Phalangodidae
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The Phalangodidae are a family of
harvestmen The Opiliones (formerly Phalangida) are an Order (biology), order of arachnids, Common name, colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs (see below). , over 6,650 species of harvestmen have been discovered w ...
with about 30
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
and more than 100 described
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, distributed in the Holarctic region. It is not to be confused with the harvestman family Phalangiidae, which is in the
suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
Eupnoi The Eupnoi are a suborder of harvestmen, with more than 200 genera, and about 1,700 described species. They consist of two superfamilies, the Phalangioidea with many long-legged species common to northern temperate regions, and the small group ...
.


Name

The name of the type genus ''Phalangodes'' is derived from
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
('), a line of soldiers in formation. This probably refers to the rows of
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
al spines.


Description

Body length ranges from less than 1 mm to about . The pedipalps are armed with large spines. While some species have legs eight times the body length, others have shorter legs around twice the body length. Most species are yellowish to orange brown.
Troglobitic A troglobite (or, formally, troglobiont) is an animal species, or population of a species, strictly bound to underground habitats, such as caves. These are separate from species that mainly live in above-ground habitats but are also able to live u ...
(cave-dwelling) species are depigmented.


Distribution

More than two-thirds of the species occur in the western
Nearctic The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface. The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...
, especially in California] with about 50 species in four endemism, endemic genera. About 10 species are known from the eastern Nearctic. Few occur in the
Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...
, with one species each in the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
and
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
, and about 20 species in the Mediterranean region.


Relationships

The family seems to be largely
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
, with the exceptions of a few genera such as ''Guerrobunus'' and ''Glennhuntia''. Although the relationship of the Phalangodidae to other
Grassatores The Grassatores Kury, 2002 are the most diverse infraorder of the Laniatores. It includes over 3,700 species distributed mainly in the tropicsKury, A.B.(2003). Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). '' Revi ...
is currently unresolved, the family seems relatively basal inside the Grassatores.


Genera

The family Phalangodidae contains these genera:Joel Hallan's Biology Catalog
Phalangodidae
/ref> * '' Ausobskya'' Martens, 1972 c g
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
(four species) * '' Banksula'' Roewer, 1949 i c g b — USA (
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
) (10 or 11 species) * '' Bishopella'' Roewer, 1927 i c g b — USA (southeastern states) (two species) * '' Bogania'' Forster, 1955 c g — Australia (five species) * '' Bunofagea'' Staręga, 1992 c g — Madagscar (two species) * '' Calicina'' Ubick & Briggs, 1989 i c g b — USA (California) (25 species) * '' Chinquipellobunus'' Goodnight and Goodnight, 1944 i c g * '' Crosbyella'' Roewer, 1927 i c g b — southeastern US (five species) * '' Enigmina'' Ubick & Briggs, 2008 i c g b * '' Glennhuntia'' Shear, 2001 — western Australia (one species; probably misplaced) * '' Guerrobunus'' Goodnight & Goodnight, 1945
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
(three species, probably misplaced) * ''
Lola Lola may refer to: Places * Lolá, a or subdistrict of Panama * Lola Township, Cherokee County, Kansas, United States * Lola Prefecture, Guinea * Lola, Guinea, a town in Lola Prefecture * Lola Island, in the Solomon Islands People * Lol ...
'' Kratochvil, 1937 g — Israel (one species) * '' Maiorerus'' Rambla, 1993 g
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
(one species) * '' Megacina'' Ubick & Briggs, 2008 i c g b * '' Microcina'' Briggs & Ubick, 1989 i c g b — California (six species) * '' Microcinella'' Ubick & Briggs, 2008 i c g b * '' Neoparalus'' Özdikmen, 2006 i c g * '' Paralola'' Kratochvil, 1951 g — Bulgaria (one species) * '' Phalangodes'' Tellkampf, 1844 i c g b — Kentucky, Cuba? (two species; one species possibly misplaced) * '' Phalangomma'' Roewer, 1949 — Virginia (one species; probably misplaced) * '' Proscotolemon'' Roewer, 1916
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
(one species) * '' Ptychosoma'' Sørensen, 1873 — Spain, Italy, North Africa (two species) * '' Scotolemon'' Lucas, 1860 g — Mediterranean (13 species) * '' Sitalcina'' Banks, 1911 i c g b — California (9 species) * '' Spalicus'' Roewer, 1949 i c g * '' Texella'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Texas, California, Oregon, New Mexico (28 species) * '' Tolus'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Tennessee (one species) * '' Tularina'' Ubick & Briggs, 2008 i c g b * '' Undulus'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Alabama (one species) * '' Wespus'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Arkansas (one species) * '' Alpazia'' Özdikmen & Kury, 2006 i c g * '' Haasus'' Roewer, 1949 (See Pyramidopidae) Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net


References

{{Authority control Harvestmen Harvestman families