The pennaceous feather is a type of

feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...

present in most modern

birds Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

and in some other species of

maniraptoriform Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyran ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

Description

A pennaceous

has a stalk or quill. Its basal part, called a ''calamus'', is embedded in the skin. The calamus is hollow and has pith formed from the dry remains of the feather pulp, and the calamus opens below by an ''inferior umbilicus'' and above by a ''superior umbilicus''. The stalk above the calamus is a solid ''

rachis In biology, a rachis (from the [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this case the ''rachi ...

'' having an umbilical groove on its underside. Pennaceous feathers have a rachis with ''vanes'' or ''vaxillum '' spreading to either side. These vanes are composed of a high number of flattened ''

barb Barb or the BARBs or ''variation'' may refer to: People * Barb (name), a list of people and fictional characters with the given name or surname * Barb, a term used by fans of Nicki Minaj to refer to themselves * The Barbs, a band Places * Barb, ...

s'', that are connected to one another with '' barbules''. The barbules are tiny strands that criss-cross on the flattened sides of the barbs. This forms a miniature

velcro Velcro IP Holdings LLC, trading as Velcro Companies and commonly referred to as Velcro (pronounced ), is a British privately held company, founded by Swiss electrical engineer George de Mestral in the 1950s. It is the original manufacturer of ho ...

-like mesh that holds all the barbs together, stabilizing the vanes. Pennaceous feathers on the wing, and elsewhere, where stresses related to flight, or other activities, are high, are accordingly attached especially strongly. This strong attachment is accomplished by ligaments under the skin, which in some birds and other

feathered dinosaurs A feathered dinosaur is any species of dinosaur possessing feathers. That includes all species of birds, and in recent decades evidence has accumulated that many non-avian dinosaur species also possessed feathers in some shape or form. The exte ...

results in raised bumps or marks along the rear forelimb bone (

ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...

). These bumps, called quill knobs (ulnar papillae), are often used as an indirect indication of strongly-attached forelimb feathers in

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

species, and can also indirectly indicate the number of secondary flight feathers in a given specimen.

Flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...

s (

remiges Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the ta ...

and

rectrices Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the Bird wing, wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those ...

) are specialized types of pennaceous feathers, adapted for high loadings and often strongly asymmetric for improved

flight Flight or flying is the motion (physics), motion of an Physical object, object through an atmosphere, or through the vacuum of Outer space, space, without contacting any planetary surface. This can be achieved by generating aerodynamic lift ass ...

performance.

References

Feathers {{ornithology-stub

Description

See also

References