Phaceloolithidae
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''Phaceloolithus'' is an
oogenus Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains ...
of
dinosaur egg Dinosaur eggs are the organic vessels in which a dinosaurs, dinosaur embryo develops. When the first scientifically documented remains of non-avian dinosaurs were being described in England during the 1820s, it was presumed that dinosaurs had la ...
found in the Fenshui'ao Formation of the
Dongting Dongting Lake () is a large, shallow lake in northeastern Hunan Province, China. It is a flood basin of the Yangtze River, so its volume depends on the season. The provinces of Hubei and Hunan are named after their location relative to the la ...
Basin of the
Hunan Province Hunan is an inland province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangxi to the east, Guangdong and Guangxi to the south, and Guizhou and Chon ...
of
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. The eggs have a subspherical shape, measuring up to 168 mm on the long axis, and having a very thin shell.D. Zeng and J. Zhang. (1979).
On the dinosaurian eggs from the western Dongting Basin, Hunan
. ''Vertebrata PalAsiatica'' 17(2):131–136
Carpenter, K. (1999). ''Eggs, Nests, and Baby Dinosaurs: A Look at Dinosaur Reproduction'' (Life of the Past). Indiana University Press, Bloomington, Indiana.


Distribution

''Phaceloolithus'' is found at the Dongting Basin in Hunan. They are dated, on the basis of other fossil eggs,
ostracods Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 33,000 species (only 13,000 of which are extant) have been identified,Brandão, S.N.; Antonietto, L.S; Nery, D.G.; Santos, S.G.; Karano ...
(seed shrimp), and
charophytes Charophyta () is a Paraphyly, paraphyletic group of freshwater green algae, called charophytes (), sometimes treated as a Division (taxonomy), division, yet also as a Division (taxonomy), superdivision. The terrestrial plants, the Embryophyta em ...
(algae) found at the same site, to the
late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
.


Description

''Phaceloolithus'' is nearly spherical, measuring 167–168 mm long and 140–150 mm across. The eggshell is very thin, measuring from 0.5 to 0.7 mm thick. Like most other non- avian dinosaur eggs,Laura E. Wilson, Karen Chin, Frankie D. Jackson, and Emily S. Bray. (2012).
Fossil eggshell: Fragments from the past
UCMP The University of California Museum of Paleontology (UCMP) is a paleontology museum located on the campus of the University of California, Berkeley. The museum is within the Valley Life Sciences Building (VLSB), designed by George W. Kelham a ...
's online fossil egg exhibit.
''Phaceloolithuss shell is stratified into two layers: the prismatic layer on the outside, and the mammillary layer on the inside. The mammillae are compressed into groups of two or three, and the pore canals are well-developed.


History

''Phaceloolithus'' was first described as the sole species of the oofamily Phaceloolithidae in
1979 Events January * January 1 ** United Nations Secretary-General Kurt Waldheim heralds the start of the ''International Year of the Child''. Many musicians donate to the ''Music for UNICEF Concert'' fund, among them ABBA, who write the song ...
by Chinese paleontologists Zeng Demin and Zhang Jinjian, following its discovery in the Dongting basin of
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
.


Parataxonomy

While ''Phaceloolithus'' has always been considered an oospecies of Phaceloolithidae, there is dispute about whether or not this oofamily is synonymous with the later-named
Dendroolithidae ''Dendroolithus'' is an oogenus of Dendroolithid dinosaur egg found in the late Cenomanian Chichengshan Formation ( Tiantai Group), in the Gong-An-Zhai and Santonian Majiacun Formations of China and the Maastrichtian Nemegt and Campanian B ...
. The oofamilies were first synonymized by Fang ''
et al. References Notes References Further reading * * External links * {{Latin phrases E ...
'' in
2003 2003 was designated by the United Nations as the International Year of Fresh water, Freshwater. In 2003, a Multi-National Force – Iraq, United States-led coalition 2003 invasion of Iraq, invaded Iraq, starting the Iraq War. Demographic ...
.Fang, Xiao-si, et al. "Cretaceous fossil eggs from the Tiantai basin of Zhejiang, with a discussion on the extinction of dinosaurs." ''Geological Bulletin of China'' 22.7 (2003): 1–24. This position was supported by Ribeiro ''et al.'' in
2014 The year 2014 was marked by the surge of the Western African Ebola epidemic, West African Ebola epidemic, which began in 2013, becoming the List of Ebola outbreaks, most widespread outbreak of the Ebola, Ebola virus in human history, resul ...
,Ribeiro, V., Mateus, O., Holwerda, F., Araújo, R., & Castanhinha, R. (2014). "Two new theropod egg sites from the Late Jurassic Lourinhã Formation, Portugal." ''Historical Biology'', 26(2), 206–217. but other authors have considered the two oofamilies distinct,Barta, D. E., Brundridge, K. M., Croghan, J. A., Jackson, F. D., Varricchio, D. J., Jin, X., & Poust, A. W. (2014). "Eggs and clutches of the Spheroolithidae from the Cretaceous Tiantai basin, Zhejiang Province, China." ''Historical Biology'', 26(2), 183–194. and in his master's thesis, Daniel Barta argued against synonymy because the brevity of the description of Phaceloolithidae makes any further referrals to that oofamily uncertain.Barta, D. E. (2014). "A phylogenetic approach to understanding dinosaur egg diversity and the evolution of reproductive traits within Dinosauria" (Master's thesis, Montana State University-Bozeman, College of Letters & Science). If the synonymy is correct, then ''Phaceloolithus'' would be a sister taxon to ''
Dendroolithus ''Dendroolithus'' is an oogenus of Dendroolithid dinosaur egg found in the late Cenomanian Chichengshan Formation ( Tiantai Group), in the Gong-An-Zhai and Santonian Majiacun Formations of China and the Maastrichtian Nemegt and Campanian B ...
'' and '' Paradendroolithus''. If not, then ''Phaceloolithus'' would be the sole oogenus of Phaceloolithus.Fang Xiao-si, Yue Zhao, Ling Hong. (2009). "近十五年来蛋化石研究概况 eview of the Past 15 years’ Research on Fossil Eggs in China. 地球学报, 30(4), 523–542.


See also

*
List of dinosaur oogenera Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains of t ...


References

{{Taxonbar, from=Q7179941 Dinosaur reproduction Fossil parataxa described in 1979