Penstemon Anguineus
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''Penstemon anguineus'' is a species of
penstemon ''Penstemon'' , the beardtongues, is a large genus of roughly 280 species of flowering plants native to North America from northern Canada to Central America. It is the largest genus of flowering plants endemic to North America. As well as bein ...
known by the common name Siskiyou penstemon. It is native to the mountains of southern
Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...
and northern
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
, where it grows in
coniferous forests Conifers () are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All e ...
, often in open areas left by
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidder, skidding, on-site processing, and loading of trees or trunk (botany), logs onto logging truck, trucksherbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...
, one without woody stems, that may reach as much as 90 centimeters in height. Though it may be occasionally as short as 10 cm at maturity it is more often taller than 30 cm. Both
basal leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...
, ones attached directly to the base of the plant, and the lower
cauline A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...
leaves, ones attached to the stems are 30 to 150 millimeters long and 6 to 40 mm wide. Basal leaves are
lanceolate The following terms are used to describe leaf plant morphology, morphology in the description and taxonomy (biology), taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade ...
, shaped like a spear head, to narrowly
ovate Ovate may refer to: * Ovate (egg-shaped) leaves, tepals, or other botanical parts *Ovate, a type of prehistoric stone hand axe * Ovates, one of three ranks of membership in the Welsh Gorsedd * Vates or ovate, a term for ancient Celtic bards ...
, egg shaped but thinner. Leaf edges are dentate, having symmetrical teeth, to
serrate Serration is a saw-like appearance or a row of sharp or tooth-like projections. A serrated cutting edge has many small points of contact with the material being cut. By having less contact area than a smooth blade or other edge, the applied pr ...
, with distinct forward pointing saw teeth along the edge. Some plants may have smooth edged leaves or only have very fine ones. There are two to four pairs of leaves attached to the stems. They are arranged oppositely with ones higher on the stem clasping it, with the base of the leaf partly wrapped around it. They may be 11 to 90 millimeters long and 5–50 mm in width. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
produces several light blue or purple flowers. The flowers have violet
nectar guides Nectar guides are markings or patterns seen in flowers of some angiosperm species, that guide pollinators to their rewards. These markings may appear as lines, spots, or "blotches". Such patterns are also known as "pollen guides" and "honey gui ...
and 13–18 millimeters long. The
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s and flowers are covered in glandular hairs. The inside of the flower has many white hairs. The smooth to sparely hairy
staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co ...
is 10–13 mm long and extends out of the opening of the flower tube. When present the staminode hairs are yellow.


Taxonomy

This species was scientifically described for the first time in 1880 as a
variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
under the name '' Penstemon rattanii'' var. ''minor'' by the botanist Asa Gray. Independently,
Alice Eastwood __NOTOC__ Alice Eastwood (January 19, 1859 – October 30, 1953) was a Canadian American botanist. She is credited with building the botanical collection at the California Academy of Sciences in San Francisco. She published over 310 scient ...
described it as a species named ''Penstemon anguineus'' in 1905. Because of the differing descriptions the earlier variety and the 1940 description of it by David D. Keck as ''Penstemon minor'' are considered
heterotypic synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently ...
s. The specimen described as
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...
of the species was collected from Shelley Creek in
Del Norte County, California Del Norte County ( Spanish for "Of The North") is a county located at the far northwest corner of the U.S. state of California, along the Pacific Ocean adjacent to the Oregon border. Its population was 27,743 as of the 2020 census, down from ...
.


Names

The
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
s in English for ''Penstemon anguineus'' include Siskiyou penstemon and tongue-leaved penstemon.


Range and habitat

The native range for Siskiyou penstemon is in the US states of California and Oregon. In California it grows in the northern part of the state in the higher parts of the
North Coast Ranges The Coast Ranges of California span from Del Norte or Humboldt County, California, south to Santa Barbara County. The other three coastal California mountain ranges are the Transverse Ranges, Peninsular Ranges and the Klamath Mountains. Phy ...
and
Klamath Ranges The Klamath Mountains are a rugged and lightly populated mountain range in northwestern California and southwestern Oregon in the western United States. As a mountain system within both the greater Pacific Coast Ranges and the California Coast R ...
. The distribution of the species continues northward to the area around
Crater Lake Crater Lake ( Klamath: ) is a volcanic crater lake in south-central Oregon in the Western United States. It is the main feature of Crater Lake National Park and is a tourist attraction for its deep blue color and water clarity. T ...
in southern Oregon including Douglas County. In
coniferous forest Conifers () are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All e ...
s it grows in forest openings including those produced by logging activities. It also grows on mountainsides and ridges and the transition to
chaparral Chaparral ( ) is a shrubland plant plant community, community found primarily in California, southern Oregon, and northern Baja California. It is shaped by a Mediterranean climate (mild wet winters and hot dry summers) and infrequent, high-intens ...
habitats.


See also

List of ''Penstemon'' species


External links

*


References

{{Taxonbar, from=Q7164832 anguineus Flora of California Flora of Oregon