
The Phalangodidae are a family of
harvestmen
The Opiliones (formerly Phalangida) are an Order (biology), order of arachnids Common name, colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. , over 6,650 species of harvestmen have been discovered worldwide, alth ...
with about 30
genera
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...
and more than 100 described
species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
, distributed in the Holarctic region.
It is not to be confused with the harvestman family
Phalangiidae
The Phalangiidae are a family of harvestmen with about 380 known species. The best known is ''Phalangium opilio''. '' Dicranopalpus ramosus'' is an invasive species in Europe.
It is not to be confused with the harvestman family Phalangodidae, w ...
, which is in the
suborder
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
Eupnoi
The Eupnoi are a suborder of harvestmen, with more than 200 genera, and about 1,700 described species.
They consist of two superfamilies, the Phalangioidea with many long-legged species common to northern temperate regions, and the small group C ...
.
Name
The name of the type genus ''Phalangodes'' is derived from
Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
('), a line of soldiers in formation. This probably refers to the rows of
pedipalp
Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") a ...
al spines.
Description
Body length ranges from less than 1 mm to about . The pedipalps are armed with large spines. While some species have legs eight times the body length, others have shorter legs around twice the body length. Most species are yellowish to orange brown.
Troglobitic
A troglobite (or, formally, troglobiont) is an animal species, or population of a species, strictly bound to underground habitats, such as caves. These are separate from species that mainly live in above-ground habitats but are also able to live u ...
(cave-dwelling) species are depigmented.
[
]
Distribution
More than two-thirds of the species occur in the western Nearctic
The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface.
The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...
, especially in California] with about 50 species in four endemism, endemic genera. About 10 species are known from the eastern Nearctic. Few occur in the Palearctic
The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa.
The realm consists of several bioregions: the Euro-Si ...
, with one species each in the Canary Islands
The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Mo ...
and Japan, and about 20 species in the Mediterranean region
In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and ...
.[
]
Relationships
The family seems to be largely monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
, with the exceptions of a few genera such as ''Guerrobunus'' and ''Glennhuntia''. Although the relationship of the Phalangodidae to other Grassatores
The Grassatores are the most diverse infraorder of the Laniatores. It includes over 3,500 species distributed mainly in the tropicsKury, A.B.(2003). Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). '' Revista Ibéri ...
is currently unresolved, the family seems relatively basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
inside the Grassatores.[
]
Genera
The family Phalangodidae contains these genera:[Joel Hallan's Biology Catalog]
Phalangodidae
/ref>[
* '' Alpazia'' Özdikmen & Kury, 2006 i c g
* '' Ausobskya'' Martens, 1972 g — ]Greece
Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wit ...
(four species)
* '' Banksula'' Roewer, 1949 i c g b — California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
(10 species)
* '' Bishopella'' Roewer, 1927 i c g b — southeastern US (two species)
* '' Bogania'' Forster, 1955 g
* '' Calicina'' Ubick & Briggs, 1989 i c g b — California (25 species)
* '' Chinquipellobunus'' C.J. Goodnight and M.L. Goodnight, 1944 i c g
* '' Crosbyella'' Roewer, 1927 i c g b — southeastern US (five species)
* '' Enigmina'' Ubick & Briggs, 2008 i c g b
* '' Glennhuntia'' Shear, 2001 — western Australia (one species; probably misplaced)
* '' Guerrobunus'' Goodnight & Goodnight, 1945 — Mexico
Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
(three species, probably misplaced)
* '' Haasus'' Roewer, 1949 g
* ''Lola
Lola may refer to:
Places
* Lolá, a or subdistrict of Panama
* Lola Township, Cherokee County, Kansas, United States
* Lola Prefecture, Guinea
* Lola, Guinea, a town in Lola Prefecture
* Lola Island, in the Solomon Islands
People
* Lol ...
'' Kratochvil, 1937 g — Israel (one species)
* '' Maiorerus'' Rambla, 1993 g — Canary Islands
The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Mo ...
(one species)
* '' Megacina'' Ubick & Briggs, 2008 i c g b
* '' Microcina'' Briggs & Ubick, 1989 i c g b — California (six species)
* '' Microcinella'' Ubick & Briggs, 2008 i c g b
* '' Neoparalus'' Özdikmen, 2006 i c g
* '' Paralola'' Kratochvil, 1951 g — Bulgaria (one species)
* '' Phalangodes'' Tellkampf, 1844 i c g b — Kentucky, Cuba? (two species; one species possibly misplaced)
* '' Phalangomma'' Roewer, 1949 — Virginia (one species; probably misplaced)
* '' Proscotolemon'' Roewer, 1916 — Japan (one species)
* '' Ptychosoma'' Sørensen, 1873 — Spain, Italy, North Africa (two species)
* '' Scotolemon'' Lucas, 1860 g — Mediterranean (13 species)
* '' Sitalcina'' Banks, 1911 i c g b — California (9 species)
* '' Spalicus'' Roewer, 1949 i c g
* ''Texella
''Texella'' is a genus of armoured harvestmen in the family Phalangodidae. There are more than 20 described species in ''Texella''.
Species
These 21 species belong to the genus ''Texella'':
* ''Texella bifurcata'' (Briggs, 1968)
* ''Texella b ...
'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Texas, California, Oregon, New Mexico (28 species)
* '' Tolus'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Tennessee (one species)
* '' Tularina'' Ubick & Briggs, 2008 i c g b
* '' Undulus'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Alabama (one species)
* '' Wespus'' C.J. Goodnight & M.L. Goodnight, 1942 i c g b — Arkansas (one species)
Data sources: i = ITIS,[ c = Catalogue of Life,][ g = GBIF,][ b = Bugguide.net]
References
{{Taxonbar, from=Q3377994
Harvestmen
Harvestman families