extensive single-species stands along the margins of lakes and streams and in wet open areas, with a wide distribution in Europe, Asia, northern ...
'' (reed canary grass) differs from Harding grass in having more distinct rhizomes, and an inflorescence that is compact at first but becomes more open as the branches spread. Hybrids of Harding grass and reed canary grass have been produced. Varieties include 'AQ1', 'Uneta', and 'Australis'. ''P. aquatica'' is a quick-growing grass which incorporates and utilises soil nitrogen rapidly.


Geography

''Phalaris aquatica'' originated from
Southern Europe Southern Europe is also known as Mediterranean Europe, as its geography is marked by the Mediterranean Sea. Definitions of southern Europe include some or all of these countries and regions: Albania, Andorra, Bosnia and Herzegovina, Bulgaria, C ...
and the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
. It is naturalized in South Africa, Australia, New Zealand and the USA. Although very recently introduced there, its
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Types of pasture Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, c ...
value was first recognised in Australia. Domesticated cultivation then spread to the United States, Argentina and several other countries in South America, and New Zealand.


Chemical constituents

Leaves and seedlings contain
gramine Gramine (also called donaxine) is a naturally occurring indole alkaloid present in several plant species. Gramine may play a defensive role in these plants, since it is toxic to many organisms. Occurrence Gramine has been found in the giant reed, ...
. And
dimethyltryptamine Dimethyltryptamine (DMT), also known as ''N'',''N''-dimethyltryptamine (''N'',''N''-DMT), is a Psychedelic drug, serotonergic hallucinogen and Investigational New Drug, investigational drug of the substituted tryptamine, tryptamine family tha ...


Forage

''Phalaris aquatica'' is a pasture species grazed by ruminants. The grass can also be cut, providing good quality fodder for grazing livestock for 8 to 12 months a year.


Toxicity

Some ''Phalaris'' species contain
gramine Gramine (also called donaxine) is a naturally occurring indole alkaloid present in several plant species. Gramine may play a defensive role in these plants, since it is toxic to many organisms. Occurrence Gramine has been found in the giant reed, ...
(also found in Avena sativa 'Oats') which was ''claimed '' to cause brain damage, other organ damage,
central nervous system The central nervous system (CNS) is the part of the nervous system consisting primarily of the brain, spinal cord and retina. The CNS is so named because the brain integrates the received information and coordinates and influences the activity o ...
damage and death in sheep (when injected in very large amounts) although ''Phalaris aquatica'' is ''said'' to be non-toxic to them. In
Victoria Victoria most commonly refers to: * Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India * Victoria (state), a state of Australia * Victoria, British Columbia, Canada, a provincial capital * Victoria, Seychelles, the capi ...
, Australia, it is reported that
kangaroos Kangaroos are marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use, the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern grey ...
grazing on ''Phalaris aquatica'' may develop a condition known familiarly as "Phalaris staggers", in which coordination and mobility are affected.


Invasive species

Harding grass is an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
in
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
, oak woodland,
chaparral Chaparral ( ) is a shrubland plant plant community, community found primarily in California, southern Oregon, and northern Baja California. It is shaped by a Mediterranean climate (mild wet winters and hot dry summers) and infrequent, high-intens ...
, and
riparian A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripar ...
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s. Native grasses and grassland habitat in California are affected.


References


External links


Jepson Manual Treatment
{{Taxonbar, from=Q1750612 Forages aquatica Psychedelic tryptamine carriers Taxa named by Carl Linnaeus Flora of Malta Grasses of Lebanon